Turing Hopf Bifurcation Research Articles

The study of non-constant steady states and pattern formation for an interacting population model in a spatial environment

This manuscript accounts for an investigation of the complex dynamics of a spatial model for interacting populations. We discuss the existence and boundedness of solutions for the proposed spatio-temporal system. The global stability of the co-existing steady state of the proposed system is analyzed with the help of a suitable Lyapunov function. We provide results on the existence and non-existence of positive non-constant solutions of the model. The priori estimate for the positive steady state is obtained for the nonexistence of the non-constant positive steady state by using the maximum principle. The existence of a non-constant positive steady state is studied with the help of Leray–Schauder degree theory. The stability and Hopf bifurcation are briefly revisited for the co-existing steady state in the corresponding temporal model, where a bubble-like structure is observed. The onset of Hopf bifurcation has been analyzed, and different conditions for the formation of the Turing pattern have been established through diffusion-driven instability analysis. Numerical simulations are performed in detail to figure out the effects of saturated harvesting on Turing patterns. The Turing as well as non-Turing patterns in their respective domains are also examined. Finally, the criteria of Turing–Hopf bifurcation is briefly demonstrated with relevant numerical examples and corresponding plots that give a better illustration of this work.

Turing–Hopf Bifurcation Coinduced by Diffusion and Delay in Gierer–Meinhardt Systems

This paper investigates the impact of gene expression time delay and diffusion on the dynamic behavior of a class of Gierer–Meinhardt systems under Neumann boundary conditions. It provides necessary and sufficient conditions for the emergence of Hopf, Turing, and Turing–Hopf bifurcations. Utilizing the normal form of the Turing–Hopf bifurcation, the spatiotemporal dynamics close to the bifurcation point are categorized into six types, encompassing spatially inhomogeneous and homogeneous periodic solutions, as well as spatially homogeneous and inhomogeneous steady states, along with their transitions. Notably, it’s observed that the systems may lack stable spatially inhomogeneous periodic solutions under specific parameters, despite the emergence of Turing–Hopf bifurcation. These findings are supported by numerical simulations.

Turing–Hopf bifurcation in a diffusive predator–prey model with schooling behavior and Smith growth

This paper explores the dynamics of a diffusive predator–prey model, considering schooling behavior and Smith growth in prey. Initially, we have formulated the pertinent characteristic equations. Subsequently, We proceed to examine the existence of the Turing bifurcation and Hopf bifurcation, phenomena that describe the emergence of spatial and temporal patterns due to diffusion and oscillations, respectively, and focusing on the parameters of the intrinsic growth rate γ and the diffusion coefficient d2 of the prey. Finally, we conduct numerical simulations to validate our theoretical findings and further illustrate the dynamics of the predator–prey system, considering schooling behavior and Smith growth in prey.

Turing-Hopf bifurcation analysis and normal form in delayed diffusive predator–prey system with taxis and fear effect

Turing-Hopf bifurcation analysis and normal form in delayed diffusive predator–prey system with taxis and fear effect

Evolution of Cooperation in Spatio-Temporal Evolutionary Games with Public Goods Feedback.

In biology, evolutionary game-theoretical models often arise in which players' strategies impact the state of the environment, driving feedback between strategy and the surroundings. In this case, cooperative interactions can be applied to studying ecological systems, animal or microorganism populations, and cells producing or actively extracting a growth resource from their environment. We consider the framework of eco-evolutionary game theory with replicator dynamics and growth-limiting public goods extracted by population members from some external source. It is known that the two sub-populations of cooperators and defectors can develop spatio-temporal patterns that enable long-term coexistence in the shared environment. To investigate this phenomenon and unveil the mechanisms that sustain cooperation, we analyze two eco-evolutionary models: a well-mixed environment and a heterogeneous model with spatial diffusion. In the latter, we integrate spatial diffusion into replicator dynamics. Our findings reveal rich strategy dynamics, including bistability and bifurcations, in the temporal system and spatial stability, as well as Turing instability, Turing-Hopf bifurcations, and chaos in the diffusion system. The results indicate that effective mechanisms to promote cooperation include increasing the player density, decreasing the relative timescale, controlling the density of initial cooperators, improving the diffusion rate of the public goods, lowering the diffusion rate of the cooperators, and enhancing the payoffs to the cooperators. We provide the conditions for the existence, stability, and occurrence of bifurcations in both systems. Our analysis can be applied to dynamic phenomena in fields as diverse as human decision-making, microorganism growth factors secretion, and group hunting.

