Quaternary History Research Articles

Cosmogenic 10Be exposure dating of glacial erratics on Horseshoe Island in western Antarctic Peninsula confirms rapid deglaciation in the Early Holocene

AbstractThe rapid warming observed in the western Antarctic Peninsula gives rise to a fast disintegration of ice shelves and thinning and retreat of marine-terminating continental glaciers, which is likely to raise global sea levels in the near future. In order to understand the contemporary changes in context and to provide constraints for hindcasting models, it is important to understand the Late Quaternary history of the region. Here, we build on previous work on the deglacial history of the western Antarctic Peninsula and we present four new cosmogenic 10Be exposure ages from Horseshoe Island in Marguerite Bay, which has been suggested as a former location of very fast ice stream retreat. Four samples collected from erratic pink granite boulders at an altitude of ~80 m above sea level yielded ages that range between 12.9 ± 1.1 ka and 9.4 ± 0.8 ka. As in other studies on Antarctic erratics, we have chosen to report the youngest erratic age (9.4 ± 0.8 ka) as the true age of deglaciation, which confirms a rapid thinning of the Marguerite Trough Ice Stream at the onset of Holocene. This result is consistent with other cosmogenic age data and other proxies (marine and lacustrine 14C and optically stimulated luminescence) reported from nearby areas.

The bones beneath the streets: drifting through London’s Quaternary

This article reflects on a Situationist dérive in Central London, which mobilised a creative engagement with the city’s Quaternary history (the last 2.6 million years). The aim was to animate paleoecological knowledge in the resistant, opaque and frenetic environment of a dense urban centre. This brief excursion into an alternative London is offered as a model for contemporary drifting that stretches out beyond our immediate situation to connect to successive geological and biological strata, reframing experiences of the urban environment through shifting scales and chronologies. The Situationists had declared ‘sous les pavés, la plage!’ (under the paving stones, there is a beach!), evoking the playful space of possibility behind the veneer of the city’s systems and structures. This drift aimed to search even deeper, encountering spectral inhabitations revealed by the bones beneath the streets. The article argues that uncovering these hidden ecologies has the potential to counter the urban prevalence of spectacular representations of wildlife and develop an eco-politics of co-existence.

Quaternary evolution of the river Danube in the central Pannonian Basin and its possible role as an ecological barrier to the dispersal of ground squirrels

ABSTRACT Large rivers can act as important ecological barriers for small mammals, but their exact role is debated. Here we examine the effect of the river Danube on populations of the European ground squirrel, an important representative of steppe communities, by studying the Quaternary history both of ground squirrels and of the Danube in the Pannonian Basin, in Hungary. The Danube experienced considerable channel changes during the Quaternary, with >100 km westward shift of its middle reach, interrupted by temporary eastward movements. Besides autogenic river processes, tectonically induced avulsions have played a role in re-routing channels. Three of the four Spermophilus species certainly extended their ranges on both sides of the Danube, referring to repeated river crossings during the Quaternary. Hydrological events, most importantly rapid river course changes (avulsions) but also meander cut-offs are viable mechanisms to enable ground squirrel populations to cross large rivers. On longer timescales, oscillating river course changes can maintain gene exchange across the river.

The earliest fossil record of the bandicoot rat (Bandicota indica) from the early Middle Pleistocene of Taiwan with discussion on the Quaternary history of the species

Three molar fossils of murine rodents were newly discovered from an outcrop in the Tsailiao area of Tainan City, ROC. They were dated to the early Middle Pleistocene (ca. 0.7 Ma) based on the stratigraphic and chronological data for the layers distributed at and around the outcrop. A detailed systematic analysis of the fossils reliably referred them to Bandicota indica. This species is now distributed widely in the Oriental Region, including Taiwan, where it usually lives in human-associated environments. Therefore, it is generally believed that this species was introduced artificially into Taiwan very recently. However, the fossils demonstrate that B. indica inhabited Taiwan in the early Middle Pleistocene. They also imply that this species has inhabited Taiwan continuously since the early Middle Pleistocene, although this inference must be tested with nonmorphological methods. The literature on the fossil record of Bandicota was examined to understand its chronospatial distribution throughout the Oriental Region, and indicates that the fossils from the Tsailiao area represent the earliest record of B. indica. The history of the species is discussed on the basis of these data.

