In the investigation of the floral anatomy of the Cruciferae and some allied families, with the object of determining the fuindamental floral structure of the groups, the writers have obtained evidence which compels them to reopen a subject of long controversy-the nature of the ovary of the Cruciferae. Further, light is also shed upon the recently proposed theory of carpel polymorphism as applied to this family. The structure of the crucifer gynoecium hlas long been a subject of discussion, especially in regard to the number and type of carpels, and the nature of the septum, or false partition. Students of this subject have held several theories as to the nature of the ovary. Of these the chief are, (i) that the pistil consists of two carpels; (2) that there are four carpels, either in one whorl or in two whorls of two each. That the number of carpels is six has also been maintained; and recently as many as IO-20 have been considered to compose the pistil in some genera. The two-carpel theory seems to be commonly held today. Accordiulg to this understanding there are two open carpels fused edge to edge forming the ovary. The fused margins of these carpels bear the placentae, and outgrowths of the placentae divide the really one-celled ovary. The septum is thus false. Those students holding that there are four carpels lhave generally believed that two of these are reduced or aborted. In such cases the ovules are said to be borne on placenital surfaces (of which there are four), placed between the normal and the reduced carpels; and opinions have varied as to the carpel to which the ovules belong, and hence which type of carpel is sterile and which is fertile. A prominent theory, put forth and emphasized by older writers, and recently renewed, and supported with new facts by Miss Saunders (26), is that the so-called valves of the ovary make up the sterile carpels, the fertile carpels being composed of the segments between the valves. The relationship of these two types of carpels can be understood by reference to text figure I. Many supporters of this theory, as well as of the two-carpel theory, have considered the septum, or false partition, to be a secondary placental outgrowth. A few have considered this dissepiment to represent carpellary tissue of the fertile carpels. If the septum is truly carpellary, then it must represent either the venitral part of the carpel itself or a much expanded margin of the carpel. If the dissepiment is false, that is, of placental
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