Whole-tree Harvesting Research Articles

Comparing biomass and nutrient removals of stems and fresh and predried whole trees in thinnings in two Norway spruce experiments

In Denmark, thinning trees used for energy purposes are cut and left to dry in the stand before they are removed as whole trees. This practice causes shedding of needles and reduces nutrient removals for the benefit of long-term site fertility. It is uncertain, however, to what extent needles are shed and actual nutrient loss is affected by this practice. To address this question, we compared biomass and nutrient removals in two Norway spruce ( Picea abies (L.) Karst) experiments in western Denmark. Three contemporary thinning harvest intensities were examined: harvesting of fresh whole trees, predried whole trees, and stems only. The whole trees were chipped individually, and samples were removed to determine moisture and nutrient contents, whereas sample discs were removed from harvested stems. The biomass content of the cut whole trees was estimated to decrease 17% during predrying, whereas nutrient contents decreased 35%–60% for N, P, and K and <32% for Ca and Mg. The biomass content of stems was estimated to be 35%–42% lower than that of fresh whole trees. The corresponding differences in nutrient contents were in the range 84%–89% for N, P, and K and 73%–80% for Ca and Mg. Predrying and technological methods to reduce nutrient removals were compared and discussed.

Validation of an Integrated Estimation of Loblolly Pine (Pinus taeda L.) Leaf Area Index (LAI) Using Two Indirect Optical Methods in the Southeastern United States

Abstract Quality assessment of satellite-derived leaf area index (LAI) products requires appropriate ground measurements for validation. Since the National Aeronautics and Space Administration launch of Terra (1999) and Aqua (2001), 1-km, 8-day composited retrievals of LAI have been produced for six biome classes worldwide. The evergreen needle leaf biome has been examined at numerous validation sites, but the dominant commercial species in the southeastern United States, loblolly pine (Pinus taeda), has not been investigated. The objective of this research was to evaluate an in situ optical LAI estimation technique combining measurements from the Tracing Radiation and Architecture of Canopies (TRAC) optical sensor and digital hemispherical photography (DHP) in the southeastern US P.taeda forests. Stand-level LAI estimated from allometric regression equations developed from whole-tree harvest data were compared to TRAC–DHP optical LAI estimates at a study site located in the North Carolina Sandhills Region. Within-shoot clumping, (i.e., the needle-to-shoot area ratio [γE]) was estimated at 1.21 and fell within the range of previously reported values for coniferous species (1.2–2.1). The woody-to-total area ratio (α = 0.31) was within the range of other published results (0.11–0.34). Overall, the indirect optical TRAC–DHP method of determining LAI was similar to LAI estimates that had been derived from allometric equations from whole-tree harvests. The TRAC–DHP yielded a value 0.14 LAI units below that retrieved from stand-level whole-tree harvest allometric equations. DHP alone yielded the best LAI estimate, a 0.04 LAI unit differential compared with the same allometrically derived LAI.

Response of litter decomposition and soil C and N transformations in a Norway spruce thinning stand to removal of logging residue

Increasing demand for production of bioenergy has led to an interest in forest management which uses logging residue from both clear-cuttings and thinning stands. The aim of this study was to investigate the effects of removal of logging residue in a thinning Norway spruce stand on (1) litter decomposition and (2) soil microbial processes in C and N cycling and the quality of soil organic matter. The study site was a 40-year-old Norway spruce stand growing on a relatively fertile site. During thinning, logging residue was either removed (whole-tree harvest) or left on the site (stem-only harvest). Different types of material in the logging residue, from main branches to needles, were weighed separately into mesh bags. The bags were placed above the moss layer in the whole-tree harvest treatment and in the logging residue layer in the stem-only harvest treatment, and decomposition was monitored for 5 years after treatment. From the humus layer, samples were taken 10 years after treatment. Harvest method affected the mass loss of the litter material very little but the C-to-N ratio of the remaining material was slightly higher in whole-tree harvest than in stem-only harvest, particularly in the needle material. In the humus layer samples, taken 10 years after treatment, the rate of C mineralization was lower in whole-tree harvest than in stem-only harvest; also the rate of net N mineralization and the amounts of C and N in the microbial biomass tended to be lower, although not statistically significantly. Removal of logging residue had no effect on pH ( p H H 2 O 3.9 in both treatments) or C-to-N ratio (28 in both treatments) in the humus layer. The concentrations of total water-soluble phenols and an important group of phenols, condensed tannins, were both lower in the humus layer of whole-tree harvest than in that of stem-only harvest. Concentrations of sesqui-, di- or triterpenes in the humus layer were similar in both treatments. In conclusion, 10 years after harvest, soil microbial activities and organic matter characteristics in whole-tree harvest differed from those in stem-only harvest.

