Presence Of Elephants Research Articles

Effects of illegal grazing and invasive Lantana camara on Asian elephant habitat use

Protected areas provide some of the last refuges for Asian elephants in the wild. Managing these areas for elephants will be critical for elephant conservation. Scientists know little about elephant habitat use in Asia and how invasive species or livestock grazing influence habitat use. We studied these issues in two protected areas in Sri Lanka, Udawalawe National Park and Hurulu Eco-Park. These areas contain some of Sri Lanka's largest remaining grasslands. These grasslands are threatened by the invasive and toxic shrub, Lantana camara, and are used for illegal livestock grazing. To measure habitat use by elephants and livestock, we conducted dung surveys along over 50 km of transects stratified across grassland, scrub, and forest. We surveyed 159 vegetation plots along these transects to assess plant composition, and mapped habitat types based on satellite images. We used mixed-effect models to determine the relative importance of habitats, livestock presence, and plant associations for elephant use. Elephant presence was greatest in scrub and grassland habitats, positively associated with both livestock presence and short graminoids, and unaffected by L. camara, which was widespread but at low densities. Given the importance of these areas to elephants, we recommend a precautionary management approach that focuses on curbing both illegal grazing and the spread of L. camara.

Lethal Fence Electrocution: A Major Threat to Asian Elephants in Assam, India

India has the largest population of Asian elephants ( Elephas maximus) worldwide. Habitat fragmentation and loss of habitat have diminished food resources, and wild elephants have resorted to raiding crops grown within or adjacent to their home range. Elephants are often deliberately electrocuted for foraying into human-used areas, and this is a key reason for elephant mortalities in India. We collated data on elephant mortalities for a 13-year period (2003–2016) from the Forest Department records. We conducted surveys across Sonitpur District (East and West Forest Division), Assam, where electric fences are installed and documented their location, properties, and elephant presence. Overall, 138 elephants died between 2003 and 2016 due to retaliation, electrocution, accidental or natural death, and unknown reasons. We recorded 47 electric fences (27 lethal and 20 nonlethal) of which 49% were situated within notified forest boundaries. Most lethal fences (63%) protected agriculture fields and were seasonal installations, whereas nonlethal fences protected settlements and forest edges (25% each) and were permanent. Individuals controlled 52% of all lethal fences, while nonlethal fences were primarily controlled by the communities (50%). Most lethal fences (83%) were less than 1 km, whereas 80% of nonlethal fences were over 1 km. Elephant presence was seasonal in 56% of lethal fence locations and year-round in 85% of nonlethal fence locations. We postulate habitat loss and encroachment as two key drivers of fence installations. We recommend rehabilitation of encroachers, monitoring of areas where electricity is tapped illegally, sensitization of local communities, and involving multiple stakeholders to help reduce elephant mortalities because of electrocution.

