Tourism destinations are cultural heritage and spatial landscape systems of organic coexistence between humans and the Earth, and are formed through the long historical evolution of a specific geographical environment. With the development of large-scale cultural and tourism projects and the construction of scenic areas, many tourist destinations face conflicts between people and the environment and the crisis of discontinuity in the landscape and the cultural context. The concept of tourist destination personality provides a perspective for studying the interaction between humans and the environment in tourist destinations. However, existing research has not delved into the mechanisms of temporal–spatial interaction and spatial representation of regional cultural heritage in regional systems. Therefore, from the perspectives of geography and urban–rural planning, this study selects traditional villages in ancient Huizhou as the research object and employs relevant theories from cultural ecology to construct a paradigm for analyzing the formation path of tourist destination personality based on a cultural core from a regional systemic perspective. Building on this, this study develops a coupling analysis framework for the “accumulation anchoring” of heritage landscape representation in traditional villages in ancient Huizhou based on a cognitive understanding of tourist destination heritage landscape and a ‘time–space’ interactive model. The research reveals that the formation of personality in traditional villages in ancient Huizhou centers around cultural cores such as production methods, social organizations, construction mechanisms, and social beliefs. It is driven by basic forces such as resource endowment vitality, a social structure driving force, and a historical choice regulatory force, with the logic of forming dominant functions and obtaining expected benefits such as the core. The spatial representation of traditional village heritage in ancient Huizhou exhibits a synergistic evolution mechanism between ‘culture and landscape’. The research process and conclusions provide a basic framework and methodological system for the study of tourist destination personality and heritage revitalization, expanding the understanding of the process of human–environment interaction and spatial patterns in tourist destinations.