Effect of intestinal permeability and phagocytes diffusion rate on pattern structure of Crohn’s disease based on the Turing–Hopf bifurcation

Effect of intestinal permeability and phagocytes diffusion rate on pattern structure of Crohn’s disease based on the Turing–Hopf bifurcation

Spatiotemporal Dynamics of a General Two-Species System with Taxis Term

In this paper, we investigate the spatiotemporal dynamics in a diffusive two-species system with taxis term and general functional response, which means the directional movement of one species upward or downward the other one. The stability of positive equilibrium and the existences of Turing bifurcation, Turing–Hopf bifurcation and Turing–Turing bifurcation are investigated. An algorithm for calculating the normal form of the Turing–Hopf bifurcation induced by the taxis term and another parameter is derived. Furthermore, we apply our theoretical results to a cooperative Lotka–Volterra system and a predator–prey system with prey-taxis. For the cooperative system, stable equilibrium becomes unstable by taxis-driven Turing instability, which is impossible for the cooperative system without taxis. For a predator–prey system with prey-taxis, the dynamical classification near the Turing–Hopf bifurcation point is clearly described. Near the Turing–Hopf point, there are spatially inhomogeneous steady-state solution, spatially homogeneous/nonhomogeneous periodic solution and pattern transitions from one spatiotemporal state to another one.

Spatial Dynamics of a Competitive and Cooperative Model with Multiple Delay Effects: Turing Patterns and Hopf Bifurcation

Competing populations within an ecosystem often release chemicals during the interactions and diffusion processes. These chemicals can have diverse effects on competitors, ranging from inhibition to stimulation of species’ growth. This work constructs a competition model that incorporates stimulatory substances, spatial effects, and multiple time lags to investigate the combined impact of these phenomena on competitors’ growth. When the stimulation rate from the produced chemicals falls within a suitable threshold interval, all species within the system can coexist. Under the species’ coexistence, their diffusive phenomenon leads to a spatially heterogeneous distribution, resulting in patchy structures (Turing patterns) within their habitat. As the parameter values exceed their thresholds, species begin to exhibit spatially periodic oscillations (spatial Hopf bifurcation). The presence of multiple delays and competitors’ diffusion contributes to spatially complex and heterogeneous behaviors (Turing–Hopf bifurcation). The results help us understand the underlying mechanisms behind these heterogeneous behaviors and enable us to mitigate their negative impact on species’ growth and harvest. Numerical simulations are used to measure the dynamics of competitors under different parameter conditions.

Spatiotemporal patterns in a diffusive resource–consumer model with distributed memory and maturation delay

In this paper, we propose a diffusive consumer–resource model in which the consumer is involved with spatiotemporal memory and the resource has maturation delay. Firstly, for the case without maturation delay, the influence of the spatiotemporal distributed memory on the stability of the positive steady state is investigated. It has been shown that memory delay can induce Turing bifurcation for the negative memory-based diffusion coefficient and induce Hopf bifurcation for the positive memory-based diffusion coefficient. Then the joint effect of the memory delay and the maturation delay on the stability of the positive steady state is investigated and it has been shown that the joint effect of the memory delay and the maturation delay can induce more complicated spatiotemporal dynamics via Turing–Hopf bifurcation and double Hopf bifurcation. Finally, we apply our theoretical analysis results to a consumer–resource model with Holling-II type functional response to illustrate the existence of the different spatiotemporal patterns.

Dynamics of neural fields with exponential temporal kernel.

We consider the standard neural field equation with an exponential temporal kernel. We analyze the time-independent (static) and time-dependent (dynamic) bifurcations of the equilibrium solution and the emerging spatiotemporal wave patterns. We show that an exponential temporal kernel does not allow static bifurcations such as saddle-node, pitchfork, and in particular, static Turing bifurcations. However, the exponential temporal kernel possesses the important property that it takes into account the finite memory of past activities of neurons, which Green's function does not. Through a dynamic bifurcation analysis, we give explicit bifurcation conditions. Hopf bifurcations lead to temporally non-constant, but spatially constant solutions, but Turing-Hopf bifurcations generate spatially and temporally non-constant solutions, in particular, traveling waves. Bifurcation parameters are the coefficient of the exponential temporal kernel, the transmission speed of neural signals, the time delay rate of synapses, and the ratio of excitatory to inhibitory synaptic weights.