Quaternary history of the Lake Magadi Basin, southern Kenya Rift: Tectonic and climatic controls

Sediments from the Magadi Basin (south Kenya Rift) preserve a one-million-year palaeoenvironmental record that reflects interactions between climatic, volcanic and tectonic controls. Climate changes that impacted sedimentation include wet-dry cycles on variable timescales and an overall progressive trend towards greater aridity. Volcanic influences involved inputs of tephra to the basin, significant inflow of geothermal fluids, and the effects of weathering, erosion and transportation of clastics from trachyte and basalt terrains. Tectonic controls, which were often step-like, reflect the influence of faults that provided pathways for fluids and which controlled accommodation space and drainage directions.Intensified aridity and evaporative concentration resulted in salinity and pH increasing with time, which led to a change from calcite deposition in mildly saline lakes before 380 ka to the later formation of zeolites from reactions of volcaniclastic debris with highly alkaline lake and pore water. After 105 ka, hyperalkaline conditions led to trona accumulation and increasingly variable rare earth elements (REEs). The presence of mixed saline and freshwater diatom taxa between 545 and 16 ka indicates climate variability and episodic inputs of fresh water to saline lakes. Calcrete formed in lake marginal settings during semi-arid periods.Tectonic controls operated independently of climate, but they interacted together to determine environmental conditions. Aquatic deposition was maintained during periods of increasing aridity because fault-controlled ambient and geothermal springs continued to flow lakewards. This recharge, in turn, limited pedogenesis: palaeosols are common in other rift floor sequences. Trona formed when aridity and evapoconcentration increased, but its precipitation also reflects increased magmatic CO2 that ascended along faults. Basin fragmentation and north-south fractures caused loss of cross-rift (east-west) drainage from rift-marginal basalts, resulting in reduced transition metals after 545 ka. The Magadi Basin demonstrates how a careful reconstruction of these complex tectono-climatic interactions is essential for accurate palaeoenvironmental reconstruction in continental rifts and in other tectonic settings.

Late Weichselian glacial history of Forlandsundet, western Svalbard: an inter-ice-stream setting

The last glacial maximum (LGM) and post-glacial Quaternary history of Forlandsundet, the strait between western Spitsbergen and Prins Karls Forland, are enigmatic. Previous terrestrial field studies report contradicting evidence for an ice sheet either overriding the entire strait or completely absent during the LGM. Here, we present a multi-proxy investigation of marine sediments, high-resolution bathymetric data and aerial imagery from Forlandsundet. We reveal glacial till present at 15 cal. ka BP and geomorphological landforms characteristic to an inter-ice-stream glaciated margin. This new evidence implies that the Forlandsundet region was fully glaciated during the LGM. This glaciation was followed by a stepwise retreat of glacial ice during the Bolling–Allerod (14.7–12.7 cal. ka BP) and Younger Dryas (12.7–11.7 cal. ka BP). The Holocene record from the marine sediments is incomplete, with a hiatus from approximately 11.8 to 7.4 cal. ka BP, interpreted as an erosive event. By the mid-Holocene (7.4 cal. ka BP), more temperate, Atlantic conditions based on the benthic foraminiferal assemblages prevailed, and are followed by gradual cooling into the late Holocene (< 4 cal. ka BP). This study provides new data to resolve the LGM extent of the Svalbard–Barents Sea ice sheet in the Forlandsundet region and sheds light on the deglacial ice dynamics in a palaeo-inter-ice-stream area driven by the inflow of warm Atlantic water.