About the Cover

About the Cover Catherine Raymond Click for larger view View full resolution Front Cover. Cover design: Sophia Varcados, Northern Illinois University Cover: Port et bourg de Mergui (Port and Town of Mergui), Jacques Nicolas Bellin (1703– 1772), Map, 1764, 22x18 cm. Burma Art Collection, Rare Books and Special Collections at Northern Illinois University, Founders Memorial Library. G7724.M41764.B45 This black and white image was originally a rendering of a beautiful hand colored engraved maritime chart of the port and city of Mergui (now Myeik), located in Southern Myanmar in Tanintharyi Region. This chart, drawn by Jacques Nicolas Bellin (1703–1772), was initially a component of Le Petit Atlas Maritime: Recueil de Cartes et Plans des Quatre Parties du Monde en Cinq Volumes, published in Paris in 1764. Bellin was a renowned cartographer and hydrographer. He trained at a very young age at the famous Dépôt des Cartes et Plans de la Marine (also known as Dépôt de la Marine, where navigational charts were made and kept in France under royal patronage). He produced numerous maritime atlases; especially the Petit Atlas Maritime, which comprised 580 individual prints. He was well-known for taking extreme care to render the finest details with the highest accuracy. Much of his work was more widely distributed and reproduced in numerous other atlases and travelogues, such as Histoire Générale des Voyages d’Antoine Prévost, which assembled numerous small format maps and charts, and was published in fifteen volumes. Myeik (Mergui) is located in Southern Myanmar on the Bay of Bengal. Its port is well-protected by Kadan Island. Ancient Ptolemaic maps indicated Regnum Mursuli or Mergui (Tenasserim). It was visited in the 1420s by Nicolo de Conti, who, while entering the river called Tenaserin, noted the presence of numerous elephants (Suarez 1999: 122). But, the oldest description of the actual city of Mergui, where “. . . one can find silk-making, cockfights, Brazilwood and benzoin,” is attributed to Varthema. In the first decade of the sixteenth century, he elaborated, “(t)he city of Tarnassari is situated near the sea. It is a level place and well-watered, and has a good port that is a river on the side toward the north . . . The houses of this city are well-surrounded by walls (Suarez 1999: 123).” Mergui later became an important port for European traders, particularly for the Portuguese, as they could source merchandise from Pegu and Siam, including wine stored in the famous Martaban jars. From there, such products could be exported to India. By the 18th century, when this chart was originally surveyed, France and England were in competition. Mergui was considered by the British officer Alexander Hamilton as being an area “infested by the pirates,” hence not as desirable an entrepôt as Rangoon. That explains—as indicated here—the presence of a French fort and also places where the French could build boats and careen their ships. As a French settlement, it did not last long. In 1824-26, under King Bagyidaw, the first Anglo-Burmese war ensued and Tenasserim, along with Arakan, were annexed by the British. Today, centuries later, the recent interest of Mergui Archipelago is moving toward a completely new development. It is appropriate to consider it in our ratiocinations of “Progress from Whom? And for What?” Catherine Raymond curator of the Burma Art Collection at Northern Illinois University Reference Suarez, Thomas. Early Mapping of Southeast Asia. Indonesia: Periplus Editions, 1999. Copyright © 2018 Center for Burma Studies, Northern Illinois University

Applied Geographical Information System (GIS): Overlay analysis

Technological limitations have posed challenges in integrating wildlife conservation needs with human land uses and other infrastructural development needs. The advent of the Global Position System (GPS) technology has made it possible to collect elephant location and human land use data. Geographic Information System (GIS) software has allowed overlay analysis of elephant location data and human land uses. This study, therefore demonstrates how the overlay analysis function in GIS can be employed to link human land use data layers and elephant location. To achieve this, the overlay analysis function was used to combine elephant location data with distance maps of each human land use factor. The distance values for each human land use factor were extracted at each elephant location using the map value function in ILWIS. The resultant table show extracted elephant location values and corresponding distances values for each human land use factor. Further research can be conducted by exporting elephant location values and corresponding land use values to SPSS to predict the human land uses which significantly influence the presence or absence of elephants. This study also recommends the use of the overlay analysis tool in Environmental Impact Assessment projects to model the most suitable site for a proposed project such as dam construction.

Environmental factors affecting the distribution of African elephants in the Kasigau wildlife corridor, SE Kenya

AbstractThe African elephant, Loxodonta africana, is under threat from habitat loss, poaching and human–elephant conflict. To mitigate for impact of habitat loss and reduce conflict, connectivity between elephant habitats can be improved through the protection of corridor areas. This study looks at elephant distribution and movement patterns within the Kasigau Wildlife Corridor (KWC) within the Tsavo Conservation Area in South‐east Kenya. Elephant presence data were obtained from observations by rangers during routine patrols across KWC, and were analysed in MaxEnt. The environmental factors predicting elephant distribution and density were tested, as well as the relationship between elephant maximum entropy and the presence and abundance of other wildlife. Seasonal variations in temperature and precipitation, plus presence of waterholes were found to play significant roles in elephant distribution across KWC. Higher elephant densities were not found to correlate with lower densities of other wildlife species; indeed, during the dry seasons, elephant presence was associated with greater wild herbivore densities. Besides illustrating the importance of the KWC for elephant conservation in the Tsavo ecosystem, both as a key corridor and habitat, this study also hopes to highlight the untapped utility of routine ranger patrol data, and encourage the use of such presence‐only data for deducing important knowledge for conservation of biodiversity.