Effects of Mechanized Thinning and Prescription Fire on Stand Structure, Live Crown, and Mortality in Jeffrey Pine

ABSTRACT Forest thinning, using cut-to-length and whole-tree harvesting systems with subsequent underburning were assessed for their influence on stand structure, health, and fire resilience in uneven-aged Jeffrey pine (Pinus jeffreyi Grev. & Balf). Stand attributes, derived from measuring trees ≥ 10.2 cm diameter at breast height (DBH), were collected from permanent plots. Trees were divided into three size classes that generally corresponded to dominant/codominant, intermediate, and suppressed crown classes. Comparisons of post- to pre-burning mortality revealed significant thinning and fire main treatment effects as well as significant interaction between these two treatments in the two larger size classes. Mortality increased by 250% in the intermediate crown class within the burned stand portion of the whole-tree treatment, whereas among dominant/codominant trees mortality rose by 160% in the burned cut-to-length treatment combination. Pre- to post-burning shifts in live crown, expressed as a percentage of total tree height, were significantly influenced by both thinning and fire main treatments in the two larger size classes, while the interaction of these treatments was also significant among the largest trees. Within both of these size classes, decreases in live crown percentage were greatest in the burned portion of the unthinned treatment, where intermediate crown class trees lost over 20% of their crowns, while reductions in dominant/codominant trees averaged nearly 25%. The second highest losses for both size classes occurred within the burned cut-to-length treatment. In the smallest trees, mortality rose sharply and live crown decreased substantially after burning in both thinning treatments and in the unthinned control. Within the two larger size classes, preburn live crown size was negatively correlated with changes in crown size subsequent to underburning while DBH was negatively correlated with postburning changes in mortality, but only in intermediate crown class trees. These results present land managers with outcomes of differing management practices presently being evaluated for their potential to enhance forest health and reduce wildfire risk in the Sierra Nevada and similar dry forest regions.

Modelling the hydro-geochemistry of acid-sensitive catchments in Finland under atmospheric deposition and biomass harvesting scenarios

The dynamic hydro-geochemical Model of Acidification of Groundwater in Catchments (MAGIC) was used to predict the response of 163 Finnish lake catchments to historic and future atmospheric deposition (1880–2100) and future tree harvesting practices. Deposition was assumed to follow current legislated European emission reduction policies (CLe) and a scenario based on maximum (technically) feasible reductions (MFR). Future harvesting was assumed to shift from stem-only harvesting (SOH) to whole-tree harvesting (WTH) owing to the potential increased utilisation of biofuels. Despite the modest changes in atmospheric deposition under CLe (compared to current day), these reductions are predicted to halt the decline in soil base saturation; however, further reductions are required to improve soil and lake water chemistry. The MFR scenario predicted a significant long-term improvement in soil base saturation leading to continued long-term recovery in surface waters (all lakes with ANC > 0 by 2100). However under the WTH scenario, significant long-term impacts (re-acidification) were predicted for soil and surface water chemistry (14 lakes with ANC < 0 by 2100). To some extent the long-term negative impacts were reduced under MFR, indicating that increased utilisation of biofuels will necessitate ‘trading emissions for timber’, or soil amendment, to maintain ecosystem quality and sustainable forest growth. The current practice of SOH is close to the sustainable maximum harvesting under current (legislated) atmospheric deposition in Finland.

