Lectotype Specimen Research Articles

The genus Toyoides Matsumura (Hemiptera: Auchenorrhyncha: Delphacidae): new combination and lectotype designations

In this study, we revised the planthopper genus Toyoides Matsumura (Hemiptera: Auchenorrhyncha: Delphacidae) to include three species. The genus Epunka Matsumura be synonymized with Toyoides based on habitus and genital characteristics. The lectotype specimens have been designated for both T. albipennis and T. bilineata comb. n. Morphological information on T. bilineata comb. n. is presented, with photographs of the habitus and male genitalia provided. We have also provided the keys for the identification of related genera and species.

Virtual reconstruction of the Canis arnensis type (Canidae, Mammalia) from the Upper Valdarno Basin (Italy, Early Pleistocene)

Taphonomic deformation, whether it be brittle or plastic, is possibly the most influential process hindering the correct understanding of fossil species morphology. This is especially true if the deformation affects type specimens or applies to or obscures taxonomically diagnostic or functionally significant traits. Target Deformation, a recently developed virtual manipulation protocol, was implemented to address this issue by applying landmark-guided restoration of the original, deformed fossils, using undeformed specimens (or parts thereof) of the same species as a reference. The enigmatic Early Pleistocene canid Canis arnensis provides a typical example of a fossil species in dire need of virtual restoration. Its lectotype specimen is heavily deformed and none of the few known skulls are well preserved, obscuring the recognition of its systematic and phylogenetic position. Our results indicate that the algorithm effectively countered the lectotype skull’s laterolateral compression and its concomitant rostrocaudal elongation. Morphometrically, comparison of the retrodeformed cranium (IGF 867_W) with other specimens of the same species, and to other fossil and extant canid material, confirms IGF 867_W consistently clusters within C. arnensis variability. Overall, the evidence presented here confirms that Target Deformation provides a powerful tool to better characterize complex taxa like C. arnensis, whose knowledge is severely affected by the state of preservation of its fossil material.

On the lectotype designation of Bryconamericus rubropictus (Berg) (Characiformes, Characidae, Stevardiinae).

The lectotype specimen of Bryconamericus rubropictus (Berg) and its designation have remained imprecisely documented since its publication. The lack of a photograph or an unambiguous illustration, correct size, inaccurate labelling, and proper specimen separation has led to an uncertainty about the identity and nomenclatural status of the lectotype. We recovered and provided detailed morphological data on the specimen that must be recognised as the lectotype. This contribution brings stability and clarity on the nomenclatural status of the species and its type material.

Towards a nomenclatural clarification of the Peltigera ponojensis/monticola clade including metagenomic sequencing of type material and the introduction of P. globulata Miadl. & Magain sp. nov.

AbstractPeltigera globulata Miadl. & Magain, a new species in the P. ponojensis/monticola species complex of section Peltigera, is formally described. This clade was previously given the interim designation Peltigera sp. 17. It is found in sun-exposed and xeric habitats at high altitudes in Peru and Ecuador. Peltigera globulata can be easily recognized by its irregularly globulated margins covered mostly by thick, white pruina, somewhat resembling the sorediate thallus margins of P. soredians, another South American species from section Peltigera. The hypervariable region of ITS1 (ITS1-HR), which is in general highly variable among species of section Peltigera, does not have diagnostic value for species identification within the P. ponojensis/monticola complex. Nevertheless, no significant level of gene flow was detected among eight lineages representing a clade of putative species (including P. globulata) within this complex. ITS sequences from the holotype specimens of P. monticola Vitik. (collected in 1979) and P. soredians Vitik. (collected in 1981) and lectotype specimens of P. antarctica C. W. Dodge (collected in 1941) and P. aubertii C. W. Dodge (collected in 1952) were successfully obtained through Sanger and Illumina metagenomic sequencing. BLAST results of these sequences revealed that the type specimen of P. monticola falls within the P. monticola/ponojensis 7 clade, which represents P. monticola s. str., and confirmed that the type specimen of P. aubertii falls within a clade identified previously as P. aubertii based on morphology. The ITS sequence from the type specimen of P. soredians, which superficially resembles P. globulata, confirms its placement in the P. rufescens clade. Finally, we discovered that the name P. antarctica was erroneously applied to a lineage in the P. ponojensis/monticola clade. The ITS sequence from the type specimen of P. antarctica represents a lineage within the P. rufescens clade, which is sister to the P. ponojensis/monticola clade.