Predicting the factors which significantly predict the presence of elephants

Predicting the factors which significantly predict the presence of elephants

Patch-occupancy survey of elephant (<i>Loxodonta africana</i>) surrounding Livingstone, Zambia

Wild elephants represent the biggest human–wildlife conflict issue in Livingstone, Zambia. However, little is known about their movements. This survey investigated elephants’ habitat use outside a core protected and fenced zone that forms part of Mosi-oa-Tunya National Park, Zambia. Using ‘patch-occupancy’ methodology, indications of elephant presence (feeding behaviour, dung and tracks) were surveyed. The survey aimed to assist proposed future monitoring exercises by defining the geographical extent that should be considered to improve accuracy in species abundance estimates. Results were supplemented using collected indications of elephant presence from prior monitoring exercises, and during this survey. Elephant presence was confirmed up to 8 km from the boundary of the protected core habitat, focussed in: (1) an unfenced zone of the national park, (2) along a road leading from the national park to the Dambwa Forest to the north and (3) along two rivers located to the west (Sinde River) and east (Maramba River) of the core area. Detection probability of elephant presence was high using these methods, and we recommend regular sampling to determine changes in habitat use by elephants, as humans continue to modify land-use patterns.Conservation implications: Identification of elephant ranging behaviour up to 8 km outside of the Mosi-oa-Tunya National Park in southern Zambia will assist in managing human– elephant conflict in the area, as well as in assessing this seasonal population’s abundance.

Humans and elephants as treefall drivers in African savannas

Humans have played a major role in altering savanna structure and function, and growing land‐use pressure will only increase their influence on woody cover. Yet humans are often overlooked as ecological components. Both humans and the African elephant Loxodonta africana alter woody vegetation in savannas through removal of large trees and activities that may increase shrub cover. Interactive effects of both humans and elephants with fire may also alter vegetation structure and composition. Here we capitalize on a macroscale experimental opportunity – brought about by the juxtaposition of an elephant‐mediated landscape, human‐utilized communal harvesting lands and a nature reserve fenced off from both humans and elephants – to investigate the influence of humans and elephants on height‐specific treefall dynamics. We surveyed 6812 ha using repeat, airborne high resolution Light Detection and Ranging (LiDAR) to track the fate of 453 685 tree canopies over two years. Human‐mediated biennial treefall rates were 2–3.5 fold higher than the background treefall rate of 1.5% treefall ha–1, while elephant‐mediated treefall rates were 5 times higher at 7.6% treefall ha–1 than the control site. Model predictors of treefall revealed that human or elephant presence was the most important variable, followed by the interaction between geology and fire frequency. Treefall patterns were spatially heterogeneous with elephant‐driven treefall associated with geology and surface water, while human patterns were related to perceived ease of access to wood harvesting areas and settlement expansion. Our results show humans and elephants utilize all height classes of woody vegetation, and that large tree shortages in a heavily utilized communal land has transferred treefall occurrence to shorter vegetation. Elephant‐ and human‐dominated landscapes are tied to interactive effects that may hinder tree seedling survival which, combined with tree loss in the landscape, may compromise woodland sustainability.

Elephants in the understory: opposing direct and indirect effects of consumption and ecosystem engineering by megaherbivores.

Positive indirect effects of consumers on their resources can stabilize food webs by preventing overexploitation, but the coupling of trophic and non-trophic interactions remains poorly integrated into our understanding of community dynamics. Elephants engineer African savanna ecosystems by toppling trees and breaking branches, and although their negative effects on trees are well documented, their effects on small-statured plants remain poorly understood. Using data on 117 understory plant taxa collected over 7 yr within 36 1-ha experimental plots in a semi-arid Kenyan savanna, we measured the strength and direction of elephant impacts on understory vegetation. We found that elephants had neutral effects on most (83-89%) species, with a similar frequency of positive and negative responses among the remainder. Overall, estimated understory biomass was 5-14% greater in the presence of elephants across a range of rainfall levels. Whereas direct consumption likely accounts for the negative effects, positive effects are presumably indirect. We hypothesized that elephants create associational refuges for understory plants by damaging tree canopies in ways that physically inhibit feeding by other large herbivores. As predicted, understory biomass and species richness beneath elephant-damaged trees were 55% and 21% greater, respectively, than under undamaged trees. Experimentally simulated elephant damage increased understory biomass by 37% and species richness by 49% after 1yr. Conversely, experimentally removing elephant damaged branches decreased understory biomass by 39% and richness by 30% relative to sham-manipulated trees. Camera-trap surveys revealed that elephant damage reduced the frequency of herbivory by 71%, whereas we detected no significant effect of damage on temperature, light, or soil moisture. We conclude that elephants locally facilitate understory plants by creating refuges from herbivory, which countervails the direct negative effects of consumption and enhances larger-scale biomass and diversity by promoting the persistence of rare and palatable species. Our results offer a counterpoint to concerns about the deleterious impacts of elephant "overpopulation" that should be considered in debates over wildlife management in African protected areas: understory species comprise the bulk of savanna plant biodiversity, and their responses to elephants are buffered by the interplay of opposing consumptive and non-consumptive interactions.

