Taphonomic Analysis Research Articles

Archeozoology and taphonomy of bird remains from Grotta di Castelcivita (Salerno, Italy) and clues for human-bird interactions

Grotta di Castelcivita (Campania, Southern Italy) is a cave-site containing a key archaeological sequence for the study of the Middle to Upper Palaeolithic transition in Southern Italy. A Late Mousterian occupation, whose modelled dates span from 47.6 to 41.1 ka cal BP, is followed by layers bearing evidence of the Uluzzian techno-complex, which underlie an articulated Protoaurignacian sequence. The prehistoric deposit is sealed by volcanic sediments attributed to the Campanian Ignimbrite (dated to 39.85 ± 0.14 ka BP), which represent a terminus ante quem for the Palaeolithic occupation of the cave. We present here the study of the avifauna collected during the excavations carried out at Castelcivita by the University of Siena in the years 1975–88. The examined sample is composed of 631 specimens, out of which 486 have been identified according to species, genus, family or order. Bird remains belong to 36 species and at least to 175 individuals. In the Late Mousterian the abundance of species adapted to open environments indicates a cool-temperate climate; water birds, and wood and forest birds are present as well. During the Uluzzian a shift towards colder climatic conditions is testified by the increase in steppe grassland species. In the Protoaurignacian the presence of birds of open and dry environments is more marked, even if climate seems to shift toward milder conditions at the end of this phase. Taphonomic analyses have provided significant evidence for the exploitation of birds by humans across the whole sequence. Clues of human activity on bird remains are attested both in the Mousterian and, more rarely, in the Protoaurignacian by traces, possibly indicating the intentional removal of feathers (in the Mousterian) and other kinds of carcass manipulation. The Uluzzian sample is the richest in human modifications. Some of them are related to an interest for feathers (on Pyrrhocorax graculus, Falco subbuteo and an Accipitriformes of large size). Other modifications (fresh bone fractures, burnt bones, peeling, arrachement) testify to carcass treatments of Galliformes, Anseriformes, Columbiformes, Charadriiformes and Passeriformes. In the Protoaurignacian traces due to anthropogenic activity are rare and there is an increase in bone modifications caused by carnivores. Results allow us to assume that at Castelcivita humans consistently hunted birds for several purposes and exploited (especially during the Uluzzian) some species to acquire an exclusive and ethnographically well-documented resource such as feathers.

A NEW ANALYTICAL PROCEDURE TO GRAPHICALLY CHARACTERIZE THE TAPHONOMIC PROPERTIES OF SKELETAL CARBONATES. AN EXAMPLE FROM MIOCENE LIMESTONES OF NEW ZEALAND

ABSTRACT A new graphical representation of the taphonomy of skeletal carbonates observed at the thin-section scale is proposed, demonstrating their utility in drawing information about the effects of early sea-floor processes on the post-mortem fate of fossil hard parts. The graphical representations consist of two-axis diagrams, which integrate the degree of fragmentation, abrasion, bioerosion, and encrustation as primary descriptors of the average state of fossil remains in a Miocene limestone section from southeastern North Island of New Zealand. Numerical values are expressed as percentages that represent the contribution of each factor to alteration (i.e., the four factors represent up to 25% each of the total alteration), the sum of values quantifying the amount of departure from pristine condition. Data are plotted in a cumulative form that reflects synergistic action of sea-floor processes towards hard parts destruction. One important virtue of this graphical representation is that the nature, degree, and variability of taphonomic alteration can be visualized and compared in a single diagram for several grain categories within a sample, and between samples. The proposed scheme is particularly flexible because more than four taphonomic categories can be integrated, independently from the number of alteration classes specific to each category, provided conversion of scores obtained in each taphonomic categories into percentages. Compiled results of such taphonomic analyses could be used in the future to identify specific depositional conditions, such as hydraulic regime, transportation, and residence time on the sea floor (a potential proxy to net accumulation rates).