Bifurcations and pattern formation in a host–parasitoid model with nonlocal effect

In this paper, we analyse Turing instability and bifurcations in a host–parasitoid model with nonlocal effect. For a ordinary differential equation model, we provide some preliminary analysis on Hopf bifurcation. For a reaction–diffusion model with local intraspecific prey competition, we first explore the Turing instability of spatially homogeneous steady states. Next, we show that the model can undergo Hopf bifurcation and Turing–Hopf bifurcation, and find that a pair of spatially nonhomogeneous periodic solutions is stable for a (8,0)-mode Turing–Hopf bifurcation and unstable for a (3,0)-mode Turing–Hopf bifurcation. For a reaction–diffusion model with nonlocal intraspecific prey competition, we study the existence of the Hopf bifurcation, double-Hopf bifurcation, Turing bifurcation, and Turing–Hopf bifurcation successively, and find that a spatially nonhomogeneous quasi-periodic solution is unstable for a (0,1)-mode double-Hopf bifurcation. Our results indicate that the model exhibits complex pattern formations, including transient states, monostability, bistability, and tristability. Finally, numerical simulations are provided to illustrate complex dynamics and verify our theoretical results.

Spatiotemporal dynamics of a diffusive nutrient-phytoplankton model with delayed nutrient recycling

In this paper, we investigate the spatiotemporal dynamics of a diffusive nutrient-phytoplankton model with delayed nutrient recycling. We first study the stability of positive equilibrium and Turing instability induced by diffusion. We then investigate the effect of delay, and it turns out that the value of the rate of recycling k plays an important role in the Hopf bifurcation induced by delay. The delay will and will not induce Hopf bifurcation with low and high level of k, respectively. To reveal the spatiotemporal dynamics, Turing–Hopf bifurcation is carried out, and normal form is derived. Many spatiotemporal dynamics are found, including the coexistence of two stable spatially inhomogeneous periodic solutions or two stable spatially inhomogeneous steadystate solutions.

The role of natural recovery category in malaria dynamics under saturated treatment.

In the process of malaria transmission, natural recovery individuals are slightly infectious compared with infected individuals. Our concern is whether the infectivity of natural recovery category can be ignored in areas with limited medical resources, so as to reveal the epidemic pattern of malaria with simpler analysis. To achieve this, we incorporate saturated treatment into two-compartment and three-compartment models, and the infectivity of natural recovery category is only reflected in the latter. The non-spatial two-compartment model can admit backward bifurcation. Its spatial version does not possess rich dynamics. Besides, the non-spatial three-compartment model can undergo backward bifurcation, Hopf bifurcation and Bogdanov-Takens bifurcation. For spatial three-compartment model, due to the complexity of characteristic equation, we apply Shengjin's Distinguishing Means to realize stability analysis. Further, the model exhibits Turing instability, Hopf bifurcation and Turing-Hopf bifurcation. This makes the model may admit bistability or even tristability when its basic reproduction number less than one. Biologically, malaria may present a variety of epidemic trends, such as elimination or inhomogeneous distribution in space and periodic fluctuation in time of infectious populations. Notably, parameter regions are given to illustrate substitution effect of two-compartment model for three-compartment model in both scenarios without or with spatial movement. Finally, spatial three-compartment model is used to present malaria transmission in Burundi. The application of efficiency index enables us to determine the most effective method to control the number of cases in different scenarios.

Spatiotemporal patterns induced by Turing-Hopf interaction and symmetry on a disk.

Turing bifurcation and Hopf bifurcation are two important kinds of transitions giving birth to inhomogeneous solutions, in spatial or temporal ways. On a disk, these two bifurcations may lead to equivariant Turing-Hopf bifurcations whose normal forms are given in three different cases in this paper. In addition, we analyzed the possible solutions for each normal form, which can guide us to find solutions with physical significance in real-world systems, and the breathing, standing wave-like, and rotating wave-like patterns are found in a delayed mussel-algae model.

A high accuracy compact difference scheme and numerical simulation for a type of diffusive plant-water model in an arid flat environment

<abstract><p>In this paper, we investigate the numerical computation method for a one-dimensional self diffusion plant water model with homogeneous Neumann boundary conditions. First, a high accuracy compact difference scheme for the diffusive plant water model in an arid flat environment is constructed using the finite difference method. The fourth order compact difference scheme is used for the spatial derivative term, and the Taylor series expansion and residual correction function are used to discretize the time term. We obtain a difference scheme with second-order accuracy in time and fourth-order accuracy in space. Second, the Fourier analysis method is used to prove that the above format is unconditionally stable. Then, the numerical examples provided the convergence and accuracy of the difference scheme. Finally, numerical simulations are conducted near the Turing Hopf bifurcation point of the model to obtain the spatial distribution maps of vegetation and water under small disturbances of different parameters. In this paper, the evolution law of vegetation quantity and water density at any time is observed.Revealing the impact of small changes in parameters on the spatiotemporal dynamics of plant water models will provide a basis for understanding whether ecosystems are fragile.</p></abstract>