Drivers of Change in Myanmar’s Wild Elephant Distribution

Myanmar is considered as a stronghold for wild elephants, though past estimates varied widely from 3,000 to 10,000. Results of a 2004 expert workshop showed estimates between 1,430 and 2,065. Building on surveys from 1990 to 1992, we conducted new expert interviews in townships throughout the range, with questions focusing on numbers of elephants living in townships and threats to and from elephants. We used general linear models to analyze characteristics of townships with and without elephants and to understand factors influencing changes in elephant presence. Our results show a major decrease in the geographic distribution of wild elephants in Myanmar between 1992 and 2006 with deforestation being the major driver. While forest cover is important for elephant presence, continuity with other elephant populations had a stonger influence on elephant persistence between surveys. Fragmentation of elephant populations is also a major driver of decline. Increases in forest cover increased elephant presence, while increases in edge and human population had the reverse effect. Deforestation and fragmentation lead to increased human–elephant conflict in some areas, sometimes concluding with the capture of elephants for timber operations and further draining wild populations. A national elephant action plan that includes monitoring of elephant status and threats is urgently needed and critical for Myanmar’s wild elephants, particularly as more than 50 years of political isolation are rapidly ending and pressure on the country’s natural resources is increasing exponentially.

No evidence for an elephant-termite feedback loop in Sand Forest, South Africa

Termites and mammalian herbivores might derive mutual benefit from each other through positive feedback loops, but empirical evidence is lacking. One suggested positive feedback loop is between termites and elephant, both ecosystem engineers. Termites, as decomposer organisms, contribute to nutrient cycling and soil enrichment, creating hotspots with increased forage quality. Elephant are known to select these high quality vegetation patches, depositing woody debris when foraging, but it is unknown whether elephant presence might affect termites, and if a feedback loop between these two taxa exists. We tested this hypothesis by measuring termite diversity and activity over three months in the summer wet season, inside and outside a long-term (eight years) elephant exclosure experiment in Sand Forest at Phinda Private Game Reserve, South Africa. Termites were sampled using cellulose baits and woody debris was quantified using line transects. Termite activity was not affected by eight years of elephant absence, despite a greater volume of woody debris where elephant were present. Termite assemblages were similarly unaffected by elephant absence. Apart from Schedorhinotermes, all other termite genera were sampled in both treatments. Therefore, the postulated positive feedback loop between termites and elephant is not supported in Sand Forest. However, this does not contradict findings that areas with high termite activity positively influence herbivore foraging, only that elephant foraging does not affect termite activity. This suggests that elephant disturbance (and elephant management policies) has little to no effect on termites and the fundamental roles they play in ecosystems over at least the short-term.

Predicting Hotspots of Human-Elephant Conflict to Inform Mitigation Strategies in Xishuangbanna, Southwest China.