Evidence for previously unknown mortuary practices in the Southwest of France (Fournol, Lot) during the Gravettian

European Mid Upper Paleolithic mortuary practices have been traditionally associated with primary burials, ochre, body ornaments and grave goods. Recently, evidence of the post-mortem treatment of skeletal remains, such as the displacement and removal of skeletal elements, has been reported for the Gravettian period in the Southwest of France. Here, we present the preliminary results of anthropological and taphonomic analyses of the human remains from the Gravettian site of Fournol (Soturac, Lot, France). We describe the first evidence of the scalping and disarticulation of human remains, a previously unknown post-mortem treatment in Gravettian contexts from the South West of France.

Assessing functionality during the early Acheulean in level TKSF at Thiongo Korongo site (Olduvai Gorge, Tanzania)

To understand the identity of the early Acheulean, it is necessary to discriminate between the variables that influenced the selection of technological strategies. Functionality of the archaeological sites is crucial in assessing the manufacturing strategies of lithic tools. To achieve this goal, analysis of the post-depositional processes must be evaluated. When bone remains have been preserved, anthropic animal processing can be identified through zooarchaeological and taphonomic analyses, and the spatial relationships among all the components can also be assessed, especially when bone surface is not sufficiently preserved.There are two levels present at Thiongo Korongo, ~1.3 Ma, TKLF and TKSF. These are in autochthonous position with no significant temporal diachrony but with substantial technological differences in the manufacturing of the lithic tools, which enables the analysis of the influence of human activities on technological behaviour. In order to evaluate this issue at TKSF, we present lithic, faunal, taphonomic, fabric, and spatial analyses.An assemblage of megaherbivores, among which Sivatherium is outstanding, dominate the TKSF faunal remains. Cortical preservation is poor; there is no intervention of carnivores, the rate of green fractures is low, and a few cut marks on size 5 and 3b animals were identified. Hence, apparently human intervention on the fauna was not intensive. Spatial and geostatistical analyses hints of a specific area where megaherbivores were processed by humans. Through wear use and biomarker analyses on stone tools, we are currently trying to understand the activities that were carried out in the remaining paleosurface.At paleosurface TKLF, the main anthropogenic input could be related to activities other than animal resource exploitation, in which large handaxes were necessary. Without the assessment of site functionality and chronological context, this data could have lead to the differences observed at TKLF and TKSF being attributed to different Acheulean stages.

The Use of Spatial Taphonomy for Interpreting Pleistocene Palimpsests: An Interdisciplinary Approach to the Châtelperronian and Carnivore Occupations at Cassenade (Dordogne, France)

One of the challenges commonly faced by Paleolithic archaeologists is disentangling archaeological layers in caves and rockshelters that often reflect complex palimpests. Layers defined in the field are primarly used to distinguish occupations, yet their actual nature and integrity are rarely tested or justified after excavation. Distinct occupations may become mixed together in a single field layer either following depositional and post-depositional processes (taphonomic admixture) or difficulties in reliably separating assemblages in the field (analytical lumping). Here we explore how three-dimensional spatial analyses combined with geoarchaeological and taphonomical data can be used to interpret Pleistocene palimpsests using the example of the Châtelperronian and carnivore occupations of Cassenade, a recently excavated site in Dordogne (France). We combine field observations with extensive post-excavation analysis (using spatial, geoarchaeological, lithic and faunal data, lithic particle-size distributions, fabrics, refits, and Bayesian modelling of radiocarbon dates) in order to (re)define assemblage boundaries and test their integrity. This approach resulted in a more comprehensive understanding of Cassenade sequence, including 1) increased stratigraphic resolution compared to initial field layer attributions; 2) evidence of how carnivore and human activity could be mixed by natural processes; 3) more reliable interpretations weighed against data from site formation processes; and, 4) a clearer understanding of the nature of the Châtelperronian occupations at Cassenade (short stopovers with a distinct site function?) and related mobility systems. Cassenade provides yet another example of the necessity of critically revising field layers after excavation through three-dimensional spatial and taphonomic analyses.

