Summary In recent years considerable attention has focused on metamorphic core complexes of the Basin and Range Province of the western US Cordillera. These highly extended areas are characterized by an upper plate that has been brittlely attenuated by normal faults separated by a sub-horizontal detachment surface from a lower plate that has been ductilely thinned and stretched. A study of mesoscopic structures, finite strain, microstructures, quartz c -axis fabrics and 40 Ar/ 39 Ar geochronology was undertaken in order to characterize the nature, geometry, kinematic history and timing of ductile extension in the lower plate of the northern Snake Range metamorphic core complex in E-central Nevada. These data provide new insights into the processes of deep-seated ductile strain beneath supracrustal normal fault mosaics in highly extended regions. Mesoscopic structures and finite strain analyses indicate that the lower plate underwent plane strain with a sub-vertical Z-axis and a sub-horizontal WNW-ESE X -axis. The magnitude of strain increases dramatically from W-to-E, and is constant vertically. Bedding and foliation are everywhere parallel and bedding is thinned 30–90%. The nature and geometry of microstructures and c -axis fabrics changes progressively from W-to-E. Lower strain rocks on the W flank of the range are characterized by a single bedding-parallel foliation, defined by flattened detrital grains, globular grains which have their c -axes parallel to the Z -axis and symmetrical cross-girdle c -axis fabrics. Further E to slightly higher strains, the rocks are characterized by asymmetrical cross-girdle c -axis fabrics, although globular grains with their c -axes parallel to Z are still present. The high-strain rocks on the E flank of the range are characterized by C -planes, asymmetrical mica ‘fish’ and oblique quartz foliations, and asymmetrical single-girdle c -axis fabrics. These data suggest that the lower plate has not deformed entirely by either coaxial or non-coaxial strain, and we propose an evolutionary model whereby the lower plate deformed during an early period of coaxial strain followed by a later component of non-coaxial strain accompanied by coaxial strain on the E flank of the range. 40 Ar/ 39 Ar geochronology on lower-plate hornblendes and micas indicates that temperatures of deformation were >280°C, but <530°C, and increased with depth. Geochronological data suggest that lower-plate ductile strain is post-latest Cretaceous, and biotite and muscovite data indicate that ductile deformation was probably ongoing 22–26 Ma. Our data and regional geological relations suggest that ductile extensional deformation in the northern Snake Range occurred because of localized heat input to shallow levels of the crust, and together with seismic reflection data, indicate that lower-plate rocks probably represent the top of a regional metamorphic terrane of Tertiary age.