Research on the spatial patterns of human-wildlife conflict is fundamental to understanding the mechanisms underlying it and to identifying opportunities for mitigation. In the state of Xishuangbanna, containing China’s largest tropical forest, an imbalance between nature conservation and economic development has led to increasing conflicts between humans and Asian elephants (Elephas maximus), as both elephant numbers and conversion of habitable land to rubber plantations have increased over the last several decades. We analyzed government data on the compensation costs of elephant-caused damage in Xishuangbanna between 2008 and 2012 to understand the spatial and temporal patterns of conflict, in terms of their occurrence, frequency and distribution. More than 18,261 incidents were reported, including episodes involving damage to rubber trees (n = 10,999), damage to crops such as paddy, upland rice, corn, bananas and sugarcane (n = 11,020), property loss (n = 689) and attacks on humans (n = 19). The conflict data reconfirmed the presence of elephants in areas which have lacked records since the late 1990s. Zero Altered Negative Binomial models revealed that the risk of damage to crops and plantations increased with proximity to protected areas, increasing distance from roads, and lower settlement density. The patterns were constant across seasons and types of crop damaged. Damage to rubber trees was essentially incidental as elephants searched for crops to eat. A predictive map of risks revealed hotspots of conflict within and around protected areas, the last refuges for elephants in the region, and along habitat corridors connecting them. Additionally, we analyzed how mitigation efforts can best diminish the risk of conflict while minimizing financial costs and adverse biological impacts. Our analytical approach can be adopted, adjusted and expanded to other areas with historical records of human-wildlife conflict.

Kamala tree as an indicator of the presence of Asian elephants during the dry season in the Shivalik landscape of northwestern India

The availability of forage resources during the dry season is often a critical factor in determining the distribution and movement of large herbivores. It has long been suspected that the Kamala tree (Mallotus philippinensis) can serve as an indicator of the distribution for Asian elephants during the dry season in northwestern India. However, there is little direct evidence in support of this speculation, especially at a large landscape scale. Here, we predicted the distribution of Kamala trees in the Shivalik landscape of northwestern India based on topographic and bioclimatic variables, as well as satellite-derived vegetation indices and forest canopy height data using a presence-only ecological niche model. We used the area under the receiver operating characteristic curve (AUC) and true skill statistic (TSS) to validate the model. We then examined the relationship between the occurrence probability of Kamala trees and the presence of Asian elephants with data collected during dry seasons between 2010 and 2014 using logistic regression models. Our results showed that the probability of occurrence of Kamala trees was predicted with good accuracy (AUC=0.88 and TSS=0.51). The logistic regression models showed that the presence of Asian elephants can be adequately predicted by the occurrence probability of Kamala trees. This result suggests that the distribution of Kamala trees is a good indicator of the presence of Asian elephants during the dry season in the Shivalik landscape. These findings may have major implications for the conservation of Asian elephants, especially in designing wildlife corridors and mitigating human-elephant conflicts.

Taking the elephant out of the room and into the corridor: can urban corridors work?

AbstractTransfrontier wildlife corridors can be successful conservation tools, connecting protected areas and reducing the impact of habitat fragmentation on mobile species. Urban wildlife corridors have been proposed as a potential mitigation tool to facilitate the passage of elephants through towns without causing conflict with urban communities. However, because such corridors are typically narrow and close to human development, wildlife (particularly large mammals) may be less likely to use them. We used remote-sensor camera traps and global positioning system collars to identify the movement patterns of African elephants Loxondonta africana through narrow, urban corridors in Botswana. The corridors were in three types of human-dominated land-use designations with varying levels of human activity: agricultural, industrial and open-space recreational land. We found that elephants used the corridors within all three land-use designations and we identified, using a model selection approach, that season, time of day and rainfall were important factors in determining the presence of elephants in the corridors. Elephants moved more slowly through the narrow corridors compared with their movement patterns through broader, wide-ranging corridors. Our results indicate that urban wildlife corridors are useful for facilitating elephants to pass through urban areas.

Patterns and Determinants of Habitat Occupancy by the Asian Elephant in the Western Ghats of Karnataka, India.