On seven undescribed leaf insect species revealed within the recent "Tree of Leaves" (Phasmatodea, Phylliidae).

With the recent advance in molecular phylogenetics focused on the leaf insects (Phasmatodea, Phylliidae), gaps in knowledge are beginning to be filled. Yet, shortcomings are also being highlighted, for instance, the unveiling of numerous undescribed phylliid species. Here, some of these taxa are described, including Phylliumiyadaonsp. nov. from Mindoro Island, Philippines; Phylliumsamarensesp. nov. from Samar Island, Philippines; Phylliumortizisp. nov. from Mindanao Island, Philippines; Pulchriphylliumheraclessp. nov. from Vietnam; Pulchriphylliumdelisleisp. nov. from South Kalimantan, Indonesia; and Pulchriphylliumbhaskaraisp. nov. from Java, Indonesia. Several additional specimens of these species together with a seventh species described herein, Pulchriphylliumanangusp. nov. from southwestern India, were incorporated into a newly constructed phylogenetic tree. Additionally, two taxa that were originally described as species, but in recent decades have been treated as subspecies, are elevated back to species status to reflect their unique morphology and geographic isolation, creating the following new combinations: Pulchriphylliumscythe (Gray, 1843) stat. rev., comb. nov. from Bangladesh and northeastern India, and Pulchriphylliumcrurifolium (Audinet-Serville, 1838) stat. rev., comb. nov. from the Seychelles islands. Lectotype specimens are also designated for Pulchriphylliumscythe (Gray, 1843) stat. rev., comb. nov. and Pulchriphylliumcrurifolium (Audinet-Serville, 1838) stat. rev., comb. nov. from original type material.

(2952) Proposal to conserve the name Coccidioides posadasii against Posadasia esferiformis, Coccidium posadasi, Pseudococcidioides mazzae, Geotrichum louisianoideum, Glenospora meteuropea, Glenospora metamericana and Trichosporon proteolyticum (Ascomycota)

(2952) Proposal to conserve the name <i>Coccidioides posadasii</i> against <i>Posadasia esferiformis</i>, <i>Coccidium posadasi</i>, <i>Pseudococcidioides mazzae</i>, <i>Geotrichum louisianoideum</i>, <i>Glenospora meteuropea</i>, <i>Glenospora metamericana</i> and <i>Trichosporon proteolyticum</i> (<i>Ascomycota</i>)

Redescription of Neocarus platensis Silvestri, 1905 (Acari: Opilioacaridae) and a new species from the syntype series

Neocarus platensis is redescribed from part of the syntype series collected in Argentina and lectotype specimens are designated. The remaining syntype material from Uruguay represents a new species, herein named Neocarus paraplatensis sp. nov. Taxonomic problems within the Opilioacaridae are discussed in light of a comparative morphological discussion including setae quantity variation.

Reappraisal and lectotypification of Justicia tanalensis S. Moore (Acanthaceae), rediscovered from central Madagascar more than 100 years since the last collection

Justicia tanalensis (Acanthaceae) was rediscovered from the central highland of Madagascar since the last collection in 1895. Its morphological characters are reported and amended with photographs in this paper. The species is characterized by a prostrate herbaceous habit and a simple spike with one flower per node. It has a distribution range in the central-eastern Madagascar. The conservation assessment of the species is provided. Because no holotype was indicated in the protologue and the type specimens cited by the original author were collected in two different times, they should be considered as two gatherings and they are syntypes. The lectotype specimen of the name is designated here as well.

