AbstractThe impact of assimilating in situ temperature, moisture and wind observations from a WindBorne Systems balloon, with long duration and adjustable ballast, is evaluated using a case study from the 2022 THINICE field campaign. A case is selected wherein the WindBorne balloon directly sampled a jet streak associated with a tropopause polar vortex (TPV). The observed TPV merged with another TPV at the same time as a downstream Arctic cyclone (AC) redeveloped eastward. The case is used to better understand the role of the observed TPV in the evolution of the downstream AC. The assimilation of the WindBorne observations improves the forecast track and amplitude of the TPV during the ∼1 day forecast period that the TPV can be tracked as a distinct feature. The root mean square error of temperature at 350 hPa is improved by the WindBorne assimilation throughout the 2.5 day forecast period. The surface cyclone track forecast is improved by the WindBorne assimilation during the period of eastward redevelopment of the surface cyclone, and the sea level pressure RMSE in the surrounding region is reduced during the first ∼1.5 days of forecast lead time. Results demonstrate the capability of the long‐duration controllable WindBorne balloons to improve the analysis of the TPV associated with the jet streak, leading to improved forecast of both tropopause‐level and surface pressure features. Additionally, the importance of observing the upstream TPV amplitude for improving forecasts of the process of TPV merging and its impact on the evolution and longevity of the mature AC is confirmed.