Early Quaternary history of the genus Archidiskodon (Proboscidea, Elephantidae) in Western Siberia: to the question of intermediate links in mammoth lineage

The paper is concerned with the early representatives of the Archidiskodon–Mammuthus lineage from the sediments of Lower Pleistocene of the South of Western Siberia (Kuznetsk Basin). Several forms were being described within the genus Archidiskodon (A. meridionalis, A. m. tamanensis, A. m. voigtstedtensis, A. aff. wusti) which represent consecutive steps of progressive development of the lineage on the early stage of its evolution. Morphometric features of molars (M3) have been studied using the author’s original method designed for this group, which allowed specifying systematic attribution of the taxa under study and their position in the structure of the mammoth lineage. It was demonstrated that the form previously distinguished as Wust’s elephant and later attributed to the species M. trogontherii significantly differs from the latter in frequency and length of the plates. This form also stands out well statistically and is regarded as a separate and the latest taxon among the Archidiskodon group of elephants. A. wusti occupies a distinct position of an intermediate form (a ‘bridge’ as H. Pohlig put it) between the genera Archidiskodon and Mammuthus in the structure of the mammoth lineage. Thus, the multi-dimensional diagrams used in our method permit to take a step close to revealing of so-called intermediate links in the mammoth lineage. Direction of phyletic development and some adaptations of the group in the second half of Early Pleistocene are also discussed. Account is also given to the accompanying fauna and the existence period of these species on the territory under study.

Comment on paper by Ritter et al. (2018), Evidence for multiple Plio-Pleistocene lake episodes in the hyperarid Atacama Desert, published in Quaternary Geochronology: v. 44, p. 1–12. DOI: 10.1016/j.quageo.2017.11.002

Abstract Ritter et al. (2018) analyzed the Quaternary history of a purported paleo-lake in the Atacama Desert of northern Chile. They provide a much more thorough analysis than previous publications of the field relationships where there are landform benches surrounding a bedrock knob, Cerro Soledad. They conclude that the benches formed due to shoreline erosion by a paleo-lake while Cerro Soledad rose in elevation due to tectonic activity. Ritter et al. (2018) also used terrestrial cosmogenic nuclide (Neon, Beryllium, and Aluminum) data to determine a history of the landforms, and thus what they conclude is a history of former lake levels and uplift rate. This Comment examines the distribution of mapped geological units, especially Pliocene lake deposits, relative to the locations sampled and described by Ritter et al. (2018). We find that some of the benches sampled for cosmogenic nuclide dating occur on Miocene and Pliocene lacustrine strata, which opens the possibility that clasts were recycled from Miocene-Pliocene lake deposits into the Pleistocene bench clast population. We reach the alternative interpretation that differential resistance to erosion by hillslope processes created most benched forms on the flanks of Cerro Soledad, rather than generation of the benches by shoreline processes. An alternative exhumation scenario which utilizes the cosmogenic exposure ages better explains many of the landforms. In this scenario, the cosmogenic nuclide exposure dates reveal valley-scale denudation in response to tectonic uplift of Cerro Soledad, reduction of valley floor elevation by dissolution of evaporite units, and capture of the valley by the Loa River, rather than shoreline erosion. Irrespective of the alternative interpretation, the exposure history chronological information provided by Ritter et al. (2018) provides new opportunities to examine environmental fluctuations in the central Atacama Desert.

The Quaternary rivers of the Jurassic Coast region: From the Neogene to the Anthropocene

The Jurassic Coast World Heritage Sites (JCWHS) is not only a 95 km long coastline and remarkable Mesozoic geological section, but also a slice through a Quaternary landscape. For the majority of the last two million years this landscape lay in the periglacial zone, just south of a waxing and waning ice margin and just north of an Atlantic inlet which eventually became the English Channel. This paper reviews how the previous landscape inherited from the Cenozoic, was modified through uplift, climatically driven fluvial activity and periglaciation. Much evidence of this Quaternary history can be seen today in sections along the JCWHS coast which cuts through a number of headwater valleys the largest of which are the Exe and Axe. Recent studies, largely funded from the Aggregate Levy Tax, have produced the first independent chronologies for the Exe and Axe valleys and a model of how periglaciation interacted with the layer-cake stratigraphy of the Mesozoic bedrocks of the JCWHS. The Quaternary history of the JCWHS is also preserved in raised beaches on the Isle of Portland, coastal landforms, and in Holocene alluvial sediments associated with human activity and which may constitute part of the putative Anthropocene. An appreciation of the Quaternary history of the JCWHS is also important in understanding modern geological hazards from landslides to flooding.