Chemical composition of forest floor and consequences for nutrient availability after wildfire and harvesting in the boreal forest

In boreal forests of eastern Canada, wildfire has gradually been replaced by clearcut harvesting as the most extensive form of disturbance. Such a shift in disturbance may influence the chemical properties of the forest floor and its capacity to cycle and supply nutrients, with possible implications for forest productivity. We compared the effects of stem-only harvesting (SOH), whole-tree harvesting (WTH) and wildfire on the chemical composition of forest floor organic matter and nutrient availability for plants, 15–20 years after disturbance in boreal coniferous stands in Quebec (Canada). The forest floor on plots of wildfire origin was significantly enriched in aromatic forms of C with low solubility, whereas the forest floor from SOH and WTH plots was enriched with more soluble and labile C compounds. The forest floor of wildfire plots was also characterized by higher N concentration, but its high C:N and high concentration of 15N suggest that its N content could be recalcitrant and have a slow turnover rate. Total and exchangeable K were associated with easily degradable organic structures, whereas total and exchangeable Ca and Mg were positively correlated with the more recalcitrant forms of C. We suggest that the bulk of Ca and Mg cycling in the soil–plant system is inherited from the influx of exchangeable cations in the forest floor following disturbance. The buildup of Ca and Mg exchangeable reserves should be greater with wildfire than with harvesting, due to the sudden pulse of cation-rich ash and to the deposition of charred materials with high exchange capacity. This raises uncertainties about the long-term availability of Ca and Mg for plant uptake on harvested sites. In contrast, K availability should not be compromised by either harvesting or wildfire since it could be recycled rapidly through vegetation, litter and labile organic compounds.

The economics of alternative fuel reduction treatments in western United States dry forests: Financial and policy implications from the National Fire and Fire Surrogate Study

We collected data at seven sites in the western US, on the costs of fuel reduction operations (prescribed fire, mechanical treatment, mechanical plus fire), and measured the effects of these treatments on surface fuel and stand parameters. We also modeled the potential behavior of wildfire in the treated and control stands. Gross costs of mechanical treatments were more expensive than those of prescribed fire, but net costs of mechanical treatments after deducting the values of harvested products were, on most sites, less than those of fire. The fire-only treatment reduced surface fuels, while most mechanical treatments (with the probable exception of whole-tree removal) increased these loads. Most mechanical-plus-fire treatments had little net effect on surface fuels. All treatments reduced the number of live trees, on average by about 300, 500 and 700 stems per hectare respectively for fire-only, mechanical, and mechanical-plus-fire. As intended by prescription, the mechanical treatments reduced basal area per hectare significantly. In most cases the fires – either alone or following mechanical treatment – killed mostly small trees, having essentially no impact on basal area. The mechanical-plus-fire treatment was the most effective, followed by fire-only, at reducing the modeled severity of wildfire effects under extreme weather conditions. The effectiveness of mechanical-only treatments depended on how much surface fuel remained on site. A whole-tree harvesting system removed the tops and limbs along with the felled trees, thereby reducing potential fire severity more than methods which left slash and/or masticated material within the stands. The various treatments created different conditions, and therefore the treatment intervals needed to maintain desired fire resilience would probably differ as well, being shorter for fire-only than for mechanical-only or mechanical-plus-fire treatments. Decisions about which treatments to prescribe, where, and when, will generally consider not only the financial costs and entry intervals, but other societal benefits and costs of the treatments and of wildfires as well.

Effects of harvest management practices on forest biomass and soil carbon in eucalypt forests in New South Wales, Australia: Simulations with the forest succession model LINKAGES

