The presence of the fern Weichselia in a restricted sedimentary interval of a drilling core in the Parnaíba Basin (Brazil) was used as a lead for paleoenvironmental interpretation integrating paleobotanical, palynofacies, palynological, organic petrography and clay mineralogy analyses. The fern paleobiogeography was amplified and its association with terrestrial bryophyte gametophytes (Muscites) indicated depositional conditions corresponding to marginal areas of freshwater bodies subjected to frequent flooding under the general fluvial-deltaic conditions so far accepted for the intermediary portion of the Codó Formation. The high dominance of non-opaque phytoclasts and very scarce autochthonous non-marine palynomorphs pointed to a shallow water body linked to marginal areas of fluvial systems in the river outlets, channel margins in estuarine systems, and/or shallow floodplain lakes connected/open to fluvial canals. The very scarce marine palynomorphs (dinoflagellate cysts) suggest a limited influence from unstable environments on coastal margins or estuarine canals. The terrestrial vegetation surrounding the depositional setting, deciphered by palynological analysis, reflected distinct environmental conditions prevailing simultaneously in 1) humid areas dominated by fern communities, and 2) dry-xerophytic areas dominated mainly by Araucariaceae and Cheirolepidiaceae gymnosperms, both as components of a wider Aptian paleoenvironment within the periequatorial latitudes in South America. The results were supported by clay mineralogy that showed abundance of detrital kaolinite over montmorillonite and suggest the dominance of a climatic humidification process and a less expressive semiarid climate in a regional context.