Bifurcations in a General Delay Sel’kov–Schnakenberg Reaction–Diffusion System

The dynamics of a delay Sel’kov–Schnakenberg reaction–diffusion system are explored. The existence and the occurrence conditions of the Turing and the Hopf bifurcations of the system are found by taking the diffusion coefficient and the time delay as the bifurcation parameters. Based on that, the existence of codimension-2 bifurcations including Turing–Turing, Hopf–Hopf and Turing–Hopf bifurcations are given. Using the center manifold theory and the normal form method, the universal unfolding of the Turing–Hopf bifurcation at the positive constant steady-state is demonstrated. According to the universal unfolding, a Turing–Hopf bifurcation diagram is shown under a set of specific parameters. Furthermore, in different parameter regions, we find the existence of the spatially inhomogeneous steady-state, the spatially homogeneous and inhomogeneous periodic solutions. Discretization of time and space visualizes these spatio-temporal solutions. In particular, near the critical point of Hopf–Hopf bifurcation, the spatially homogeneous periodic and inhomogeneous quasi-periodic solutions are found numerically.

Turing-Hopf Bifurcation Analysis in a Diffusive Ratio-Dependent Predator-Prey Model with Allee Effect and Predator Harvesting.

This paper investigates the complex dynamics of a ratio-dependent predator-prey model incorporating the Allee effect in prey and predator harvesting. To explore the joint effect of the harvesting effort and diffusion on the dynamics of the system, we perform the following analyses: (a) The stability of non-negative constant steady states; (b) The sufficient conditions for the occurrence of a Hopf bifurcation, Turing bifurcation, and Turing-Hopf bifurcation; (c) The derivation of the normal form near the Turing-Hopf singularity. Moreover, we provide numerical simulations to illustrate the theoretical results. The results demonstrate that the small change in harvesting effort and the ratio of the diffusion coefficients will destabilize the constant steady states and lead to the complex spatiotemporal behaviors, including homogeneous and inhomogeneous periodic solutions and nonconstant steady states. Moreover, the numerical simulations coincide with our theoretical results.

Turing-Hopf bifurcation and bi-stable spatiotemporal periodic orbits in a delayed predator-prey model with predator-taxis

When Turing-Hopf bifurcation occurs, it is naturally worth discussing the types of spatiotemporal patterns and their stabilities. In this paper, for partial functional differential equations with nonlinear diffusion, we establish the explicit formulae for the coefficients of normal form associated with Turing-Hopf bifurcation. This provides an explicit connection between the original system and the third-order ordinary differential equations (i.e. normal form) restricted on center manifold. Subsequently, with the aid of extended formulae, we discuss a delayed predator-prey model with predator-taxis and find that Turing-Hopf bifurcation leads to the emergence of a pair of stable single-peak spatially inhomogeneous steady states, a pair of stable multi-peak spatially inhomogeneous periodic orbits and so on. Also, the parameter regions in which these phenomena arise are given quantitatively. It is further shown that the pair of bistable spatially inhomogeneous periodic orbits arise due to the interaction of predator-taxis and time delay, which won't emerge in the same system without predator-taxis.

Spatiotemporal Dynamics of a Diffusive Immunosuppressive Infection Model with Nonlocal Competition and Crowley–Martin Functional Response

In this paper, we introduce the Crowley–Martin functional response and nonlocal competition into a reaction–diffusion immunosuppressive infection model. First, we analyze the existence and stability of the positive constant steady states of the systems with nonlocal competition and local competition, respectively. Second, we deduce the conditions for the occurrence of Turing, Hopf, and Turing–Hopf bifurcations of the system with nonlocal competition, as well as the conditions for the occurrence of Hopf bifurcations of the system with local competition. Furthermore, we employ the multiple time scales method to derive the normal forms of the Hopf bifurcations reduced on the center manifold for both systems. Finally, we conduct numerical simulations for both systems under the same parameter settings, compare the impact of nonlocal competition, and find that the nonlocal term can induce spatially inhomogeneous stable periodic solutions. We also provide corresponding biological explanations for the simulation results.

Stability and spatiotemporal patterns of a memory-based diffusion equation with nonlocal interaction

In this paper, we investigate the spatiotemporal dynamics driven by the memory delay and nonlocal interaction for the memory-based diffusion equation, where the nonlocal interaction is characterized by the given Green function. It has been shown that this kind of nonlocal interaction can lead to inhomogeneous steady states with any modes, and the joint effect of nonlocal interaction and memory delay can lead spatially inhomogeneous Hopf bifurcation and Turing–Hopf bifurcation.