Impacts of Quaternary History on Critical Zone Structure and Processes: Examples and a Conceptual Model From the Intensively Managed Landscapes Critical Zone Observatory

The concept of a critical zone (CZ) supporting terrestrial life has fostered groundbreaking interdisciplinary science addressing complex interactions among water, soil, rock, air and life near Earth’s surface. Pioneering work has focused on the CZ in areas with residual soils and steady-state or erosional topography. CZ evolution in these areas is conceptualized as progressive weathering of local bedrock (e.g. in the flow-through reactor model). However, this model is not applicable to areas in which weathering profiles form in transported materials including the formerly glaciated portion of the Central Lowland of North America. We present a new conceptual model of CZ evolution in landscapes impacted by continental glaciation based on investigations at three study sites in the Intensively Managed Landscapes Critical Zone Observatory (IML-CZO) The IML-CZO is devoted to the study of CZ processes in a region characterized by thick surficial deposits resulting from multiple continental glaciations, with bedrock at depths of up to 150 m. Here the physical (glacial ice, loess, developing soil profiles) and biological (microbes, tundra, forest, prairie) components of the CZ vary significantly in time. Moreover, the spatial relationships between mineral components of the CZ record a history of glacial-interglacial cycles and landscape evolution. We present cross-sections from IML-CZO sites to provide specific examples of how environmental change is recorded by the structure of the mineral components of the CZ. We build on these examples to create an idealized model of CZ evolution through a glacial cycle that represents the IML-CZO sites and other areas of low relief that have experienced continental glaciation. In addition, we identify two main characteristics of CZ structure which should be included in a conceptual model of CZ development in the IML-CZO and similar settings: (1) mineral components have diverse origins and transport trajectories including alteration in past CZs, and, (2) variability in climate, ecosystems, and hydrology during glacial-interglacial cycles profoundly influence the CZ composition, creating a legacy retained in its structure. This legacy is important because the current physical CZ structure influences the occurrence and rates of CZ processes, as well as future CZ responses to land use and climate change.

The southernmost record of a large erethizontid rodent (Hystricomorpha: Erethizontoidea) in the Pleistocene of South America: Biogeographic and paleoenvironmental implications

The South American porcupines (Erethizontidae) are included in two genera: Chaetomys and Coendou. The latter is a very speciose taxon, with about 13 living species. During at least the late Pliocene–early Pleistocene, erethizontids immigrated to Central and North America during the Great American Biotic Interchange. Although some Pleistocene fossils have been reported, the Quaternary history of this clade is still understudied. The only known extinct species is Coendou magnus. In this work, a fossil of a porcupine is reported from an Upper Pleistocene fluvial sedimentary sequence cropping out in the Northern Pampa geomorphological region, Santa Fe Province, Argentina. Despite this group having different living forms widely distributed in South American Neotropical woodland habitats, the Pleistocene occurrences of Erethizontidae are scarce and limited to Upper Pleistocene deposits from Bolivia, Brazil, and Uruguay. Currently, the specimen here reported represents the only Pleistocene porcupine from Argentina with a stratigraphical context. The morphological characters as well as the dimensions indicate that it is close to the Pleistocene erethizontid Coendou magnus. In this context, the presence of this erethizontid in such a southern locality, together with other taxa recorded from this site and the associated geological and paleoenvironmental evidence, indicates subtropical conditions, compared with the current conditions, which may have allowed a southern displacement of taxa more related to woodlands and xeric subtropical environments.

