Turing Bifurcation Research Articles

Pattern formation on regular polygons and circles

We investigate the formation of Turing patterns on regular polygonal domains, as the number of edges grow, leading to the limiting case of the circle. Using linear and weakly nonlinear analysis, and evidence by simulations, we demonstrate how the domain shape can fundamentally change the expected bifurcation structure. Specifically, on the square domain we are able to derive pitchfork bifurcations for stripe and spot solutions, as well as show that both branches cannot bifurcate to produce stable patterns. This compares with the case of the equilateral triangle domain that causes the Turing bifurcation to be generically transcritical and, in some cases, none of the bifurcating branches are stable. Moreover, we find a monotonically increasing, but nonlinear relationship, between the minimal bifurcation area and the number of edges. Thus, patterns can occur on triangles with much smaller areas than circles. Overall, this work raises questions for researchers who are simulating applications on domains with simple shapes. Specifically, even small changes to domain geometry can have large impacts on the produced patterns; thus, domain perturbations should be considered in any sensitivity analyses.

A diffusive plant-sulphide model: spatio-temporal dynamics contrast between discrete and distributed delay

Abstract This paper studies the spatio-temporal dynamics of a diffusive plant-sulphide model with toxicity delay. More specifically, the effects of discrete delay and distributed delay on the dynamics are explored, respectively. The deep analysis of eigenvalues indicates that both diffusion and delay can induce Hopf bifurcations. The normal form theory is used to set up an exact formula that determines the properties of Hopf bifurcation in a diffusive plant-sulphide model. A sufficiently small discrete delay does not affect the stability and a sufficiently large discrete delay destabilizes the system. Nonetheless, a sufficiently small or large distributed delay does not affect the stability. Both delays cause instability by inducing Hopf bifurcation rather than Turing bifurcation.

A Framework to Identify Supercritical and Subcritical Turing Bifurcations: Case Study of Soai Reaction Featuring Asymmetric Autocatalysis

A Framework to Identify Supercritical and Subcritical Turing Bifurcations: Case Study of Soai Reaction Featuring Asymmetric Autocatalysis

Effects of extra resource and harvesting on the pattern formation for a predation system

We deal with a reaction–diffusion predation system with extra resource provided to predator and harvesting on prey. We first discuss the long-time behaviors of parabolic system, including the dissipativeness and persistence of positive solutions. It is shown that under certain constraints of harvesting, the quality of extra resource, the predation and transition rates of predator, the dissipativeness is compatible with persistence. Secondly, some properties of steady-state system are investigated, mainly including the existence of non-constant positive solutions, Turing and steady-state bifurcation phenomena. It is found that the extra resource, prey harvesting and diffusion have significant impacts on the pattern formations. Furthermore, some numerical simulations on Turing patterns and steady-state bifurcation solutions are performed to illustrate the theoretical analysis. We observe that when the quantity of extra resources is low, changes in the quality of extra resources can lead to significant changes in the spatial distribution of species, which is in sharp contrast to the case of high quantity of extra resource. Additionally, we conclude that different diffusion rates of predator can lead to different spatial patterns for the system.

Spatiotemporal complexity analysis of a discrete space-time cancer growth model with self-diffusion and cross-diffusion

We investigate spatiotemporal pattern formation in cancer growth using discrete time and space variables. We first introduce the coupled map lattices (CMLs) model and provide a dynamical analysis of its fixed points along with stability results. We then offer parameter criteria for flip, Neimark–Sacker, and Turing bifurcations. In the presence of spatial diffusion, we find that stable homogeneous solutions can experience Turing instability under certain conditions. Numerical simulations reveal a variety of spatiotemporal patterns, including patches, spirals, and numerous other regular and irregular patterns. Compared to previous literature, our discrete model captures more complex and richer nonlinear dynamical behaviors, providing new insights into the formation of complex patterns in spatially extended discrete tumor models. These findings demonstrate the model’s ability to capture complex dynamics and offer valuable insights for understanding and treating cancer growth, highlighting its potential applications in biomedical research.

