Main Vegetation Types Research Articles

Latitudinal distribution, co-occurring tree species and structural diversity of the threatened palm Borassus aethiopum (Arecaceae) in Benin, West Africa

Background and aims – Borassus aethiopum Mart. is a wild palm species with high subsistence importance in West Africa. Extensive agriculture and overharvesting of its stem and fruits for multiple uses have caused a decrease of its natural populations in its native range. For conservation purposes, the distribution, abundance and structural diversity of the species were investigated across ten phytodistricts in three biogeographical zones in Benin. Two hypotheses were tested (i) tree floristic composition of B. aethiopum natural habitat changes with phytodistricts and (ii) structural diversity of B. aethiopum changes with phytodistricts, both as potential adaptation strategies to changing ecological conditions. Methods – Geographical coordinates of the species occurrence were recorded. Abundance was assessed in 852 one-ha plots. Structural diversity was studied using structural indices on data from ecological inventories and neighbourhood survey in 70 one-ha plots. Key results – The two hypotheses proved true. B. aethiopum was found in all phytodistricts but with strong variations in abundance. Overall, floristic composition of its natural habitats showed dissimilarities among phytodistricts. Three main vegetation types sheltered B. aethiopum : mixed grass and shrub savannas, savanna woodlands and woodlands, all of which were found in gallery forest landscapes. The density of B. aethiopum was lower in grass savannas but larger, shorter and distant individuals were found there than in savanna woodlands and woodlands. In the latter vegetation types, its density was high with thin, tall and closely spaced individuals. B. aethiopum tolerates mingling with several other tree species but increased mingling tends to lead to positive differentiation in diameter and height. Conclusions – Borassus aethiopum is a sun-demanding species and establishes successful populations in various ecological conditions. It could be mixed with other tree species in tree plantations and modern agroforestry systems as long as water requirements are met. However, it would be preferable that the species is associated with shade tolerant or medium sun-demanding species.

Influence of flow regime on the vegetation zonation along mountain streams in the Western Cape, South Africa

Zonation patterns of riparian vegetation have been sampled and described in mountain streams in two catchments in the Hottentots-Holland Mountains, Western Cape, South Africa. Six main vegetation types that differ in structure and species composition, are dominant along these river banks: Aquatic vegetation, Wetbanks, Palmiet, Scrub, Forest and Shrubland (Fynbos). The study aims to correlate the vegetation patterns to flooding patterns, in particular the inundation frequency and stream power. A problem arises: because these catchments are ungauged, like most mountain catchments, with the only weirs at the downstream end of the catchment. Discharge data at the weirs are extrapolated to the sites upstream by multiplication with a factor based on the size of the subcatchment that drains through a sample site. In this way, recurrence intervals for floods in mountain streams are derived. Discharges at sites are also calculated using bed roughness (Manning’s n) and slope in straight sections with uniform flow conditions. Stream power is derived from the discharges calculated in this manner. The combination of stream power and recurrence intervals explains the occurrence of most vegetation types occurring on the banks, except for one type: Afromontane Forest. This type is probably more dependent on other factors, such as protection from fire and the depth of the groundwater table.

Land cover changes assessment using object‐based image analysis in the Binah River watershed (Togo and Benin)