Lateglacial environmental change in Scotland

ABSTRACTThis paper reviews the evidence for environmental change during the Lateglacial period (c.14.7–11.7 ka), perhaps the most intensively studied episode in the Quaternary history of Scotland. It considers first the stratigraphic subdivision and nomenclature of the Lateglacial, before proceeding to a discussion of the various lines of proxy evidence that have been used to reconstruct the spatial and temporal patterns of environmental change during this time period. These include pollen and plant macrofossil data; coleopteran and chironomid records; diatom data; stable isotope and geochemical records; and evidence for human activity. The paper then considers the principal methods that have been employed to date and correlate Lateglacial events: radiocarbon dating; surface exposure dating; varve chronology; and tephrochronology. This is followed by an examination of the constraints imposed on environmental reconstructions, an account of the ways in which the evidence can be employed in the development of an event stratigraphy for the Lateglacial in Scotland, and a proposal for a provisional Lateglacial type sequence (stratotype) at Whitrig Bog in SE Scotland. Emphasis is placed throughout on the potential linkages between the Scottish records and the isotopic signal in the Greenland ice cores, which forms the stratigraphic template for the N Atlantic region. The paper concludes with a discussion of the strategies and approaches that should underpin future research programmes on Lateglacial environmental change in Scotland.

Palynology and the Ecology of the New Zealand Conifers

The New Zealand conifers (20 species of trees and shrubs in the Araucariaceae, Podocarpaceae and Cupressaceae) are often regarded as ancient Gondwanan elements, but mostly originated much later. Often thought of as tall trees of humid, warm forests, they are present throughout in alpine shrublands, tree lines, bogs, swamps, and in dry, frost-prone regions. The tall conifers rarely form purely coniferous forest and mostly occur as an emergent stratum above evergreen angiosperm trees. During Maori settlement in the 13th century, these fire-sensitive trees succumbed rapidly, most of the drier forests being lost. As these were also the more conifer-rich forests, ecological research has been skewed towards conifer dynamics of forests wetter and cooler than the pre-human norm. Conifers are well represented in the pollen record and we here we review their late Quaternary history in the light of what is known about their current ecology with the intention of countering this bias. During glacial episodes, all trees were scarce south of c. 40°S, and extensive conifer-dominant forest was confined to the northern third of the North Island. Drought- and cold-resistant Halocarpus bidwillii and Phyllocladus alpinus formed widespread scrub in the south. During the deglacial, beginning 18,000 years ago, tall conifers underwent explosive spread to dominate the forest biomass throughout. Conifer dominance lessened in favour of angiosperms in the wetter western lowland forests over the Holocene but the dryland eastern forests persisted largely unchanged until settlement. Mid to late Holocene climate change favoured the more rapidly growing Nothofagaceae which replaced the previous conifer-angiosperm low forest or shrubland in tree line ecotones and montane areas. The key to this dynamic conifer history appears to be their bimodal ability to withstand stress, and dominate on poor soils and in cool, dry regions but, in wetter, warmer locations, to slowly grow thorough competing broadleaves to occupy an exposed, emergent stratum where their inherent stress resistance ensures little effective angiosperm competition.

Eolian sand and loess deposits indicate west-northwest paleowinds during the Late Pleistocene in western Wisconsin, USA

AbstractOur study adds to the Quaternary history of eolian systems and deposits in western Wisconsin, USA, primarily within the lower Chippewa River valley. Thickness and textural patterns of loess deposits in the region indicate transport by west-northwesterly and westerly winds. Loess is thickest and coarsest on the southeastern flanks of large bedrock ridges and uplands, similar in some ways to shadow dunes. In many areas, sand was transported up and onto the western flanks of bedrock ridges as sand ramps, presumably as loess was deposited in their lee. Long, linear dunes, common on the sandy lowlands of the Chippewa valley, also trend to the east-southeast. Small depressional blowouts are widespread here as well and often lie immediately upwind of small parabolic dunes. Finally, in areas where sediment was being exposed by erosion along cutbanks of the Chippewa River, sand appears to have been transported up and onto the terrace treads, forming cliff-top dunes. Luminescence data indicate that this activity has continued throughout the latest Pleistocene and into the mid-Holocene. Together, these landforms and sediments paint a picture of a locally destabilized landscape with widespread eolian activity throughout much of the postglacial period.

