This paper presents a comprehensible neural network tree (CNNTREE). CNNTREE is a proposed general modular neural network structure, where each node in this tree is a comprehensible expert neural network (CENN). One advantage of using CNNTREE is that it is a “gray box”; because it can be interpreted easily for symbolic systems; where each node in the CNNTREE is equivalent for symbolic operator in the symbolic system. Another advantage of CNNTREE is that it can be trained as any normal multi layer feed forward neural network. An evolutionary algorithm is given for designing the CNNTREE. Back propagation is also checked as local learning algorithm that fits for real time learning constraints. The tree generalization and training performance are examined using experiments with a digit recognition problem.