Widespread and ongoing increases in park and protected area (PPA) visitation presents managers with the imminent challenge of preserving ecological conditions while also maintaining accessibility in the open spaces that are fundamental to the health and wellness needs of society. In the field of recreation resource management, recreation specialization, the selective channeling of interests and abilities into a specific recreational activity, has contributed to a more comprehensive understanding of recreation behavior, site preference, management perceptions and conservation support. Contributing to historical understandings of specialization can inform recreation planning on the diversity of uses occurring in urban-proximate PPAs. In this study, survey and GPS data collected in urban and peri-urban parks in the Nature Reserve of Orange County, CA, USA, were used to classify recreationists into three specialization types based on empirically derived dimensions of involvement, commitment and skill. For each survey participant, thirteen spatio-temporal metrics (STM) were calculated. A principle component analysis (PCA) reduced STMs into 3 factors representing expressions of spatial behavior and a one-way analysis of variance indicated unique patterns between specialization types and time spent recreating, elevation gained, speed traveled and stopping behavior. Additionally, GPS point data were analyzed with an analytical approach adopted from the field of wildlife movement ecology; a resource selection function (RSF). The RSF assisted in quantifying spatial distribution patterns specific to each specialization type across activity types at four park locations and demonstrated a landscape-based statistical analysis of probability of use in relation to change in elevation, distance to starting points and park amenities. Committed hikers and mountain bikers largely demonstrated longer trip durations, more elevation gain, less overall stopping, faster overall speeds and farther total distances, while e-bikers in this study demonstrated unintuitive specialization dynamics; casual specialization types travelling farther from starting points and dispersing more across trail systems. Further spatial results revealed spatial behavior to be inherently complex and influenced by numerous confounding factors (i.e. activity type/bike type, starting points, park topography and trail design). This novel spatial examination of recreation specialization according to activity type and park location, is useful in understanding recreation behavior and park use in a spatial context. This ability is helpful in a predictive managerial context when reviewed in correspondence with historical evidence identifying behaviors with increased potential for ecological impact.