Paleobiological forensic study of an upper Miocene Pelobates tadpole (Anura, Pelobatidae) from the Tresjuncos lagerstätte of central Spain

ABSTRACTA paleobiological reconstruction of the last hours of life of an exceptionally well-preserved fossil tadpole is carried out through morphological and taphonomic analyses, allowing inferences to be made about its identification, developmental stage, physical condition, health, behavior, death, and burial. The tadpole was recovered from an upper Miocene (Turolian, MN13) diatomaceous konservat-lagerstätte deposit near Tresjuncos (Cuenca Province, Spain). The specimen belongs to an unnamed species of Pelobates (Pelobatidae) of the westernmost Palearctic phylogenetic clade. Its gigantic size was due neither to metabolic (hypothyroidism) nor to ecological (overwintering) causes but merely reflects an optimum living environment. Dying at Gosner developmental stage 41, the ossification pattern of the tadpole is slightly discordant with that of extant Pelobates cultripes, a greater degree for the skull and vertebral column in the fossil. The tadpole, seemingly in excellent health, had an intestine containing abundant remains of an epiphytic/epipelic diatom, different from the planktonic species surrounding the fossilized body. Death was probably violent, as revealed by a fossil hollow trace similar to hematomas made by egret beak strikes. Rapid burial coincided with an occurrence of massive planktonic diatom sedimentation distant from the lakeshore.

Late Mid Devonian Sawdonia (Zosterophyllopsida) from Venezuela

Premise of research. Fossil plants from the Late Middle to early Late Devonian age Campo Chico Formation, Sierra de Perijá, western Venezuela, are determined to represent a new species of the zosterophyll genus Sawdonia. While mentioned in previous publications, this plant has not been previously described or formally named. A conclusive taxonomic assignment was not possible until clarification of the sporangial morphology of the type species of Sawdonia, S. ornata (Gensel and Berry 2016), from the Early Devonian of Canada.Methodology. The adpressions were prepared using dégagement, including serial dégagement and photography at each stage. Permineralized remains were prepared using established techniques for pyrite sectioning and polishing. Taphonomic analysis included preparing models and subjecting them to compression to aid in understanding their shape and orientation in life.Pivotal results. The Venezuelan plants are adpressions of spiny axes that exhibit two types of branching (especially in vegetative ones), circinate tips, anatomy with G-type tracheids, and lateral sporangia with unequal-sized sporangial valves, with the abaxial valve being spinous as is characteristic of the genus Sawdonia Hueber emend Gensel and Berry 2016. Sporangia are irregularly arranged along some axes and appear to be located in medial to subdistal regions. Spores were not obtained.Conclusions. Differences in length and density of spines, the presence of two types of branching related to its growth habit, and details of sporangial morphology distinguish the Venezuelan plants from those of the type species and S. deblondii, thus leading to establishing a new species. This discovery demonstrates the longevity of the genus (ca. 20 Myr) during a time of profound vegetational change, including the rise of forests, and is the first record of a fertile zosterophyll recorded from the Middle Devonian of South America.

New data on avifaunal remains associated with the Middle Stone Age layers from Diepkloof Rock Shelter, Western Cape, South Africa

New bird remains were identified in the faunal assemblage recovered from the Middle Stone Age layers of Diepkloof Rock Shelter in the Western Cape Province of South Africa. The bird assemblage, while small, is composed of 12 distinct taxa, whose presence is linked to (1) the recovery of the material within a large rock shelter offering various roosting and/or nesting places (passerines and birds of prey such as Verreaux's eagle); the proximity of (2) the Verlorenvlei River (heron, white-breasted cormorant and African swamphen) and (3) the Atlantic Ocean coastline (African penguin and Cape cormorant). The occurrence of the coastal birds in the sequence seems to be stratigraphically consistent with other lines of evidence suggesting occasional human visits to the coast and correlates with putative periods of higher sea level and associated closer proximity of the shelter to the coastline (i.e. during the interglacial Marine Isotopic Stages 5 and 3). The taphonomic analysis shows a weak anthropogenic signal and suggests, rather, a primarily natural origin for most of the bird remains, possibly partly accumulated by a large raptor such as a Verreaux's eagle. By comparing data from Diepkloof Rock Shelter with data from Sibudu Cave, another large rock shelter located in a different part of South Africa but where the same techno-complexes have been recognized -namely the Still Bay and Howiesons Poort-, I touch on the possible influence of environmental and geographical factors on animal exploitation variability amongst late Pleistocene hunters-gatherers.