AbstractIn this study, land cover changes between 1972 and 2013 were investigated in the Binah River watershed (North of Togo and Benin) using remote sensing and geographic information system technologies. Multitemporal satellite images—Landsat MSS (1972), TM (1987), and OLI‐TIRS (2013)—were processed using object‐based image analysis and post–classification comparison methods including landscape metrics and changes trajectories analysis. Land cover maps referring to five main land cover classes, namely, agricultural land, forest land, savannah, settlements, and water bodies, were produced for each acquisition date. The overall accuracies were 76.64% (1972), 83.52% (1987), and 88.84% (2013) with respective Kappa statistics of 0.69, 0.78, and 0.86. The assessment of the spatiotemporal pattern of land cover changes indicates that savannah, the main vegetation type, has undergone the most dominant change, decreasing from 67% of the basin area in 1972 to 56% in 1987 and 33% in 2013. At the same time, agricultural land has significantly increased from 15% in 1972 to 24% in 1987 and 43% in 2013, while some proportions of agricultural land were converted to savannah relating to fallow agriculture. In total, more than 55% of the landscape experienced changes between 1972 and 2013. These changes are primarily due to human activities and population growth. In addition, agricultural activities significantly contributed to the increase in the number of patches, degree of division, and splitting index of forest and savannah vegetations and the decrease in their effective mesh sizes. These results indicate further fragmentation of forest and savannah vegetations between 1972 and 2013. Further research is needed to quantitatively evaluate the influences of individual factors of human activities and to separate these from the impacts of climate change‐driven disturbances.

Habitat and diet of Bhutan takin Budorcas taxicolor whitei during summer in Jigme Dorji National Park, Bhutan

Little is known about the ecology of the rare and vulnerable Bhutan takin Budorcas taxicolor whitei, a large and basically forest-dwelling goat-antelope inhabiting mountain valleys in northern Bhutan. In Tsharijathang valley (3800 m asl) in Jigme Dorji National Park, we described the summer habitat of this takin subspecies through vegetation transects and by studying diet via microhistological analysis of faeces supplemented by examination of feeding sites. The habitats utilized by the c.250 animals occupying the valley consisted of six main vegetation types with roughly equal coverages, viz. three forest types (dominated by Juniperus, Betula or Abies), alpine scrub (mainly low Rhododendron shrubs), open alpine meadow (mainly graminoids and forbs) and semi-open willow shrub (mainly Salix and forbs). Sixty-eight food species were identified in the feeding sites, but the bulk of the diet consisted of ≈10 species, the same species that also dominated the faecal material. With >50% of the diet consisting of shrubs, mainly Salix myrtillacea, the takins were mainly browsers. When foraging, they exhibited a weak form of selection by choosing sites with a slightly higher coverage of the most important food species and, while at these, proportionally more feeding signs were recorded. Domestic yak Bos grunniens are allowed to graze in the study area during winter when the takins have descended to lower elevation. At the present grazing pressure, the wintering yaks probably benefit the takin by maintaining the open vegetation structure, but their numbers and impact on the habitat should be monitored to prevent adverse effects.

Spatial patterns of cultural ecosystem services provision in Southern Patagonia

Context Although there is a need to develop a spatially explicit methodological approach that addresses the social importance of cultural ecosystem services for regional planning, few studies have analysed the spatial distribution on the cultural ecosystem services based on social perceptions. Objective The main objective of this study was to identify cultural ecosystem service hot-spots, and factors that characterize such hot-spots and define the spatial associations between cultural ecosystem services in Southern Patagonia (Argentina). Methods The study was carried out in Southern Patagonia (243.9 thousand km 2 ) located between 46 and 55 SL with the Andes mountains on the western fringe and the Atlantic Ocean on the eastern fringe of the study area. The study region has a range of different vegetation types (grasslands, shrub-lands, peat-lands and forests) though the cold arid steppe is the main vegetation type. We used geo-tagged digital images that local people and visitors posted in the Panoramio web platform to identify hot-spots of four cultural ecosystem services (aesthetic value, existence value, recreation and local identity) and relate these hot-spots with social and biophysical landscape features.

Dynamic Response of Satellite-Derived Vegetation Growth to Climate Change in the Three North Shelter Forest Region in China