Late Quaternary fluvial dynamics and landscape evolution at the lower Shulaveris Ghele River (southern Caucasus)

AbstractThe southern Caucasus was intensively settled through the Neolithic period to present. Studies of late Quaternary fluvial dynamics and landscape development at the lower Shulaveris Ghele River in southeastern Georgia aid understanding of the Quaternary history of this region. Our studies show that following river aggradation to form a fan-shaped alluvial surface during the late Pleistocene, fluvial sedimentation shifted west, leaving a fan surface subject only to local fluvial and colluvial processes as well as late Neolithic settlement. At about 6 ka, the river avulsed to the east and eroded some late Neolithic settlements occupying the eastern portion of the fan. The avulsion was followed by 3 distinct episodes of aggradation at about 6 ka, 3–2 ka, and since 1.0 ka. No such aggradation is evident in the early Holocene. Regional Holocene fluvial activity in this area was apparently influenced by vegetation dynamics, possibly controlled by regional climatic and/or anthropogenic factors, and significantly differs from the eastern Mediterranean region. All late Neolithic settlements were originally built far from an active river, possibly indicating late Neolithic water management systems.

Early Holocene wild boar remains from Tsudupisuki-abu Cave on Miyako Island of the Southern Ryukyus, Japan

A large number of Sus remains were recently collected from early Holocene sediments of Tsudupisuki-abu Cave on Miyako Island of the Southern Ryukyus, Japan, during excavations. Wild boar (Sus) are now absent from the island, where Sus remains were also uncommon in Quaternary fossil localities before the excavations. Taxonomic analyses of the Sus remains, especially metrical analysis of third molars, have revealed that the remains are divided into two clusters (A and B) of different size. Cluster A consists of larger specimens and cluster B of smaller specimens and are referred, respectively, to Sus scrofa and Sus sp. On the basis of the information of Sus remains from this cave and another locality (Pinza-abu Cave), we reconstruct a late Quaternary history of Sus on Miyako Island. It is inferred that large-sized Sus scrofa was introduced by humans onto the island in the late Late Pleistocene, and then reduced in size to be Sus scrofa of Tsudupisuki-abu Cave by a mechanism of insular dwarfism until the early Holocene. On the other hand, the second human introduction of a miniature Sus form (= Sus sp.) seems to have taken place in the earliest Holocene.

The Quaternary history of effusive volcanism of the Nevado de Toluca area, Central Mexico

Andesite and dacite lava flows and domes, and intermediate-mafic cones from the Nevado de Toluca area were classified into five groups using field data and 40Ar/39Ar geochronology constraints. Thirty-four lava units of diverse mineralogy and whole-rock major-element geochemistry, distributed between the groups, were identified. These effusive products were produced between ∼1.5 and ∼0.05 Ma, indicating a mid-Pleistocene older-age for Nevado de Toluca volcano, coexisting with explosive products that suggest a complex history for this volcano. A ∼0.96 Ma pyroclastic deposit attests for the co-existence of effusive and explosive episodes in the mid-Pleistocene history. Nevado de Toluca initiated as a composite volcano with multiple vents until ∼1.0 Ma, when the activity began to centralize in an area close to the present-day crater. The modern main edifice reached its maximum height at ca. 50 ka after bulky, spiny domes erupted in the current summit of the crater. Distribution and geochemical behavior in major elements of lavas indicate a co-magmatic relationship between different andesite and dacite domes and flows, although unrelated to the magmatism of the monogenetic volcanism. Mafic-intermediate magma likely replenished the system at Nevado de Toluca since ca. ∼1.0 Ma and contributed to the eruption of new domes, cones, as well as effusive-explosive activity. Altogether, field and laboratory data suggest that a large volume of magma was ejected around 1 Ma in and around the Nevado de Toluca.

No obvious genetic erosion, but evident relict status at the westernmost range edge of the Pontic-Pannonian steppe plant Linum flavum L. (Linaceae) in Central Europe.