Understanding long-term changes in forest ecosystem carbon stocks under forest management practices such as timber harvesting is important for assessing the contribution of forests to the global carbon cycle. Harvesting effects are complicated by the amount, type, and condition of residue left on-site, the decomposition rate of this residue, the incorporation of residue into soil organic matter and the rate of new detritus input to the forest floor from regrowing vegetation. In an attempt to address these complexities, the forest succession model LINKAGES was used to assess the production of aboveground biomass, detritus, and soil carbon stocks in native Eucalyptus forests as influenced by five harvest management practices in New South Wales, Australia. The original decomposition sub-routines of LINKAGES were modified by adding components of the Rothamsted (RothC) soil organic matter turnover model. Simulation results using the new model were compared to data from long-term forest inventory plots. Good agreement was observed between simulated and measured above-ground biomass, but mixed results were obtained for basal area. Harvesting operations examined included removing trees for quota sawlogs (QSL, DBH >80 cm), integrated sawlogs (ISL, DBH >20 cm) and whole-tree harvesting in integrated sawlogs (WTH). We also examined the impact of different cutting cycles (20, 50 or 80 years) and intensities (removing 20, 50 or 80 m 3). Generally medium and high intensities of shorter cutting cycles in sawlog harvesting systems produced considerably higher soil carbon values compared to no harvesting. On average, soil carbon was 2–9% lower in whole-tree harvest simulations whereas in sawlog harvest simulations soil carbon was 5–17% higher than in no harvesting.

Influences of Mechanized Thinning and Prescribed Underburning on Radial Growth of Jeffrey Pine

Forest thinning utilizing cut-to-length and whole-tree harvesting systems with subsequent underburning were assessed for their impacts on several components of radial increment in uneven-aged Jeffrey pine (Pinus jeffreyi Grev. & Balf.) on the east slope of the Sierra Nevada during a period of extended drought. At the conclusion of three growing seasons following thinning and two seasons after burning, increment cores were extracted from dominant and codominant crown class trees over a century old as well as from intermediate crown class trees that were approximately 20 years younger. Separate measurements of early wood and late wood ring width during each of the 10 growing seasons preceding treatment and three seasons subsequent to thinning were summed for each year to obtain total ring width and permitted the calculation of proportions of total increment of the respective components. Comparisons of posttreatment values or averages of each component as well as of the magnitude of change, in both absolute and relative terms, in their values following treatment from 10-year pretreatment means were made among treatments for three posttreatment phases. Thinning treatment differences in direct measurements of the several components were marginal and uncommon. However, disparities in the magnitude of shifts in radial increment resulting from thinning were highly significant. Total ring width in dominant and codominant trees averaged 14% below to 13% above the pretreatment level in thinned portions of the stand over the three postthinning phases examined, while in the unthinned control total increment was 30-53% lower than the pretreatment mean, with the lowest values in all treatments occurring during the driest period of the study and highest observed after annual precipitation increased nearer to normal. In intermediate crown class trees, changes in ring width among the thinning treatments were similar to those in the dominant and codominant trees. Differences between the cut-to-length and whole-tree treatments were more pronounced in the dominant/codominant trees and comparisons generally revealed higher relative growth in the latter than in the former, while in the intermediate crown class, trees in the cut-to-length treatment performed marginally better in this regard than those in the whole-tree treatment. Influences of prescribed fire were limited to only a few components of radial growth in both sets of trees considered. The magnitude of change in several components of radial growth following thinning was found to be negatively correlated with residual basal area, but only in dominant and codominant trees. Live crown percentage influenced radial growth preceding thinning in both the dominant/ codominant and intermediate crown classes, in which it was a positive factor, as well as changes in increment subsequent to thinning in the former alone, when it was negative. Overall, results of this study demonstrate that considerable improvement in stand productivity can be realized with density management in an older, dry site forest type, which at minimum is not diminished by subsequent underburning.

Fluctuations dimensionnelles chez Pinus jeffreyi Grev. &amp; Balf. et variations dans le stockage d’eau influencées par l’éclaircie et le feu prescrit