Since the late 1970s, the Chinese government has initiated ecological restoration programs in the Three North Shelter Forest System Project (TNSFSP) area. Whether accelerated climate change will help or hinder these efforts is still poorly understood. Using the updated and extended AVHRR NDVI3g dataset from 1982 to 2011 and corresponding climatic data, we investigated vegetation variations in response to climate change. The results showed that the overall state of vegetation in the study region has improved over the past three decades. Vegetation cover significantly decreased in 23.1% and significantly increased in 21.8% of the study area. An increase in all three main vegetation types (forest, grassland, and cropland) was observed, but the trend was only statistically significant in cropland. In addition, bare and sparsely vegetated areas, mainly located in the western part of the study area, have significantly expanded since the early 2000s. A moisture condition analysis indicated that the study area experienced significant climate variations, with warm-wet conditions in the western region and warm-dry conditions in the eastern region. Correlation analysis showed that variations in the Normalized Difference Vegetation Index (NDVI) were positively correlated with precipitation and negatively correlated with temperature. Ultimately, climate change influenced vegetation growth by controlling the availability of soil moisture. Further investigation suggested that the positive impacts of precipitation on NDVI have weakened in the study region, whereas the negative impacts from temperature have been enhanced in the eastern study area. However, over recent years, the negative temperature impacts have been converted to positive impacts in the western region. Considering the variations in the relationship between NDVI and climatic variables, the warm–dry climate in the eastern region is likely harmful to vegetation growth, whereas the warm–wet conditions in the western region may promote vegetation growth.

Cambio climático y sus efectos en la vegetación de Veracruz, México: una aproximación mediante modelado de nicho ecológico

De acuerdo con el último informe del Panel Intergubernamental sobre Cambio Climático, las actividades humanas son las principales responsables del calentamiento global observado en las últimas décadas y se debe al incremento en la concentración atmosférica de gases de efecto invernadero. Este proceso ya está causando cambios en la distribución geográfica de las especies, así como alteraciones en la fenología. En tal contexto, es interesante estudiar las posibles tendencias de cambio que podrían presentar la flora y fauna para hacer frente a las nuevas condiciones del medio ambiente. En este trabajo se exploran los posibles cambios en la composición de los principales tipos de vegetación en el estado de Veracruz, México, ante un escenario de cambio climático, mediante el análisis de los patrones de alteración geográfica y ecológica de un grupo de 51 especies comúnmente asociadas al bosque tropical perennifolio, el bosque de coníferas y el bosque mesófilo de montaña, presentes en el estado. El enfoque adoptado se basa en la técnica de modelado de nicho ecológico realizado con el Algoritmo Genético para la Producción de grupos de Reglas (GARP) y variables climatológicas estimadas para un escenario A2 que corresponde a una humanidad que genera emisiones medias-altas y derivan de proyecciones hechas a partir de las condiciones actuales hacia el año 2050. Los resultados obtenidos para la superficie actual de la vegetación del estado de Veracruz sugieren una afectación potencial de 53% del área de distribución para el bosque tropical perennifolio, de 16% para el bosque de coníferas y de 49% para el bosque mesófilo de montaña. Además, apuntan a que algunas especies tienen alta probabilidad de resultar extirpadas de la entidad bajo las condiciones esperadas en el 2050, mientras que otras encontrarán las condiciones adecuadas para incrementar su distribución.

Modern pollen assemblages from surface lake sediments in northwestern China and their importance as indicators of vegetation and climate

The modern pollen assemblages of surface lake sediments and topsoils in northwestern China were studied to understand the relationship of modern pollen data with contemporary vegetation and climate, and the differences between the pollen assemblages of surface lake sediments and topsoils. The results show that Chenopodiaceae and Artemisia are dominant elements in the pollen assemblages of northwestern China. Additionally, Ephedra, Cyperaceae, Asteraceae, Poaceae, Picea, Pinus, and Betula are also important pollen taxa. Both pollen assemblages and principal component analysis indicate that pollen data from surface lake sediments and topsoils can be used to differentiate the main vegetation types of this region (desert, steppe, meadow and forest). However, differences exist between modern pollen assemblages of the two types of sediments due to the different relevant source areas of pollen and degrees of pollen preservation. For example, the larger relevant source area of surface lake sediment results in a higher abundance of Betula in pollen assemblage from surface lake sediment, whereas the tendency to disintegrate thin-walled pollen types in topsoil leads to a higher proportion of resistant pollen, such as Asteraceae. Linear regression analysis indicates that the Artemisia/Chenopodiaceae (A/C) ratio in pollen assemblages of surface lake sediments can be used to indicate humidity changes in the study area. However, the A/C ratio in topsoils should be used carefully. Our results suggest that pollen data from surface lake sediments would be better references for interpreting the fossil pollen assemblages of lake cores or lacustrine profiles.