We investigate patterns of genetic variation along an east–west transect of Central European populations of Linum flavum and interpret the Quaternary history of its peripheral populations, especially those at the westernmost isolated range edge, discussing their migrations and possible relict status. We defined our peripheral transect across three study regions from Central Hungary, eastern Austria to southwestern Germany. Using AFLP fingerprinting and cpDNA sequence variation (rpL16 intron, atpI‐H), we analyzed 267 and 95 individuals, respectively, representing each study region by four populations. Hierarchical AMOVA (AFLPs) indicated significant variation among study regions (12% of total variance) and moderate differentiation between populations (10%). Population differentiation was high at the westernmost range edge (11.5%, Germany), but also in the east (13.4%, Hungary), compared to the Austrian study region (8.6%). Correspondingly, AFLP diversity was highest in the center of the study transect in eastern Austria. CpDNA haplotypes support a pattern of regional structuring with the strongest separation of the westernmost range edge, and some haplotype sharing among Austrian and Hungarian individuals. Equilibrating nucleotide versus haplotype diversity patterns, the highly diverse populations at the Pannonian range edge (Austria) indicate long‐term persistence, while Central Pannonian populations are obviously effected by recent bottlenecks. Intermediate nucleotide, but high haplotype diversity within the westernmost exclave (Swabian Alb), is indicative of a founder bottleneck during its pre‐LGM or early postglacial migration history, followed by sufficient time to accumulate cpDNA variation. The not obviously reduced genetic diversity and distinctiveness of L. flavum at the westernmost range edge suggest a long‐term persistence (relict status) of populations in this region, where the species has survived probably even the Würm glaciation in extra‐Mediterranean refugia. This genetic relict variation represents an important part of the overall genetic diversity found in the western periphery of this steppe plant and highlights the high conservation priority of respective gene pools.

Quaternary history, population genetic structure and diversity of the cold-adapted Alpine newt Ichthyosaura alpestris in peninsular Italy

Mediterranean peninsulas are major biodiversity hotspots, and cold-adapted species are an important component of this biodiversity. However, cold-adapted species contributed surprisingly little to our knowledge of the intimate links between Quaternary environmental changes, species’ responses to these changes, and current patterns of intraspecific biodiversity. Here, we investigated the genetic structure and evolutionary history of a cold-adapted amphibian, the Alpine newt Ichthyosaura alpestris, within the Italian peninsula. Nuclear and mitochondrial markers consistently identified three distinct genetic lineages, whose divergence dates to the Early Pleistocene (1.9 and 0.8 million years ago). Our results show that the Italian peninsula provided multiple Pleistocene refugia to this cold-adapted species, and suggest that allopatric fragmentation followed by secondary admixture have been key events in the formation of its current pattern of genetic diversity. Indeed, estimates of population genetic diversity clearly identified contact populations as those achieving the highest levels of diversity. Such concordance among cold-adapted and temperate species in terms of processes triggering the formation of regional patterns of genetic diversity provides strong support for the hypothesis that gene exchange between divergent lineages, rather than long-term stability of refugial populations, has been the main step toward the formation of hotspots of intraspecific biodiversity.

Geochemistry and related studies of Clyde Estuary sediments

ABSTRACTGeochemical and related studies have been made of near-surface sediments from the River Clyde estuary and adjoining areas, extending from Glasgow to the N, and W as far as the Holy Loch on the W coast of Scotland, UK. Multibeam echosounder, sidescan sonar and shallow seismic data, taken with core information, indicate that a shallow layer of modern sediment, often less than a metre thick, rests on earlier glacial and post-glacial sediments. The offshore Quaternary history can be aligned with onshore sequences, with the recognition of buried drumlins, settlement of muds from quieter water, probably behind an ice dam, and later tidal delta deposits. The geochemistry of contaminants within the cores also indicates shallow contaminated sediments, often resting on pristine pre-industrial deposits at depths less than 1m. The distribution of different contaminants with depth in the sediment, such as Pb (and Pb isotopes), organics and radionuclides, allow chronologies of contamination from different sources to be suggested. Dating was also attempted using microfossils, radiocarbon and 210Pb, but with limited success. Some of the spatial distribution of contaminants in the surface sediments can be related to grain-size variations. Contaminants are highest, both in absolute terms and in enrichment relative to the natural background, in the urban and inner estuary and in the Holy Loch, reflecting the concentration of industrial activity.