Level U3.1, a new archaeological level discovered at BK (upper bed II, Olduvai Gorge) with evidence of megafaunal exploitation

Excavations at Bell Korongo (BK) have yielded important evidence to infer different behaviours of early hominins in several archaeological levels since 1935. The present study shows the results for a newly geological and archaeological of BK (Level U3.1). This paper describes the geology of this newly discovered level, along with a taphonomical analysis of the archaeological remains to better understand the assemblage formation processes. This new level was deposited at the base of LA Unit 3, in decantation facies over a point-bar. Although these facies, found throughout the site furnish ideal conditions for preservation, this is the first time in BK level U3.1 that any archaeological or paleontological remains have been found embedded within them. The taphonomic study suggests that the assemblage is the result of a short time span involving processes accumulating remains from different thanatocoenoses created over a time span of less than one year, consistent with a sedimentation process triggered by flooding of the channel during the annual heavy rains in the wet season. Furthermore, a large percentage of bones, including megafaunal remains, appear cutmarked, percussion marks and anthropgenic green fractures. Level U3.1 is the youngest level at BK, and adds to the evidence of the megafaunal exploitation behaviour of early Pleistocene hominins, which has already been extensively reported in BK Levels 4 and 5 and other African Lower Pleistocene sites.

Early Upper Paleolithic subsistence in the Levant: Zooarchaeology of the Ahmarian–Aurignacian sequence at Manot Cave, Israel

The Early Upper Paleolithic period in the Levant is essential in the studies of the establishment of modern human communities outside Africa, and corresponding archaeological evidence may be used to shed light on human ecology, economy and demography. Specifically, cultural differences between two Early Upper Paleolithic entities, the Early Ahmarian and the Levantine Aurignacian, raise the question of differing adaptations. In this article we use archaeofaunal remains from the Early Upper Paleolithic sequence at Manot Cave (Western Galilee, Israel), to track human hunting patterns, carcass transport and processing within the Early Ahmarian (46–42 ka) and Levantine Aurignacian (38–34 ka) phases. We test two hypotheses: 1) the Ahmarian and Aurignacian represent adaptations to different environments; and 2) the two entities differ in mobility patterns and site use.Our multivariate taphonomic analysis showed subtle differences in depositional processes between the two phases and demonstrated a primarily anthropogenic complex. In both phases, human subsistence was based on two ungulate species, mountain gazelle (Gazella gazella) and Mesopotamian fallow deer (Dama mesopotamica), with some contribution from birds, tortoises and small mammals. Among the gazelles, it appears that female herds were targeted, and that hunting took place close to the cave. The results of the research show great similarity in environmental exploitation between the Ahmarian and Aurignacian phases concerning prey spectrum and choice, carcass transport and processing. These patterns occupy a middle position between the Middle Paleolithic and the late Epipaleolithic of the region. Despite this, there are also several significant differences between the phases such as increased exploitation of small game (especially birds) and faster accumulation and higher densities of material in the Aurignacian. This may indicate greater occupation intensity during the Aurignacian compared to the Ahmarian, and thus could explain the outstanding character of this entity in the Levant.

Taphonomic analysis of the level 3b fauna at BK, Olduvai Gorge

The association of stone tools and modified bones at archaeological African Plio-Pleistocene sites is essential to understanding the socio-economics of early hominin behavior. Previous research at BK suggests the site was repeatedly visited by hominins for short time periods. During these occupations, the hominins had a primary role in the exploitation of small, medium, and large size carcasses. The present study analyzes the faunal material recovered from level 3b at BK (BK3b). The results indicate that BK3b operated as a place consciously selected by hominins to carry out specific activities, which resulted in a large accumulation of lithic materials and bone remains. The high frequency of cut-marked specimens, even more than is documented at FLK Zinj, supports the idea that hominins at BK3b butchered and demarrowed the bones from animals of all sizes.