Forest thinning utilizing cut-to-length and whole-tree harvesting systems with subsequent underburning were assessed for their impacts on water storage in the extensible tissues of dominant and codominant trees in an uneven-aged Jeffrey pine (Pinus jeffreyi Grev. & Balf.) stand on the east slope of the Sierra Nevada. Prior to the onset of the third growing season following thinning and the second season after burning, manual band dendrometers were installed at breast height on the selected trees and readings of diurnal fluctuation in stem circumference, an indication of bole water status, were taken monthly for one year. Diameter and relative diameter fluctuation were calculated from the circumference measurements. Overall, thinning had a positive influence on stem water recharge capacity, with the most pronounced effects evident in the latter part of the growing season. During this period, bole contraction in thinned stand portions was 49 to 55% greater than in the unthinned control, suggesting that both a greater volume of stored water was available for transpiration and was transpired in trees of the former treatment. There was no clear evidence that harvesting method affected stem water storage and influences of underburning were also absent entirely. Seasonal effects on diurnal changes in stem diameter were prominent, as the extent to which boles contracted generally increased over the course of the growing season, whereas fluctuations were at a minimum during the colder months. The magnitude of stem dimensional flux was found to be negatively correlated with initial tree DBH in one instance, while negative relationships between the former and live crown length as well as percentage were also revealed, albeit infrequently. Changes in bole size were positively correlated with residual basal area in some cases. These results suggest that improvement in water relations can be realized from density management in a dry site forest type with no apparent compromise of this benefit by broadcast underburning.

The influence of N load and harvest intensity on the risk of P limitation in Swedish forest soils

Nitrogen (N) is often considered to be the major factor limiting tree growth in northern forest ecosystems. An increased N availability, however, increases the demand for other nutrients such as base cations and phosphorous (P) which in turn may change which nutrient is the limiting factor. If P or base cations become limiting, N will start to leach which means a risk of increased eutrophication of surface waters. As many studies focus on base cations, this study instead aims at estimating P budgets on a regional scale for different harvesting scenarios relevant for Swedish conditions. P budget calculations were carried out for 14,550 coniferous sites from the Swedish National Forest Inventory, as weathering+deposition–harvesting–leaching. Three scenarios with different harvest intensities were used: 1) no harvesting, 2) stem harvesting and 3) whole-tree harvesting. The input data were derived from measurements and model results. The P budget estimates indicate that harvesting, especially whole-tree harvesting, result in net losses of P in large parts of Sweden. The highest losses were found in southern Sweden due to high growth rate in this area. In the whole-tree harvesting scenario the losses exceeded 1 kg ha − 1 y − 1 on many sites. N budget calculations on the same sites indicate that N generally accumulates in the whole country and especially in the southern parts. Consequently, the N and P budget calculations indicate that the forests in southern Sweden are in a transition phase from N-to P-limitation to growth. This transition will proceed as long as the accumulation of N continues. These results are important in a sustainable forestry context, as a basis for assessing the risk of future N leaching, and in designing recommendations for abatement strategies of N deposition and for application of wood ash recycling and N fertilization.

Biopower generation from mountain pine infested wood in Canada: An economical opportunity for greenhouse gas mitigation

Biomass is considered carbon neutral, and displacement of fossil fuel-based power by biomass-based power is one means to mitigate greenhouse gases. Large forest areas in British Columbia (BC), Canada, are infested by the mountain pine beetle (MPB). Dead wood from the infestation is expected to vastly exceed the ability of the pulp and lumber industry to utilize it; current estimates are that 200–600 million m 3 of wood will remain unharvested over the next 20 years. Regions where the damaged wood is not harvested will experience loss of jobs in the forestry sector, increased risk of forest fire hazard, carbon emissions from burned or decaying wood, and uncertainty about timing of replanting since this usually occurs at harvest. This paper reports the results of a detailed preliminary techno-economic analysis of producing power from MPB killed wood. Power plant size and location are critical factors affecting overall power cost. Overall cost of power rises steeply at sizes below 300 MW net power output. By locating the power plant in an area of high infestation, transportation distances can be minimized. A 300 MW power plant would consume 64 million m 3 of wood over a 20-year lifetime, and hence is a significant sink for otherwise unharvestable wood. Cost estimates are based on harvesting of whole dead trees with roadside chipping and transport to a central power plant located in either the Nazko or Quesnel regions of BC. A circulating fluidized bed boiler with a conventional steam cycle is a currently available technology demonstrated at 240 MW in Finland. The estimated power cost is $68 to $74 per MWh, which is competitive with other “green power” values in BC. Given recent values of export power in the Pacific Northwest, a 300 MW MPB power plant is viable with a carbon credit below $15 per tons of CO 2.