Understanding species distribution patterns has direct ramifications for the conservation of endangered species, such as the Asian elephant Elephas maximus. However, reliable assessment of elephant distribution is handicapped by factors such as the large spatial scales of field studies, survey expertise required, the paucity of analytical approaches that explicitly account for confounding observation processes such as imperfect and variable detectability, unequal sampling probability and spatial dependence among animal detections. We addressed these problems by carrying out ‘detection—non-detection’ surveys of elephant signs across a c. 38,000-km2 landscape in the Western Ghats of Karnataka, India. We analyzed the resulting sign encounter data using a recently developed modeling approach that explicitly addresses variable detectability across space and spatially dependent non-closure of occupancy, across sampling replicates. We estimated overall occupancy, a parameter useful to monitoring elephant populations, and examined key ecological and anthropogenic drivers of elephant presence. Our results showed elephants occupied 13,483 km2 (SE = 847 km2) corresponding to 64% of the available 21,167 km2 of elephant habitat in the study landscape, a useful baseline to monitor future changes. Replicate-level detection probability ranged between 0.56 and 0.88, and ignoring it would have underestimated elephant distribution by 2116 km2 or 16%. We found that anthropogenic factors predominated over natural habitat attributes in determining elephant occupancy, underscoring the conservation need to regulate them. Human disturbances affected elephant habitat occupancy as well as site-level detectability. Rainfall is not an important limiting factor in this relatively humid bioclimate. Finally, we discuss cost-effective monitoring of Asian elephant populations and the specific spatial scales at which different population parameters can be estimated. We emphasize the need to model the observation and sampling processes that often obscure the ecological process of interest, in this case relationship between elephants to their habitat.

Synergistic effects of fire and elephants on arboreal animals in an African savanna.

Disturbance is a crucial determinant of animal abundance, distribution and community structure in many ecosystems, but the ways in which multiple disturbance types interact remain poorly understood. The effects of multiple-disturbance interactions can be additive, subadditive or super-additive (synergistic). Synergistic effects in particular can accelerate ecological change; thus, characterizing such synergies, the conditions under which they arise, and how long they persist has been identified as a major goal of ecology. We factorially manipulated two principal sources of disturbance in African savannas, fire and elephants, and measured their independent and interactive effects on the numerically dominant vertebrate (the arboreal gekkonid lizard Lygodactylus keniensis) and invertebrate (a guild of symbiotic Acacia ants) animal species in a semi-arid Kenyan savanna. Elephant exclusion alone (minus fire) had negligible effects on gecko density. Fire alone (minus elephants) had negligible effects on gecko density after 4 months, but increased gecko density twofold after 16 months, likely because the decay of fire-damaged woody biomass created refuges and nest sites for geckos. In the presence of elephants, fire increased gecko density nearly threefold within 4 months of the experimental burn; this occurred because fire increased the incidence of elephant damage to trees, which in turn improved microhabitat quality for geckos. However, this synergistic positive effect of fire and elephants attenuated over the ensuing year, such that only the main effect of fire was evident after 16 months. Fire also altered the structure of symbiotic plant-ant assemblages occupying the dominant tree species (Acacia drepanolobium); this influenced gecko habitat selection but did not explain the synergistic effect of fire and elephants. However, fire-driven shifts in plant-ant occupancy may have indirectly mediated this effect by increasing trees' susceptibility to elephant damage. Our findings confirm the importance of fire × elephant interactions in structuring arboreal wildlife populations. Where habitat modification by megaherbivores facilitates co-occurring species, fire may amplify these effects in the short term by increasing the frequency or intensity of herbivory, leading to synergy. In the longer term, tree mortality due to both top kill by fire and toppling by large herbivores may reduce overall microhabitat availability, eliminating the synergy.

Change in Mesoherbivore Browsing Is Mediated by Elephant and Hillslope Position.

Elephant are considered major drivers of ecosystems, but their effects within small-scale landscape features and on other herbivores still remain unclear. Elephant impact on vegetation has been widely studied in areas where elephant have been present for many years. We therefore examined the combined effect of short-term elephant presence (< 4 years) and hillslope position on tree species assemblages, resource availability, browsing intensity and soil properties. Short-term elephant presence did not affect woody species assemblages, but did affect height distribution, with greater sapling densities in elephant access areas. Overall tree and stem densities were also not affected by elephant. By contrast, slope position affected woody species assemblages, but not height distributions and densities. Variation in species assemblages was statistically best explained by levels of total cations, Zinc, sand and clay. Although elephant and mesoherbivore browsing intensities were unaffected by slope position, we found lower mesoherbivore browsing intensity on crests with high elephant browsing intensity. Thus, elephant appear to indirectly facilitate the survival of saplings, via the displacement of mesoherbivores, providing a window of opportunity for saplings to grow into taller trees. In the short-term, effects of elephant can be minor and in the opposite direction of expectation. In addition, such behavioural displacement promotes recruitment of saplings into larger height classes. The interaction between slope position and elephant effect found here is in contrast with other studies, and illustrates the importance of examining ecosystem complexity as a function of variation in species presence and topography. The absence of a direct effect of elephant on vegetation, but the presence of an effect on mesoherbivore browsing, is relevant for conservation areas especially where both herbivore groups are actively managed.