Taphonomic analysis of saurischian dinosaurs from the Plottier Formation (Upper Cretaceous), Mendoza, Argentina

The uppermost Cretaceous continental deposits of the Neuquén Basin (west-central Argentina) have yielded a high diversity vertebrate assemblage, including numerous dinosaur species. The Neuquén Group is the richest unit in remains of dinosaur of the basin and comprises the Río Limay, the Río Neuquén and the Río Colorado subgroups. In this group, there is abundance of isolated, disarticulated or partially articulated sauropods and theropods. However, little is known about the taphonomic history of fossil assemblages. In this study, dinosaur remains from the Plottier Formation (late Coniacian-early Santonian) found in the Cerro Guillermo area in southern Mendoza are examined. The investigation of fossil occurrences within the study area revealed the existence of different taphonomic modes, from isolated bones until partially articulated skeletons preserved in deposits of floodplain and fluvial channel settings. SEM-EDS analysis showed the substitution of hydroxyapatite by francolite in the bone microstructure. The presence of fluorine-in one of the cases-suggested a link between the elemental composition and depositional environments: floodplain and fluvial channel. The survey of the vertebrate accumulation types and their sedimentary context allowed documenting a wide range of processes responsible for their genesis, operating within a meandering fluvial environment. This contribution represents a holistic approach about taphonomic history of Cretaceous dinosaurs for assessing the differential preservation of fossil assemblages in fluvial environments.

The Early Triassic Jurong fish fauna, South China: Age, anatomy, taphonomy, and global correlation

Abstract As the higher trophic guilds in marine food chains, top predators such as larger fishes and reptiles are important indicators that a marine ecosystem has recovered following a crisis. Early Triassic marine fishes and reptiles therefore are key proxies in reconstructing the ecosystem recovery process after the end-Permian mass extinction. In South China, the Early Triassic Jurong fish fauna is the earliest marine vertebrate assemblage in the recovery period. It is constrained as mid-late Smithian in age based on both conodont biostratigraphy and carbon isotopic correlations. The Jurong fishes are all preserved in calcareous nodules embedded in black shale of the Lower Triassic Lower Qinglong Formation, and the fauna comprises at least three genera of Paraseminotidae and Perleididae. The phosphatic fish bodies often show exceptionally preserved interior structures, including network structures of possible organ walls and cartilages. Microanalysis reveals the well-preserved micro-structures (i.e. collagen layers) of teleost scales and fish fins. Abundant small pyrite framboids, 2–5 μm in diameter, are detected from the nodules and fish body surfaces, indicating a calm, euxinic burial environment. Coccoid-like microspheroids are also very abundant in the host rocks and near the fish fossil surfaces, implying that microbes may have participated in the burial process of the fishes. Taphonomic analysis uncovers the four-step formation process of the fish nodules. (1) Fishes lived in the oxic seawater in the upper ocean, and (2) their bodies sank to the anoxic seabed after death, with the body surface being wrapped by bacteria. (3) Microbial biofilms sealed body surfaces to prevent or delay the decay of the fleshy body. The decomposition of the body cavity and interior organs produced some CO2 and H2S gases. The former formed bicarbonate ions in seawater and attracted calcium ions to facilitate the precipitation of calcium carbonate, while the H2S combined with iron ions in seawater to form pyrite framboids. (4) The fish nodule gradually grew by precipitation of calcium carbonate in layers and embedding with pyrite framboids, and later the fish fossil nodule was compacted during diagenesis. Global faunal correlations indicate that the Jurong fishes are closely related to the Early Triassic fish faunas from Chaohu, Anhui Province and Madagascar.

The Body in the Ditch: Alternative Funerary Practices on the Northern Frontier of the Roman Empire?

ABSTRACTDisarticulated human remains were recovered from a first-century fort ditch at Vindolanda on the north-west frontier of the Roman Empire. Ancient DNA analysis revealed the skeleton to be that of a male individual and forensic taphonomic analysis suggested a primary deposition of the body in a waterlogged environment with no obvious evidence of formal burial. Occurrences of disarticulated human remains outside a cemetery context are often overlooked in Roman bioarchaeology. This discovery adds to the growing body of literature regarding alternative funerary practice in the Empire, highlighting that the concept of burial and disposal of the dead is more complex than ancient historical sources suggest. Details of the DNA analysis are provided in the Supplementary Material available at https://doi.org/10.1017/S0068113X1900014X.

Did climate determine Late Pleistocene settlement dynamics in the Ach Valley, SW Germany?