Climate and vegetation change during the late Miocene in southwest Bulgaria based on pollen data from the Sandanski Basin

Late Miocene sediments from the Sandanski Basin in southwest Bulgaria are studied by means of the spore and pollen analysis. Vegetation in the studied basin is described and its dynamics and development are traced out. The main vegetation types are distinguished. The flora of this time interval was characterized by a change in the structure of communities and of the dominant species. Quercus, Castanea, Corylus, Ulmus, and Carya used to prevail in the composition of mesophytic forest paleocoenoses. Paleogeographic and climatic changes at the beginning of late Miocene had determined the emergence of suitable conditions for a broader participation of herbaceous coenoses. The first herbaceous coenoses were comparatively poor in their taxonomic composition, which became enriched for the first time in the late Tortonian. The traced-out dynamics of paleoclimatic data testify to notable climatic changes. The beginning of the late Miocene witnessed a certain drying and slight cooling, resulting in a moderately warm climate. It had a distinct seasonality with respect to temperature and precipitation.

Retrieval of grassland plant coverage on the Tibetan Plateau based on a multi-scale, multi-sensor and multi-method approach

Plant coverage is a basic indicator of the biomass production in ecosystems. On the Tibetan Plateau, the biomass of grasslands provides major ecosystem services with regard to the predominant transhumance economy. The pastures, however, are threatened by progressive degradation, resulting in a substantial reduction in plant coverage with currently unknown consequences for the hydrological/climate regulation function of the plateau and the major river systems of SE Asia that depend on it and provide water for the adjacent lowlands. Thus, monitoring of changes in plant coverage is of utmost importance, but no reliable tools have been available to date to monitor the changes on the entire plateau. Due to the wide extent and remoteness of the Tibetan Plateau, remote sensing is the only tool that can recurrently provide area-wide data for monitoring purposes. In this study, we develop and present a grassland-cover product based on multi-sensor satellite data that is applicable for monitoring at three spatial resolutions (WorldView type at 2–5m, Landsat type at 30m, MODIS at 500m), where the data of the latter resolution cover the entire plateau. Four different retrieval techniques to derive plant coverage from satellite data in boreal summer (JJA) were tested. The underlying statistical models are derived with the help of field observations of the cover at 640 plots and 14 locations, considering the main grassland vegetation types of the Tibetan Plateau. To provide a product for the entire Tibetan Plateau, plant coverage estimates derived by means of the higher-resolution data were upscaled to MODIS composites acquired between 2011 and 2013. An accuracy assessment of the retrieval methods revealed best results for the retrieval using support vector machine regressions (RMSE: 9.97%, 7.13% and 5.51% from the WorldView to the MODIS scale). The retrieved values coincide well with published coverage data on the different grassland vegetation types.

Global-Scale Evaluation of Roughness Effects on C-Band AMSR-E Observations

Quantifying roughness effects on ground surface emissivity is an important step in obtaining high-quality soil moisture products from large-scale passive microwave sensors. In this study, we used a semi-empirical method to evaluate roughness effects (parameterized here by the parameter) on a global scale from AMSR-E (Advanced Microwave Scanning Radiometer for EOS) observations. AMSR-E brightness temperatures at 6.9 GHz obtained from January 2009 to September 2011, together with estimations of soil moisture from the SMOS (Soil Moisture and Ocean Salinity) L3 products and of soil temperature from ECMWF’s (European Centre for Medium-range Weather Forecasting) were used as inputs in a retrieval process. In the first step, we retrieved a parameter (referred to as the parameter) accounting for the combined effects of roughness and vegetation. Then, global MODIS NDVI data were used to decouple the effects of vegetation from those of surface roughness. Finally, global maps of the Hr parameters were produced and discussed. Initial results showed that some spatial patterns in the values could be associated with the main vegetation types (higher values of were retrieved generally in forested regions, intermediate values were obtained over crops and grasslands, and lower values were obtained over shrubs and desert) and topography. For instance, over the USA, lower values of were retrieved in relatively flat regions while relatively higher values were retrieved in hilly regions.