New species of Plutellidae from Iran (Lepidoptera: Yponomeutoidea)

Three new species of Plutellidae are described from Iran: Baerenschenkia lamasi Mey, sp. n., Deryaxenistis nekrutenkoi Mey, sp. n., and Paraxenistis ammolofon Mey, sp. n. Adults and male and female genitalia are illustrated. The male genitalia of two lectotype specimens, deposited in the Natural History Museum, London were examined and illustrated: Paraxenistis sphenospila Meyrick, 1919 and P. pentaula (Meyrick, 1913). They are provisionally retained in Paraxenstis Meyrick, 1919.

Typification of Linnaean names in the Grewioideae (Malvaceae)

AbstractTypes are designated for five species of Grewioideae (Malvaceae) described by Linnaeus. These include three lectotypes and one neotype of names of Corchorus species and one lectotype of the name of a Triumfetta species. Three of the lectotypes are illustrations and one is a specimen in Linnaeus's herbarium. In addition, an epitype for T. lappula is designated to facilitate the interpretation of the lectotype specimen, which lacks flowers that are diagnostic for the unambiguous application of the name.

A Proposal for the Lectotype Designation of Ishige foliacea (Phaeophyceae, Ishigeaceae) Using DNA Barcoding

Three species of the genus Ishige (Phaeophyceae, Ishigeaceae) are known from Korea, Japan and Mexico; they include Ishige foliacea Okamura, I. okamurae Yendo and I. sinicola (Setchell and N.L. Gardner) Chihara. Two species, I. foliacea and I. okamurae, are present in the algal flora of Korea and Japan. The original description of I. okamurae defined two forms of branches, filiform and foliose, but later the foliose branch was recognized as a new species I. foliacea, which is epiphytic on I. okamurae but can also be free-living. The currently proposed lectotype for I. foliacea is based on a free-living form and does not reflect the intent of the original description of the species. In this study, we conducted the DNA barcoding for herbarium specimens to identify Ishige species. Additionally, the variation in cox3 sequences obtained from I. okamurae specimens with two morphological forms collected from waters around the Korean Peninsula was sufficient to separate two species, I. okamurae and its epiphyte I. foliacea. The epiphytic I. foliacea on the lectotype specimen of I. okamurae is designated as lectotype I. foliacea.

Doubts about the validity of the species name Hipparchia hermione (Linnaeus, 1764), it being associated with the two species Hipparchia alcyone ([Denis &amp; Schiffermüller], 1775) and Hipparchia genava (Fruhstorfer, 1908) following the designation of a lectotype by Kudrna (1977) (Lepidoptera: Nymphalidae, Satyrinae) - First part -

