Five new species of Solanum sect. Geminata are described from Peru and Ecuador: S. clivorum, S. goniocaulon, S. malletii, S. smithii, and S. tovarii. Relationships within section Geminata are discussed, and an illustration and distribution map are provided for each species. Solanum is one of the largest genera of flowering plants, with some 1,500-2,000 valid species. Most of these species are neotropical, with taxa occurring in all habitats throughout the tropics and subtropics. Peru and Ecuador are particularly rich in Solanum species, probably due to their great diversity in habitat (Weberbauer, 1945). Section Geminata is one of the largest of the subdivisions of the nonspiny solanums, with about 145 species, all but one of these neotropical. Species of section Geminata are distinguished by their small white flowers, woody habit, usually leaf-opposed inflorescences, and hard green fruits at maturity (in all except the S. pseudocapsicum alliance). Solanum clivorum S. Knapp, sp. nov. TYPE: Peru. La Libertad: entre Huamachuco y Cajabamba, 3,000-3,200 m, 16 Mar. 1948, Ferreyra 3059 (holotype, USM; isotypes, IBE, MO, US). Figure 1. Frutex; caules dense pubescentes; sympodia difoliata geminata vel plurifoliata; folia elliptica supra glabra subtus dense dendritico-pubescentia; inflorescentiae foliis oppositae vel terminales dendriticae; corolla alba; bacca globosa in pedicello deflexo inserta; semina complanata reniformia. Bushy shrubs, 2-3 m tall; young stems and leaves densely pubescent with dendritic trichomes ca. 0.5 mm long; bark of older stems reddish or green, glabrous. Sympodial units difoliate, geminate, or appearing plurifoliate due to rapid inflorescence growth. Leaves elliptic, commonly with several axillary shoots, glabrous (occasionally with scattered simple uniseriate trichomes on the lamina) and with a few dendritic trichomes along the main veins adaxially, densely pubescent with dendritic trichomes abaxially; major leaves 13-21 cm long, 6-9 cm wide, with 7-9 pairs of primary veins, these drying yellowish, the apex acute, the base acute; petiole 2-3 cm long; minor leaves differing from the majors only in size, 7-10 cm long, 3-4 cm wide, the apex acute, the base acute; petiole 1-2 cm long. Inflorescences opposite the leaves or appearing shootterminal, 3-7 cm long, many-times branched, 2040-flowered, densely pubescent with dendritic trichomes; pedicel scars unevenly spaced 1-2 mm apart. Buds globose, the corolla only halfway exserted from the calyx tube. Pedicels at anthesis erect, tapering, 6-8 mm long, ca. 0.5 mm diam. at the base, densely pubescent with soft dendritic trichomes like those of the rest of the inflorescence. Flowers with the calyx tube conical, 1-1.5 mm long, the lobes deltate, 0.5-1 mm long, densely pubescent with dendritic trichomes; corolla white, 1-1.1 cm diam., lobed halfway to the base, the lobes planar at anthesis, densely pubescent on abaxial lobe surf ce with minute, dendritic trichomes; anthers 1.52 mm long, 1-1.5 mm wide, poricidal at the tips, the pores not opening to slits; free portion of the filaments ca. 0.5 mm long, the filament tube ca. 0.5 mm long, with 5 triangular projections ca. 0.5 mm long alternating with the anthers and closely investing the style; ovary glabrous; style 3-4 mm long, glabrous; stigma capitate, the surface minutely papillose. Fruit a globose, green berry, 1-1.2 cm diam.; fruiting pedicels woody, erect, 1.5-1.7 cm long, ca. 1.5 mm diam. at the base. Seeds dark reddish brown, flattened-reniform with thickened margins, 3-4 mm long, 2-2.5 mm wide, the surfaces minutely pitted. Distribution. In the Cajabamba valley, department of Cajamarca, Peru, on steep rocky slopes from 2,000 to 2,800 m. Figure 5. Paratypes. PERU. CAJAMARCA: Cutervo, Lagunac, camino a Cochabamba, 2,000 m, 26 Feb. 1985, Llatas Quiroz 1178 (F); Cajabamba, Finca Zil, 2,500 m, 22 Aug. 1985, Mostacero L. & Guerra L. 63 (HUT, MO, NY); Cajamarca, entre Matara y Namora, 2,600 m, 16 Aug. 1973, Sdnchez Vega et al. 1217 (NY); NamoraMatara, 2,600 m, 16 Aug. 1973, Sagdstegui A. 7755 (HUT, MO, NY); Huamachuco-Cajabamba, 2,800 m, 16 Nov. 1983, Sagdstegui A. 11144 (HUT, NY). NOVON 2: 341-350. 1992. This content downloaded from 157.55.39.110 on Wed, 14 Dec 2016 05:03:59 UTC All use subject to http://about.jstor.org/terms