Avifaunal diversity in the peripheral areas of the Maduruoya National Park in Sri Lanka: With conservation and management implications

A survey was randomly conducted in the marginal areas of Maduruoya National Park, Sri Lanka for a period of > 7 years. These study sites are located within the dry zone and the intermediate zone. The main vegetation type of the area is dry mixed evergreen forest. We recorded 196 bird species belonging to 66 families, and they included 161 breeding residents, 25 purely migrants, nine both resident and migrants, one vagrant, 14 nationally threatened, three globally threatened, and 10 endemic species. We also report the first-ever records of Chestnut-backed Owlet, Red-faced Malkoha, and Spot-winged Thrush from this dry area. However, these precious habitats and its species are threatened because of irresponsible human activities such as forest fires, land filings, hunting, road kills, encroachments, garbage dumping, agrochemicals, granite-rock blasting, logging, and road constructions. Therefore, we recommend that relevant authorities take immediate conservation action to increase the protection of these marginal areas or buffer zone in the near future.

Coastal land degradation and character assessment of Southern Black Sea landscape

The developing economies have negatively affected sustainability of natural resources. Increasing environmental pollution, global climate change and deforestation are some examples for such effects. Therefore, some leading countries seek for a solution to overcome these problems. Within this frame, “European Landscape Convention” which is also signed by Turkey, foresees to describe and characterize of landscapes in Pan-European level by applying protection and promotion principals for management and planning the landscapes in a sustainable way. Thus, there is an urgent need to describe and characterize our available national landscapes in any scale and resolution. In this study, landscape character assessment for the city center of Rize and adjacent rural vicinity was investigated. The study area is famous for cultivation of industrial plant “CameliaSinensis” and evidently under severe usage as agricultural fields. The results of the study revealed that the area can be described by 5 different landscape characters and 544 landscape units. Spatial analysis carried out in precisely built geo-database showed that the area is under influence of two main vegetation types which are tea plantation areas and coppice cover. Moreover, the study area is highly fragmented due to triple effects; “urbanization”, “inheritance law” and “natural land cover alteration”.

Calibration of FARSITE simulator in northern Iranian forests

Abstract. Wildfire simulators based on empirical or physical models need to be locally calibrated and validated when used under conditions that differ from those where the simulators were originally developed. This study aims to calibrate the FARSITE fire spread model considering a set of recent wildfires that occurred in northern Iranian forests. Site-specific fuel models in the study areas were selected by sampling the main natural vegetation type complexes and assigning standard fuel models. Overall, simulated fires presented reliable outputs that accurately replicated the observed fire perimeters and behavior. Standard fuel models of Scott and Burgan (2005) afforded better accuracy in the simulated fire perimeters than the standard fuel models of Anderson (1982). The best match between observed and modeled burned areas was observed on herbaceous fuel models. Fire modeling showed a high potential for estimating spatial variability in fire spread and behavior in the study areas. This work represents a first step in the application of fire spread modeling in northern Iran for wildfire risk monitoring and management.