In 1977, Otakar Kudrna (*1939-†2021, obituary see Balletto and Leigheb, 2021) published his “Revision of the Genus Hipparchia”, where he classified all the known species and forms of this genus according to characters of wings, androconia, male genital armatures and further subjective criteria. Until today, Kudrna’s study is considered as the guideline of systematics of the genus Hipparchia. He selected there a lectotype specimen of a Rock Grayling male in the Linnaean collection. “Hipparchia hermione Linnaeus, 1764” is therefore, at the moment, the technically correct name to identify the species. The “International Commission of Zoological Nomenclature” (ICZN) has nothing to add at this point; it only comments on cases submitted to it in the Bulletin of Zoological Nomenclature. Within the meaning of the present study and in accordance with Verity (1913), this damaged specimen without abdomen represents the same species as Ignaz Schiffermüller – allegedly the only author of the Vienna directory (see Kudrna and B., 2005, p. 5) – has described under H. alcyone from the Vienna region by referring to a coloured copper engraving published by Rösel von Rosenhof (1755). Kudrna’s “Revision” became the starting point of an extended scientific research activity during my free time containing, as a matter of priority, the examination of problem cases of systematics by checking the preimaginal characters of many rearing series from different sites. It turned out that a number of classifications proposed by Kudrna (1977) had to be reassessed as soon as characters of the pre-imaginal stages were available. The most complex case I have verified concerned the third European Grayling species which Leraut (1990) introduced under the name of Hipparchia genava (Fruhstorfer, 1908). Kudrna (1977) failed to separate this species from H. alcyone (D. &amp; S., 1775). Throughout his life, he never agreed with Leraut’s opinion. In recent years, Kudrna had hoped that genetic examinations would make redundant every rearing attempt by amateur lepidopterists and furnish the proof that his opinion was the correct one. Since he never undertook any rearing experiments, his systematics were based only on prepared imagines being housed in museum collections. He saw himself as a person with the competency to decide within a few minutes upon complex questions of taxonomy and ignored completely the assessments of others. For verifying the effective rank of H. genava, I had to examine also the two most closely related species: Hipparchia fagi and H. alcyone by rearing them all ex ovo with material from several widely spaced sites. Already on finalising my rearing work of this group, it became apparent that Leraut (1990) had been on the absolutely right path by accepting a third Grayling species, within this group. For the first time, I published the results of my rearing attempts between 2002 and 2006 in several articles in the quarterly bulletin Linneana Belgica and I readily provided information on this case to interested colleagues. Over time only, I realized that the Rock Grayling I knew from the volume on butterflies (Diurna) by Forster and Wohlfahrt (1955) as from the guides by Higgins and Riley (1970-84) under the name of H. alcyone had become H. hermione, because of the lectotype designation by Kudrna (1977). A stony path was in front of me to substantiate the factual correctness of the view taken by Leraut (1990). Complex clarifications by Peter Russell furnished well-founded arguments on the complex scientific issue why the use of the name “hermione Linnaeus, 1764” should be rejected for the Rock Grayling previously known as H. alcyone.

Morphology of the lectotype of Chelodina oblonga Gray 1841 (Testudines: Chelidae).

Uncertainty about taxonomy and nomenclature of the Australian long-necked turtle Chelodina oblonga has ensued since its description by Gray in 1841. This has been due to the general nature of information about provenance of the lectotype specimen and the ostensible morphological similarity of the specimen to another taxon from northern Australia that confused Gray and many subsequent workers. The south-western Australian provenance of the specimen has recently been confirmed by Shea et al. (2020). The present paper provides a detailed consideration of morphology of the specimen and compares it with other specimens available to Gray and to larger series that are now available. The study elucidates morphological aspects that have underlain the history of confusion and it highlights a number of characters that consistently differentiate adults of the relevant taxa. These demonstrate that morphology of the C. oblonga lectotype conforms with the taxon from the south-west of Western Australia, as has also been confirmed by recently published investigations of mitogenomics and historical information about provenance of the lectotype.

Type specimen sequencing, multilocus analyses, and species delimitation methods recognize the cosmopolitan Corallina berteroi and establish the northern Japanese C. yendoi sp. nov. (Corallinaceae, Rhodophyta).

A partial rbcL sequence of the lectotype specimen of Corallina berteroi shows that it is the earliest available name for C. ferreyrae. Multilocus species delimitation analyses (ABGD, SPN, GMYC, bPTP, and BPP) using independent or concatenated COI, psbA, and rbcL sequences recognized one, two, or three species in this complex, but only with weak support for each species hypothesis. Conservatively, we recognize a single worldwide species in this complex of what appears to be multiple, evolving populations. Included in this species, besides C. ferreyrae, are C. caespitosa, the morphologically distinct C. melobesioides, and, based on a partial rbcL sequence of the holotype specimen, C. pinnatifolia. Corallina berteroi, not C. officinalis, is the cosmopolitan temperate species found thus far in the NE Atlantic, Mediterranean Sea, warm temperate NW Atlantic and NE Pacific, cold temperate SW Atlantic (Falkland Islands), cold and warm temperate SE Pacific, NW Pacific and southern Australia. Also proposed is C. yendoi sp. nov. from Hokkaido, Japan, which was recognized as distinct by 10 of the 13 species discrimination analyses, including the multilocus BPP.