Pattern dynamics of a Lotka-Volterra model with taxis mechanism

This paper deals with the Turing bifurcation and pattern dynamics of a Lotka-Volterra model with the predator-taxis and the homogeneous no-flux boundary conditions. To investigate the pattern dynamics, we first give the occurrence conditions of the Turing bifurcation. It is found that there is no Turing bifurcation when predator-taxis disappears, while the Turing bifurcation occurs as predator-taxis is presented. Next, we establish the amplitude equation by virtue of weakly nonlinear analysis. Our theoretical result suggests the Lotka-Volterra model admits the supercritical or subcritical Turing bifurcation. In this manner, we can determine the stability of the bifurcating solution. Finally, some numerical simulation results verify the validity of the theoretical analysis. The stripe pattern, the mixed patterns, and wave patterns are performed. Interestingly, the stable stripe patterns will be broken and become wave patterns when the predator-taxis parameter is far from the Turing bifurcation critical point.

Turing–Hopf bifurcation in a diffusive predator–prey model with schooling behavior and Smith growth

This paper explores the dynamics of a diffusive predator–prey model, considering schooling behavior and Smith growth in prey. Initially, we have formulated the pertinent characteristic equations. Subsequently, We proceed to examine the existence of the Turing bifurcation and Hopf bifurcation, phenomena that describe the emergence of spatial and temporal patterns due to diffusion and oscillations, respectively, and focusing on the parameters of the intrinsic growth rate γ and the diffusion coefficient d2 of the prey. Finally, we conduct numerical simulations to validate our theoretical findings and further illustrate the dynamics of the predator–prey system, considering schooling behavior and Smith growth in prey.

Nonlinear dynamics and pattern formation in a space–time discrete diffusive intraguild predation model

In this paper, the spatiotemporal dynamics and pattern formation of a space–time discrete intraguild predation model with self-diffusion are investigated. The model is obtained by applying a coupled map lattice (CML) method. First, using linear stability analysis, the existence and stability conditions for fixed points are determined. Second, using the center manifold theorem and the bifurcation theory, the occurrence of flip, Neimark-Sacker, and Turing bifurcations are discussed. It is shown that the patterns obtained are results of Turing, flip, and Neimark-Sacker instabilities. Numerical simulations are performed to verify the theoretical analysis and to reveal complex and rich dynamics of the model, such as times series, maximal Lyapunov exponent, bifurcation diagrams, and phase portraits. Interesting patterns like spiral pattern, polygonal pattern, and the combinations of patterns of spiral waves and stripes are formed. The CML model’s results help to understand how a spatially extended, discrete intraguild predation model forms complex patterns. Notably, the continuous reaction–diffusion counterpart of the model under study is incapable of experiencing Turing instability.

Amplitude equations for wave bifurcations in reaction–diffusion systems

A wave bifurcation is the counterpart to a Turing instability in reaction–diffusion systems, but where the critical wavenumber corresponds to a pure imaginary pair rather than a zero temporal eigenvalue. Such bifurcations require at least three components and give rise to patterns that are periodic in both space and time. Depending on boundary conditions, these patterns can comprise either rotating or standing waves. Restricting to systems in one spatial dimension, complete formulae are derived for the evaluation of the coefficients of the weakly nonlinear normal form of the bifurcation up to order five, including those that determine the criticality of both rotating and standing waves. The formulae apply to arbitrary n-component systems ( n⩾3 ) and their evaluation is implemented in software which is made available as supplementary material. The theory is illustrated on two different versions of three-component reaction–diffusion models of excitable media that were previously shown to feature super- and subcritical wave instabilities and on a five-component model of two-layer chemical reaction. In each case, two-parameter bifurcation diagrams are produced to illustrate the connection between complex dispersion relations and different types of Hopf, Turing, and wave bifurcations, including the existence of several codimension-two bifurcations.

Hopf Bifurcation and Turing Instability of a Delayed Diffusive Zooplankton–Phytoplankton Model with Hunting Cooperation

In this paper, a diffusive zooplankton–phytoplankton model with time delay and hunting cooperation is established. First, the existence of all positive equilibria and their local stability are proved when the system does not include time delay and diffusion. Then, the existence of Hopf bifurcation at the positive equilibrium is proved by taking time delay as the bifurcation parameter, and the direction of Hopf bifurcation and the stability of bifurcating periodic solutions are investigated by using the center manifold theorem and the normal form theory in partial differential equations. Next, according to the theory of Turing bifurcation, the conditions for the occurrence of Turing bifurcation are obtained by taking the intraspecific competition rate of the prey as the bifurcation parameter. Furthermore, the corresponding amplitude equations are discussed by using the standard multi-scale analysis method. Finally, some numerical simulations are given to verify the theoretical results.