LATE GLACIAL AND HOLOCENE ENVIRONMENTAL CHANGE INFERRED FROM THE PÁRAMO OF CAJANUMA IN THE PODOCARPUS NATIONAL PARK, SOUTHERN ECUADOR

To reconstruct the environmental history including vegetation, fire and climate dynamics, from the Cajanuma valley area ( 3285 m elevation) in the Podocarpus National Park, southern Ecuador , we address the following major research question: (1) How did the mountain vegetation developed during the late Glacial and Holocene? (2) Did fire played an important control on the vegetation change and was it natural or of anthropogenic origin?. Palaeoenvironmental changes were investigated using multiple proxies such as pollen, spores, charcoal analyses and radiocarbon dating. Pollen data indicated that during the late Glacial and transition to the early Holocene (ca. 16 000–10 500 cal yr BP) herb paramo was the main vegetation type around the study area, while subparamo and mountain rainforest were scarcely represented. The early and mid-Holocene (ca. 10 500 to 5600 cal yr BP) is marked by high abundance of paramo during the early Holocene followed by a slight expansion of mountain forest during the mid-Holocene. During the mid- to late Holocene (ca. 5600–1200 cal yr BP) there is a significant presence of paramo and subparamo while Lower Mountain Forest decreased substantially, although, Upper Mountain Forest remained relatively stable during this period. The late Holocene, from ca. 1200 cal yr BP to present, was characterized by paramo; however, mountain forest and subparamo presented significantly abundance compared to the previous periods. Fires became frequent since the late Holocene. The marked increased local and regional fire intensity during the wetter late Holocene strongly suggests that were of anthropogenic origin. During the late Glacial and early Holocene, the upper forest line was located at low elevations; but shifted slightly upslope to higher elevations during the mid-Holocene.

Utilizing Amerindian Hunters' Descriptions to Guide the Production of a Vegetation Map

Describing vegetation types is critical for managing natural resources and assessing ecosystem risk. Vegetation maps are historically produced by “Western experts,” often ignoring local-level groups critical to resource management. Indigenous hunters, as resource managers, have strong connections to their landscapes and their descriptions of vegetation within their homelands can be useful in the map-making process. This project examined the usefulness of vegetation descriptions from Rupununi, Southern Guyana Indigenous hunters in the map-making process and how their descriptions were influenced by biophysical environmental attributes. A Landsat TM and ASTER DEM merged imagery of the Rupununi was classified using Indigenous hunters' vegetation descriptions to train the classification and assess accuracy. Based on the hunters' vegetation descriptions an eleven-class map was produced that covered the main vegetation types they described. Whereas “expert” maps rely on organized forest inventory data, Indigenous hunters' vegetation classifications were influenced by their interactions with the biophysical environment. The final map shows that Indigenous hunters may be important partners in the map-making process and play key roles in tropical forest management decision-making processes.

Mid- and late Holocene vegetation and environmental dynamics in the Llanganates National Park, Anteojos Valley, central Ecuadorian Andes

The last ca. 4100 cal yr BP of palaeoenviromental conditions in the Llanganates National Park, central Ecuadorian Andes, has been reconstructed from the pollen record ‘Anteojos Valley’ (3984 m elevation). The pollen record, dated with four radiocarbon dates, indicates that the local páramo vegetation was relatively stable with only minor fluctuations since the mid-Holocene. The páramo vegetation was characterised mainly by Asteraceae, Cyperaceae and Poaceae. The regional lower mountain rainforest vegetation is mainly represented by Moraceae/Urticaceae, and the upper mountain rainforest by Melastomataceae, Polylepis and Weinmannia. Between ca. 4100 to 2100 cal yr BP, páramo was the main vegetation type with a low presence of mountain rainforest, probably reflecting cool conditions. Between ca. 2100 cal yr BP and the present, the proportion of páramo vegetation increased with a decreased occurrence of mountain rainforest, suggesting cooler and moister conditions. Low frequencies of fires were evidenced since the mid-Holocene. However, there is a slight increase of regional fire between ca. 4100 and 3100 cal yr BP. The low abundance of larger carbonised particles since the beginning of the record suggests a low occurrence of local fire in the study area.