Investigating the soil acid–base status in managed boreal forests using the SAFE model

A dynamic soil chemistry model, SAFE, was applied to boreal stands located in three regions of Quebec (Canada) to investigate the relative contributions of forest harvesting methods (whole-tree and stem-only harvesting), natural disturbances (wildfire and spruce budworm outbreak) and atmospheric deposition on base cation fluxes and the soil acid–base status. Model parameterization was done using measured and published soil and vegetation data, published trends of nutrient dynamics, governmental databases on atmospheric deposition and climate, the reconstruction method MAKEDEP and throughfall equations that we developed for coniferous stands. The model suggested that forest disturbances, both anthropogenic and natural, increased base saturation, and thus base cation availability, to different degrees for periods of one to five decades. However, such disturbances, no matter their interval of return and their intensity, did not appear to be the main driving force of soil chemistry in the long term. On the other hand, simulation results showed that the B horizon at all sites has undergone acidification due to acid atmospheric deposition during the 20th century. Results suggest that atmospheric deposition is the main driver of long-term trends of soil chemistry for the study sites and that the issue of sustainability of soil productivity depends more on air pollution abatement policies than on forest management strategies.

Management alternatives of energy wood thinning stands

Abstract Energy wood thinning has become a feasible treatment alternative of young stands in Finland. Energy wood thinnings have been carried out mainly in stands where precommercial thinning has been neglected and the harvesting conditions for industrial wood thinning are difficult. Despite of its positive effects on harvesting costs and on renewable energy potential, whole-tree harvesting has been constantly criticized for causing growth loss. In this paper, the profitability of energy wood thinning was studied in 20 Scots pine-dominated stands where energy wood thinning was carried out. The growth of the stands after thinning was predicted with the help of Motti-stand simulator. Entire rotation time of the stands was simulated with different management alternatives. The intensity of first thinning and recovery level of logging residues varied between alternatives. In order to attain acceptable harvesting conditions, industrial wood thinning had to be delayed. The effect of energy wood thinning on subsequent stem wood growth was almost the same as in conventional thinning. Whole-tree harvesting for energy proved to be profitable alternative if the stumpage price is around 3€ m −3 , the interest rate is 3% or 5% and the removal of pulpwood is less than 20 m 3 ha −1 . If the harvestable pulpwood yield is over 20 m 3 ha −1 , integrated harvesting of industrial and energy wood or delayed industrial wood harvesting becomes more profitable.

Influences of Thinning and Prescribed Fire on Water Relations of Jeffrey Pine I. Xylem and Soil Water Potentials

Forest thinning accomplished with cut-to-length and whole-tree harvesting systems, and prescribed underburning were assessed for their impacts on water relations in eastern Sierra Nevada Jeffrey pine (Pinus Jeffreyi Grev. & Balf.) during a period of extended drought. Predawn and midday measurements of xylem water potential in dominant and codominant crown class trees more than a century old were made on six days spread over three growing seasons, accompanied by measurements of soil water potential completed between the predawn and midday sessions of each day. With the exception of a single predawn session, the only one of a total of 12 in which xylem water potentials did not differ among treatments, the potentials in trees of thinned stand subunits were 0.67 MPa higher on average during predawn sessions and 0.71 MPa higher during midday sessions than those in trees of the unthinned treatment. Differences between the cut-to-length and whole-tree treatments were marginal and uncommon, but when they occurred, potentials were higher in the former. Prescribed fire effects on xylem water potential were also uncommon, but when occurring generally indicated lower stress levels in the burned than in the unburned treatment. Soil water potentials largely coincided with those of xylem water, with higher potentials in either the cut-to-length or whole-tree treatments, and usually both, than in the unthinned treatment on each of the six days of measurement. Underburning influences on soil water were rare, but when evident, potentials were higher in the burned than in the unburned treatment by substantial margins. For a majority of the measurement sessions, xylem water potential was found to be negatively correlated with residual basal area but positively correlated with soil water potential. In turn, coarse fragments and organic matter in the soil profile intermittently influenced soil water potential, with the former a negative factor while the latter was positive. Overall, results of this study suggest that substantial ecophysiological advantages can be derived from density management in older, dry site forests, which at minimum are not compromised by subsequent implementation of controlled underburning.