The loss of Neanderthal groups across Western and Central Europe during Oxygen Isotope Stage (OIS) 3 has held the attention of archaeologists for decades. The role that climatic change, genetic interbreeding, and interspecies competition played in the extinction of Neanderthal groups is still debated. Hohle Fels is one of several important Middle and Upper Paleolithic sites from the Ach Valley in southwestern Germany which documents the presence of Neanderthals and modern humans in the region. Chronological and stratigraphic records indicate that these two groups occupied the site with little to no overlap or interaction. This provides the opportunity to examine the behavioural variability of Swabian Neanderthal populations without the complication of cross-cultural influence. In this study we contribute a terrestrial paleoenvironmental record derived from the small mammal material from Hohle Fels Cave to the ever-growing archaeological record of this period. By reconstructing the climate and landscape of the Ach Valley during this time we can identify the effect that the OIS 3 environment had on the presence of Neanderthals in the region. Based on indicator taxa and the habitat weighing method, the small mammal record, which includes rodents, insectivores, and bats, from Hohle Fels shows that the earliest Neanderthal occupation took place on a landscape characterized by substantial woodland and forest, rivers and ponds, as well as moist meadows and grasslands. A gradual increase in cold tundra and arctic environments is clear towards the end of the Middle Paleolithic, extending to the end of the early Aurignacian which may correlate with the onset of the Heinrich 4 event (~42,000 kya). Our taphonomic analysis indicates the material was accumulated primarily by opportunistic predators such as the great grey owl, snowy owl, and European eagle owl, and therefore reflects the diversity of landscapes present around the site in the past. Importantly, at the time Neanderthals abandoned the Ach Valley we find no indication for dramatic climatic deterioration. Rather, we find evidence of a gradual cooling of the Swabian landscape which may have pushed Neanderthal groups out of the Ach Valley prior to the arrival of modern human Aurignacian groups.

Holocene small mammals hunted by owls and humans in southern Brazil: taphonomic evidence and biological significance

Understanding resource utilization and economic diversification amongst Holocene hunter‐gatherers in southern Brazil requires in‐depth taphonomic analysis of faunal assemblages. Three Early to Late Holocene archaeological sites (Garivaldino, Pilger and Sangão) in Rio Grande do Sul State, southern Brazil, revealed large assemblages of small mammal (˂1 kg) remains, composed mainly of rodents. To appreciate depositional processes of fauna in relation to human consumption, taphonomic attributes and processes were evaluated. The results indicated that the remains accumulated through different antemortem taphonomic pathways, apparently linked to the size and natural history of the species represented. The small‐ and medium‐sized (<150 g) cricetid rodents (e.g. Pseudoryzomys simplex and Sooretamys angouya) were represented by a low proportion of teeth with lightly digestive corrosion suggesting that they were derived from the predatory activity of owls, possibly Tyto alba. In contrast, large‐sized cricetids (>150 g; Kunsia tomentosus, Gyldenstolpia sp. and Holochilus sp.), and several caviomorph rodents with aggregated spatial distributions (Echimyidae (†Dicolpomys fossor, Phyllomys sp., †Clyomys riograndensis and †Euryzygomatomys mordax), Caviidae (Cavia sp.) and Ctenomyidae (Ctenomys sp.)) showed evidence of thermoalteration patterns and cut marks on bones, suggesting human exploitation. A postmortem depositional environment was deduced from dendritic and branched patterns of rootlet etching, apparent mostly at Garivaldino and Pilger. Polished areas and holes on bone surfaces, and impregnation of manganese showed moderate (Sangão) or low (Garivaldino and Pilger) incidence, pointing to water transport under moderate (Pilger) and low (Garivaldino and Sangão) energy over short distances. The results obtained here provide the first clear evidence of early human exploitation of small mammals in southern Brazil, suggesting a diversification of economies. In addition, as several of the recorded rodents are today regionally or biologically extinct, a preliminary discussion about the potential impact of humans on this process is provided.