Vegetation Succession on River Sediments along the Nakdong River, South Korea

We investigated successional vegetation on river sediments along the Nakdong River in South Korea at the time when a river regulation project, named the Four River Project, was launched. We sampled differently aged stages of succession in 63 plots of 10 × 10 m in 2010 using species cover estimations. To explain the observed species composition, we considered the following environmental variables: successional age, distance from the estuary, altitude, elevation above the river water line, texture of the substrate (silt, sand and gravel content) and cover of forest around each site within a radius of 1 km. We used ordination techniques to relate plant community characteristics to environmental variables. We distinguished three main successional vegetation types based on their floristic composition, which included especially a variety of willow species. Besides the cover of forest and certain components of substrate texture, all the variables exhibited significant effects on species composition of the vegetation over the course of succession. The most important factors determining the seral vegetation were the altitudinal gradient and successional age. Environmental factors in the riverine landscape together form a complex riverine environment and determine the vegetation pattern. Unfortunately, this environment has been largely destroyed by the massive river regulation project, but our results indicate that, under certain conditions, the riverine vegetation has a potential to at least partly recover even after such a large-scale disturbance.

Structure, spatio-temporal dynamics and disturbance regime of the mixed beech–silver fir–Norway spruce old-growth forest of Biogradska Gora (Montenegro)

The structure and the spatio-temporal dynamics of the mixed beech–silver fir–Norway spruce old-growth forest of Biogradska Gora (Montenegro) have been analysed at different spatial scales: at the landscape scale, using a high-resolution SPOT5 satellite image and at the stand level with an intensive field survey. This remote-sensing approach has been used to obtain a land cover map in order to define the main vegetation types and to detect the large canopy gaps (>150 m2). The structural characteristics have been delineated in a 50-ha study area in which a regular 120-m grid was superimposed over a 1:10,000 raster map and 30 sampling points have been obtained. The forest is characterized by a high volume of living trees (1029.6 m3 ha− 1) and coarse woody debris (420.4 m3 ha− 1) and by small-scale disturbances (individual trees to small groups) with a low incidence of intermediate disturbances (18 forest canopy gaps>150 m2 over 1230 ha). The two approaches have proved useful to delineate the spatio-temporal dynamics. The Biogradska Gora forest dynamics are dominated by very small-scale processes, which are partially autogenic and partially caused by allogenic factors. The influence of large-scale or intermediate disturbances has shown to be negligible.

Classification of Korean forests: patterns along geographic and environmental gradients

AbstractAimTo describe the main vegetation types of the Korean zonal forests and explore their relationship with the main environmental and geographic gradients.LocationKorean Peninsula.MethodsA data set of 3847 relevés of zonal (semi)natural forest stands, together with environmental factors recorded in the field or derived from interpolated climatic data was collected. Using the hierarchical ISOPAM method, the relevés were classified into main forest types and their species composition, diversity and distribution were described. Their differentiation along main environmental gradients was analysed with classification and regression trees.ResultsEleven vegetation types of forests were distinguished and assigned to five vegetation classes: Quercetea mongolicae (representing the core of the Korean temperate forests), Vaccinio‐Piceetea (subalpine coniferous forest), Fagetea crenatae (rare beech‐dominated forest included within a sub‐montane and sub‐oceanic type of oak‐dominated forest), Querco mongolicae‐Betuletea davuricae (sub‐continental drought‐ and fire‐tolerant forest, differentiated along elevational gradient) and Camellietea japonicae (oceanic evergreen forest). At a higher hierarchical level, these 11 vegetation types were grouped into three clusters, which most probably reflect different vegetation development in the Late Pleistocene and Holocene. Within the three major vegetation clusters, each forest type occupies a distinct environment. The forest types are especially well separated along elevational and temperature gradients. The distribution of individual forest types is joinly driven by total amount of incoming energy and thermal extremes, both affecting water supply to plant communities.ConclusionsAll distinguished vegetation types can be assigned to the recent syntaxonomic classification system. However, three main vegetation clusters do not correspond to traditionally distinguished syntaxonomic units. Nevertheless, the forest types within these clusters share common species with similar ecology and, especially, migration histories during the late glacial and post‐glacial periods. Therefore, we interpret these clusters as biogeographic patterns reflecting vegetation history rather than the current environmental conditions.