Patterns dynamics in a diffusive prey–predator system with ratio-dependent functional response in a planar domain

This paper is concerned with a classical two-species prey–predator reaction–diffusion system with ratio-dependent functional response and subject to homogeneous Neumann boundary condition in a two-dimensional rectangle domain. By analyzing the associated eigenvalue problem, the spatially homogeneous Hopf bifurcation curve and Turing bifurcation curve of system at the constant coexistence equilibrium are established. Then when the bifurcation parameter is in the interior of range for Turing instability and near Turing bifurcation curve, the amplitude equations of the original system near the constant coexistence equilibrium are obtained by multiple-scale time perturbation analysis. On the basis of the obtained amplitude equations, the stability and classifications of spatiotemporal patterns of the original system at the constant coexistence equilibrium are discussed. Finally, to verify the validity of the obtained theoretical results, numerical simulations are also carried out.

A two-timescale model of plankton–oxygen dynamics predicts formation of oxygen minimum zones and global anoxia

Decline of the dissolved oxygen in the ocean is a growing concern, as it may eventually lead to global anoxia, an elevated mortality of marine fauna and even a mass extinction. Deoxygenation of the ocean often results in the formation of oxygen minimum zones (OMZ): large domains where the abundance of oxygen is much lower than that in the surrounding ocean environment. Factors and processes resulting in the OMZ formation remain controversial. We consider a conceptual model of coupled plankton–oxygen dynamics that, apart from the plankton growth and the oxygen production by phytoplankton, also accounts for the difference in the timescales for phyto- and zooplankton (making it a “slow-fast system”) and for the implicit effect of upper trophic levels resulting in density dependent (nonlinear) zooplankton mortality. The model is investigated using a combination of analytical techniques and numerical simulations. The slow-fast system is decomposed into its slow and fast subsystems. The critical manifold of the slow-fast system and its stability is then studied by analyzing the bifurcation structure of the fast subsystem. We obtain the canard cycles of the slow-fast system for a range of parameter values. However, the system does not allow for persistent relaxation oscillations; instead, the blowup of the canard cycle results in plankton extinction and oxygen depletion. For the spatially explicit model, the earlier works in this direction did not take into account the density dependent mortality rate of the zooplankton, and thus could exhibit Turing pattern. However, the inclusion of the density dependent mortality into the system can lead to stationary Turing patterns. The dynamics of the system is then studied near the Turing bifurcation threshold. We further consider the effect of the self-movement of the zooplankton along with the turbulent mixing. We show that an initial non-uniform perturbation can lead to the formation of an OMZ, which then grows in size and spreads over space. For a sufficiently large timescale separation, the spread of the OMZ can result in global anoxia.

Normal form and Hopf bifurcation for the memory‐based reaction‐diffusion equation with nonlocal effect

In this paper, we provide the normal form for the Hopf bifurcation of a class of the reaction‐diffusion equation with memory‐based diffusion and nonlocal effect, where the delay is present in the differential term, similar to the chemotaxis model with time delay. The eigenvalue problems and the decomposition of the phase space are discussed in detail. Through a series of variable transformations, we obtain the third‐order truncated normal form of the model constrained on the central manifold and its equivalent equation in polar coordinates. Then, with the help of the dynamic analysis for the finite dimensional equations, the key parameters for determining the direction and stability of the Hopf bifurcation are given. These theoretical results are applied to the Bazykin's model, the stability, Turing bifurcation and Hopf bifurcation of the equilibrium are demonstrated through both theoretical and numerical methods.

Nonlinear Convective Stability of a Critical Pulled Front Undergoing a Turing Bifurcation at Its Back: A Case Study

Nonlinear Convective Stability of a Critical Pulled Front Undergoing a Turing Bifurcation at Its Back: A Case Study

Spatio-temporal complexity in a prey-predator system with Holling type-IV response and Leslie-type numerical response: Turing and steady-state bifurcations

In this study, we unravel the intricate dynamics of a spatio-temporal prey-predator model featuring a Holling type-IV functional response and Leslie-type predator numerical response under Neumann boundary conditions. Our analysis encompasses the uniform persistence and global asymptotic stability of a positive equilibrium, validated by precise numerical simulations. Additionally, we explore Turing instability and spatial pattern emergence through linear stability analysis. Our primary emphasis lies in the realm of spatio-temporal bifurcation analysis, through which we establish criteria for the presence or absence of non-constant steady states within n-dimensional diffusion models. Moreover, we discern precise conditions governing Hopf bifurcation and steady-state bifurcation in 1-dimensional diffusion models. These findings offer theoretical insights that align with the intricate dynamic patterns observed in our numerical simulations.