Vascular and non-vascular plant community response to silvicultural practices and resultant microtopography creation in a forested wetland

Forested wetlands are important ecosystems valued for their indigenous plant communities, spatial heterogeneity, wildlife habitat, water quality, and timber resources. When harvested for timber, plant composition in these wetlands may change due to alteration in microsite habitats. Harvest severity also may affect plant composition. In this study, a mineral conifer wetland was subjected to whole-tree harvesting followed by installing different site preparations (bedding, trenching, draining). The original wetland overstory was Picea mariana, Larix laricina, and Pinus banksiana, with groundcover dominated by Sphagnum russowii. Eleven to twelve years after harvest, we assessed responses of vascular and non-vascular plants to created microtopographies (pits, side slopes, mounds) to determine whether harvest severity affected species richness, diversity, and relative cover of plant communities. For all the microtopography positions, the more severe harvest treatments (drained, cut, trenched) had the highest plant richness but the lowest diversity values. Richness and relative cover of Sphagnum species were highest in reference areas and much lower in the most severe harvest treatments (drained, cut, trenched). In contrast, graminoid and, to a lesser extent, herbaceous and woody plants increased in richness and cover after harvest.

Impact of Harvest Intensity on Long-Term Base Cation Budgets in Swedish Forest Soils

The effects of harvesting on the long-term mass balances of calcium (Ca), magnesium (Mg) and potassium (K) were evaluated on a regional level in Sweden. A new high-resolution weathering database was used together with estimates of total deposition, losses through harvest and leaching. Estimates were made for pine and spruce separately and for two harvesting intensity scenarios: stem harvesting and whole-tree harvesting. The mass balance calculations showed net losses of Ca and Mg in almost the whole country for both scenarios. The losses were smaller for pine than for spruce. The K balances were mainly positive for pine but negative for spruce. Leaching was a main factor in the mass balances, especially for Ca and Mg. Whole-tree harvesting in spruce forests led to substantially higher net losses of K and Ca than stem harvesting, according to the calculations. In the whole-tree harvesting scenario in spruce forests the estimated yearly net losses of Ca, Mg and K corresponded to at least 5%, 8% and 3% of the pools of exchangeable base cations, respectively, at 25% of the analysed sites. If losses of this magnitude continue the depletion of the pools of Ca, Mg and K may lead to very low base saturation of the soils, possibly accompanied by negative effects on soil fertility, runoff water quality, tree vitality and tree growth within a forest rotation in parts of Sweden. Avoiding whole-tree harvesting can improve the situation substantially for K, but the losses of Ca and Mg will still be significant.

Thinning and Prescribed Fire Effects on Forest Floor Fuels in the East Side Sierra Nevada Pine Type

Forest thinning accomplished with cut-to-length and whole-tree harvesting systems combined with prescribed underburning were assessed for their impacts on downed and dead fuel loading of all timelag categories in eastern Sierra Nevada Jeffrey pine (Pinus jeffreyi Grev. & Balf.). Cut-to-length harvesting resulted in an approximate doubling of total fuel loads with the greatest increases occurring in the 100-hr and 1000-hr categories, while changes associated with whole tree harvesting were marginal to the extent that overall posttreatment loading differed little between whole-tree and unfhinned treatments. Following the thinning operations, 1 + 10-hr and total fuel accumulations in the cut-to-lengfh treatment and 1000-hr fuels in the whole-tree treatment were positively correlated with harvested basal area and harvested foliage, branch, bole, and total tree biomass in simple regression models. Subsequent consumption during underburning eliminated 1 + 10-hr and 100-hr fuel additions from cut-to-lengfh harvesting along with a portion of the natural loading in these categories but was much less effective in reducing the 1000-hr fuels generated by this harvesting. Consumption of 1 + 10-hr, 100-hr, and total fuels in all thinning treatments was positively correlated with the amounts present within each category before underburning. Results reported here provide insight into fuel load modifications resulting from field practices that are being increasingly integrated into comprehensive management efforts to improve forest health and fire resilience in the western United States.