Taphonomy of fish concentrations from the Upper Jurassic Solnhofen Plattenkalk of Southern Germany

The density of vertebrate fossils on bedding planes is generally very low in the Upper Jurassic plattenkalks forming the Solnhofen fossil Lagerstatte. There are, however, a few bedding planes where large concentrations of the small fis Leptolepides sprattiformis (BLAINVILLE, 1818) occur. The aim of this paper is to analyse and interpret these f sh concentrations from a taphonomic point of view. For this purpose, the taphonomic features of L. sprattiformis concentrations on three large slabs were semi-quantitatively analysed. The concentrations of individuals on bedding planes are explained as recording local mass mortality events caused by tropical storms that, for short intervals, disrupted the pycnocline which separated hypersaline, oxygen-poor to anoxic bottom waters from normal marine surface waters in the Plattenkalk basins. Mass mortality is supported by tetany features such as gaping jaws, hyperextended branchiostegal rays, and splayed f ns, which occur in the majority of the specimens. The generally excellent preservation of the f sh resulted from lack of postmortem f oatation, rapid burial and/or overgrowth by microbial f lms, and quick re-establishment of the pycnocline with inhospitable conditions on the basin floo. The post-mortem concave-arching of the backbones and various displacements of the vertebral column were most likely caused by varying decay resistance of soft tissues and the pressure of decay gases in the body cavities, respectively. The taphonomic analysis of f sh concentrations can provide very detailed palaeoenvironmental information and serves as an independent line of evidence for palaeoenvironmental reconstructions that complements geochemical and sedimentary data.

TAPHONOMY AND PALEOECOLOGY OF AN OPHIUROID-STYLOPHORAN OBRUTION DEPOSIT FROM THE LOWER DEVONIAN BOKKEVELD GROUP, SOUTH AFRICA

The taphonomy of a well preserved ophiuroid-stylophoran assemblage from the Bokkeveld Group, Lower Devonian of South Africa is described using micro-CT scanning techniques. This assemblage provides a taphonomic window into the structure of Early Devonian, echinoderm-dominated communities within the Malvinokaffric Realm of SW Gondwana. Micro-CT scanning and 3D modeling of a portion of an obrution bed revealed an assemblage of over 700 ophiuroids observed of the newly described Gamiroaster tempestatisReid 2019 associated with two species of stylophorans of two species, the mitrates Paranacystis cf. petriiCaster 1954 and Placocystella africana (Reed 1925). The 3D model permitted detailed sedimentologic and taphonomic analysis of the bed including determination of the degree of fossil articulation, orientation, and faunal counts. The results indicate that deposition took place in a storm-influenced, shallow marine environment smothering both attached and vagrant taxa of the benthic community during a high-energy storm event. Micro-CT scanning is a powerful taphonomic tool for the non-destructive analysis of delicate echinoderm assemblages that would otherwise be difficult to study.

The consumption of tortoise among Last Interglacial Iberian Neanderthals

Optimal foraging theory and diet breadth models often place large mammals in top-ranking positions due to their high-energy return. However, mass collection of small prey can result in comparable return rates, and dietary diversity is nutritionally beneficial on its own right. A growing body of evidence recovered from several sites in the Mediterranean Basin confirms Neanderthal use of small size prey. Slow-moving, tortoises are an easy catch, and human collection and consumption is demonstrated by taphonomic analysis. In Portugal, two key Middle Palaeolithic cave sites, Gruta da Oliveira and Gruta da Figueira Brava, provide pertinent evidence. Based on an improved osteometric method, in which long bone measurements are standardised and analysed together using the Logarithmic Size Index (LSI), it is possible to better gauge the contribution of tortoises to the diet and the human impact on the species’ populations. At inland Gruta da Oliveira, a tendency towards the massive collection of tortoises, affecting the local population more severely, is apparent. At coastal Gruta da Figueira Brava, the data suggest opportunistic collection upon encounter, possibly due to the availability of a more diverse range of resources.

Megamamíferos Pleistocênicos da Toca da Pedreira, Ourolândia, Bahia, Brasil: Taxonomia e Inferências Tafonômicas Pleistocenic

The present paper records the occurrence of pleistocenic megamammals fossils (indeterminate Cervidae, indeterminate Toxodontinae and Eremotherium laurillardi) in a new small cave in Ourolândia municipality, Bahia: the “Toca da Pedreira”. In addition to the taxonomic identification, a taphonomic analyses was undertaken. The results allow us to suggest that the material suffered little to medium transport, by a high energy hydraulic flow, from the thanathocoenosis to the Toca da Pedreira cave, which promoted the high fragmentation of the fossils; that the fossils of E. laurillardi and Toxodontinae suffered temporal mixing; and that the fossiliferous assembly could be classified as peripheral.