In this study of thebelicoidal occlusal plane the relationships between tooth wear, the transverse slopes of mandibular molars and dental arch breadths were examined in 74 pre-contemporary Australian Aboriginal skulls. With increasing age and tooth wear the orientation of the mandibular occlusal surfaces increased towards the buccal. The differential occlusal orientation from first to third molars, present at eruption, tended to increase progressively with tooth wear. These functionally induced changes, together with regional differences in relative breadths of the maxillary and mandibular dental arches, are important in the development of abelicoidal occlusal plane.