N FERTILIZATION EFFECTS ON DENITRIFICATION AND N CYCLING IN AN AGGRADING FOREST

We investigated N cycling and denitrification rates following five years of N and dolomite amendments to whole-tree harvested forest plots at the long-term soil productivity experiment in the Fernow Experimental Forest in West Virginia, USA. We hypothesized that changes in soil chemistry and nutrient cycling induced by N fertilization would increase denitrification rates and the N2O:N2 ratio. Soils from the fertilized plots had a lower pH (2.96) than control plots (3.22) and plots that received fertilizer and dolomite (3.41). There were no significant differences in soil %C or %N between treatments. Chloroform-labile microbial biomass carbon was lower in fertilized plots compared to control plots, though this trend was not significant. Extractable soil NO3- was elevated in fertilized plots on each sample date. Soil-extractable NH4+, NO3-, pH, microbial biomass carbon, and %C varied significantly by sample date suggesting important seasonal patterns in soil chemistry and N cycling. In particular, the steep decline in extractable NH4+ during the growing season is consistent with the high N demands of a regenerating forest. Net N mineralization and nitrification also varied by date but were not affected by the fertilization and dolomite treatments. In a laboratory experiment, denitrification was stimulated by NO3- additions in soils collected from all field plots, but this effect was stronger in soils from the unfertilized control plots, suggesting that chronic N fertilization has partially alleviated a NO3- limitation on denitrification rates. Dextrose stimulated denitrification only in the whole-tree-harvest soils. Denitrification enzyme activity varied by sample date and was elevated in fertilized plots for soil collected in July 2000 and June 2001. There were no detectable treatment effects on N2O or N2 flux from soils under anaerobic conditions, though there was strong temporal variation. These results suggest that whole-tree harvesting has altered the N status of these soils so they are less prone to N saturation than more mature forests. It is likely that N losses associated with the initial harvest and high N demand by aggrading vegetation is minimizing, at least temporarily, the amount of inorganic N available for nitrification and denitrification, even in the fertilized plots in this experiment.

Nutrient and carbon budgets in forest soils as decision support in sustainable forest management

Knowledge about the nutrient and carbon budgets in forest soils is essential to maintain sustainable production, but also in several environmental issues, such as acidification, eutrophication and climate change. The budgets are strongly influenced by atmospheric deposition as well as forestry. This study demonstrates how budget calculations for nitrogen (N), carbon (C) and base cations (BC) can be used as a basis for policy decisions on a regional level in Sweden. The study was based on existing nutrient and C budget calculations on a regional scale in Sweden. The nutrient budgets have been calculated for each square in a national 5 km × 5 km net by means of mass balances including deposition, harvest losses, leaching, weathering (BC) and fixation (N). Scenarios with different deposition and forestry intensity have been run and illustrated on maps. A simplified C budget has been estimated by multiplying the N accumulation with the C/N ratio in the organic layer, based on the assumption that the C/N ratio in the accumulating organic matter is equal to the ratio in the soil organic matter pool. The budget approaches differ from earlier budget studies since they involve regional high resolution data, combine deposition and forestry scenarios and integrate different environmental aspects. The results indicate that whole-tree harvesting will cause net losses of N and base cations in large parts of Sweden, which means that forestry will not be sustainable unless nutrients are added through compensatory fertilization. To prevent net losses following whole-tree harvesting, compensatory fertilization of base cations would be required in almost the whole country, whereas N fertilization would be needed mainly in the northern half of Sweden. The results further suggest that today's recommendations for N fertilization should be revised in southern Sweden by applying the southwest–northeast gradient of the N budget calculations. The C and N accumulation calculations show that C sequestration in Swedish forest soils is not an effective or sustainable way to decrease the net carbon dioxide emissions. A better way is to apply whole-tree harvesting and use the branches, tops and needles as biofuel replacing fossil fuels. This could reduce the present carbon dioxide emissions from fossil fuels substantially. The study shows that high resolution budget calculations that illuminate different aspects of sustainability in forest ecosystems are important tools for identifying problem areas, investigating different alternatives through scenario analyses and developing new policies. Cooperation with stakeholders increases the probability that the research will be useful.