Bifurcations of a single species model with spatial memory environment

This paper reports on the bifurcations of a single-species model with spatial memory environment. The occurrence conditions of the Hopf bifurcation and Turing bifurcation are given with/without the spatial memory effect. Especially, we show that the positive equilibrium enjoys the stability switches with spatial memory effect.

Hybrid control of Turing instability and bifurcation for spatial-temporal propagation of computer virus

ABSTRACT In this era of information technology, information leakage and file corruption due to computer virus intrusion have been serious issues. How to detect and prevent the spread of the computer virus is the major challenge we are facing now. To target this problem, a class of virus propagation models with hybrid control scheme are formulated to investigate the dynamic evolution and prevention from a spatial-temporal perspective in this paper. Diffusion-induced Turing instability is detected in response to the computer virus propagation. The introduction of hybrid control scheme can effective suppress Turing instability and turn the propagation system back to a stable state. And then, the time delay is selected as the bifurcation parameter. If the time delay exceeds the bifurcation threshold, the propagation will be destabilised and a Hopf bifurcation will occur. The hybrid control tactic can not only regulate the occurrence of Hopf bifurcation well, but also optimise the properties of bifurcating period solutions. In the end, the correctness and validity of the theoretical results are verified via numerical simulations.

Spatiotemporal Dynamics of a General Two-Species System with Taxis Term

In this paper, we investigate the spatiotemporal dynamics in a diffusive two-species system with taxis term and general functional response, which means the directional movement of one species upward or downward the other one. The stability of positive equilibrium and the existences of Turing bifurcation, Turing–Hopf bifurcation and Turing–Turing bifurcation are investigated. An algorithm for calculating the normal form of the Turing–Hopf bifurcation induced by the taxis term and another parameter is derived. Furthermore, we apply our theoretical results to a cooperative Lotka–Volterra system and a predator–prey system with prey-taxis. For the cooperative system, stable equilibrium becomes unstable by taxis-driven Turing instability, which is impossible for the cooperative system without taxis. For a predator–prey system with prey-taxis, the dynamical classification near the Turing–Hopf bifurcation point is clearly described. Near the Turing–Hopf point, there are spatially inhomogeneous steady-state solution, spatially homogeneous/nonhomogeneous periodic solution and pattern transitions from one spatiotemporal state to another one.

Spatiotemporal patterns in a diffusive resource–consumer model with distributed memory and maturation delay

In this paper, we propose a diffusive consumer–resource model in which the consumer is involved with spatiotemporal memory and the resource has maturation delay. Firstly, for the case without maturation delay, the influence of the spatiotemporal distributed memory on the stability of the positive steady state is investigated. It has been shown that memory delay can induce Turing bifurcation for the negative memory-based diffusion coefficient and induce Hopf bifurcation for the positive memory-based diffusion coefficient. Then the joint effect of the memory delay and the maturation delay on the stability of the positive steady state is investigated and it has been shown that the joint effect of the memory delay and the maturation delay can induce more complicated spatiotemporal dynamics via Turing–Hopf bifurcation and double Hopf bifurcation. Finally, we apply our theoretical analysis results to a consumer–resource model with Holling-II type functional response to illustrate the existence of the different spatiotemporal patterns.

Complex pattern evolution of a two-dimensional space diffusion model of malware spread

In order to investigate the propagation mechanism of malware in cyber-physical systems (CPSs), the cross-diffusion in two-dimensional space is attempted to be introduced into a class of susceptible-infected (SI) malware propagation model depicted by partial differential equations (PDEs). Most of the traditional reaction-diffusion models of malware propagation only take into account the self-diffusion in one-dimensional space, but take less consideration of the cross-diffusion in two-dimensional space. This paper investigates the spatial diffusion behaviour of malware nodes spreading through physical devices. The formations of Turing patterns after homogeneous stationary instability triggered by Turing bifurcation are investigated by linear stability analysis and multiscale analysis methods. The conditions under the occurence of Hopf bifurcation and Turing bifurcation in the malware model are obtained. The amplitude equations are derived in the vicinity of the bifurcation point to explore the conditions for the formation of Turing patterns in two-dimensional space. And the corresponding patterns are obtained by varying the control parameters. It is shown that malicious virus nodes spread in different forms including hexagons, stripes and a mixture of the two. This paper will extend a new direction for the study of system security theory.