Elephant movement patterns in relation to human inhabitants in and around the Great Limpopo Transfrontier Park

The presence of humans and African elephants (Loxodonta africana) in the Great Limpopo Transfrontier Park can create situations of potential human–elephant conflict. Such conflict will likely be exacerbated as elephant and human populations increase, unless mitigation measures are put in place. In this study we analysed the movement patterns of 13 collared adult African elephants from the northern Kruger National Park over a period of eight years (2006–2014). We compared the occurrence and displacement rates of elephant bulls and cows around villages in the Limpopo National Park and northern border of the Kruger National Park across seasons and at different times of the day. Elephants occurred close to villages more often in the dry season than in the wet season, with bulls occurring more frequently around villages than cows. Both the bulls and the cows preferred to use areas close to villages from early evening to midnight, with the bulls moving closer to villages than the cows. These results suggest that elephants, especially the bulls, are moving through the studied villages in Mozambique and Zimbabwe at night and that these movements are most common during the drier months when resources are known to be scarce.Conservation implications: Elephants from the Kruger National Park are moving in close proximity to villages within the Great Limpopo Transfrontier Park. Resettlement of villages within and around the park should therefore be planned away from elephant seasonal routes to minimise conflict between humans and elephants.

Not the brain alone: The nutritional potential of elephant heads in Paleolithic sites

The presence of elephants, and specifically of elephant head remains, is well demonstrated in many Paleolithic sites in Europe, Africa, and Asia. However, the possible mechanisms for the exploitation of this enormous body part are rarely discussed, and it is often suggested that elephants' heads were exploited specifically for the extraction and consumption of the brain. In this paper, we discuss the nutritional potential that lies within elephants' heads as implied by ethnographic and zoological literature, and present archaeological evidence from Paleolithic sites for the exploitation of proboscideans' heads. The data show that the prevailing view should be re-evaluated, and that the nutritional potential within the elephant's head extends far beyond the brain. We suggest that organs such as the temporal gland, the trunk, the tongue, the mandible and the skull itself were exploited routinely as an integral part of early humans' diet. The nutritional potential of the elephant head provides a parsimonious explanation for the investment early humans put into transporting and exploiting this specific body part at open-air sites but particularly at cave sites, and serves as a significant beacon in understanding Paleolithic human behavior in relation to proboscidean remains.

HABITAT UTILIZATION BY ASIATIC WILD ELEPHANT (ELEPHUS MAXIMUS) IN PARSA WILDLIFE RESERVE, NEPAL

During the habitat survey, elephants foot prints, dungs and dirct observations were performed alongeach sampling grid. GPS points of elephant's presence were recorded. Aiming to provide quantitativeanalysis of habitats, circular quadrats of 10 m, 2 m and 1m radius were used to collect informationabout trees, shrubs and herbs. Importance values of each species of trees were analyzed by addingrelative frequency, relative density and relative dominance. Name of each forest type was determinedfrom the sequencial order of the large Importance Values (IV) of tree species. Prominance valueswere calculated for shrubs and ground flora and were used for classification of the shrub and groundvegetation type. Higher area of the reserve was covered by the forest (85.9%) followed by agricultureand buildup land (5.1%), grassland (3%), water body (3%), riverbed (2%) and shrub land (1%),respectively. Based on the phyto-sociological combination, forests of this reserve incorporated fourtypes namely Sal with Tarai Mixed Hardwood, Sal, Tarai Mixed Hardwood and Riverine.Ample ofevidence found inside lowler part (Tarai, Bhabar and the base of Churia) of Parsa Wildlife Reserve. Asuitable habitat with palatable forages (eg, Mallotus philippinnesis, Bananas, bamboos and climbers)for wild elephants were found inside the reserve. These forests were used as prime habitat by theAsiatic wild elephant.DOI: http://dx.doi.org/10.3126/eco.v20i0.11440ECOPRINTAn International Journal of EcologyVol. 20, 2013Page 41-52