Genomic analysis of the lectotype specimens of European Ulva rigida and Ulva lacinulata (Ulvaceae, Chlorophyta) reveals the ongoing misapplication of names

ABSTRACT Species of Ulva Linnaeus are nearly impossible to identify using morpho-anatomy due to their simple thallus structure and phenotypic plasticity. The current solution to this problem is to sequence DNA from field-collected specimens and match these sequences to those available in public DNA databases. However, because type specimens of many species have not been sequenced, the accuracy of these identifications is highly doubtful. Ulva rigida C.Agardh is reported to be one of the most widespread and ecologically important Ulva species, but these records are based on either morpho-anatomy or, more recently, on DNA sequences. High-throughput sequencing (HTS) was performed on the lectotype specimen of U. rigida from Cádiz, Spain to determine the correct application of the name. The analysis yielded its complete plastid genome. rbcL, tufA and ITS sequences from the lectotype specimen differed at the species level from all U. rigida sequences deposited in public databases. Instead, the lectotype sequences of U. rigida were identical or very similar to sequences identified as U. rotundata Bliding (referred to by some as U. pseudorotundata Cormaci, G.Furnari & Alongi) from Ireland and Portugal, but not to the holotype of U. rotundata from Italy, which was identical to U. lactuca L. HTS of the lectotype of U. lacinulata (Kützing) Wittrock from Lesina, Croatia, a species morphologically similar to U. rigida with macroscopic marginal teeth, also yielded a complete plastid genome, with sequences identical or highly similar to GenBank U. armoricana Dion, Reviers & Coat, U. ‘laetevirens’, U. ‘rigida’ and U. scandinavica Bliding. Since U. lacinulata is the oldest validly published name, it is the correct one to apply to the globally distributed species that was previously but incorrectly known as U. rigida. Based on this genetic evidence, U. rigida is restricted to European waters and confirmed by DNA sequences from Ireland, Portugal and Spain. This analysis shows that many barcode species identifications and taxonomic conclusions in the genus Ulva are incorrect.

Taxonomic revisions based on genetic analysis of type specimens of Ulva conglobata, U. laetevirens, U. pertusa and U. spathulata (Ulvales, Chlorophyta)

SUMMARYGenetic analysis of type material of various Ulva species has shown that many names have been misapplied. DNA sequences of two of the specimens on the lectotype sheet of U. australis from South Australia showed that U. australis is conspecific with specimens called U. pertusa from Japan. However, type material of U. pertusa has not been sequenced, nor have other type specimens with type localities in Japan and Australia, including U. conglobata f. conglobata and U. conglobata f. densa from the former, and U. laetevirens and U. spathulata from the latter. To determine if these names have been correctly applied, targeted DNA sequencing of the rbcL gene was performed on (i) the lectotype specimen of U. laetevirens, (ii) the lectotype and two syntype specimens of U. pertusa, (iii) the lectotype and two syntype specimens of U. congoblata f. conglobata and (iv) two specimens on the holotype sheet of U. conglobata f. densa. In addition, high‐throughput sequencing (HTS) analysis was carried out on an isotype specimen of U. spathulata. Phylogenetic analysis of the resulting rbcL gene sequences supports the taxonomic conclusions that U. laetevirens, U. pertusa and U. spathulata are synonyms of U. australis, and U. conglobata f. conglobata and U. conglobata f. densa represent a single separate and distinct species, U. conglobata. These genetic data contribute to the taxonomy of Ulva and further demonstrate that analyzing DNA from type material is the only currently known reliable method to correctly apply names in this genus.

Australia's prehistoric 'swamp king': revision of the Plio-Pleistocene crocodylian genus Pallimnarchus de Vis, 1886.

The crocodylian fossil record from the Cenozoic of Australasia is notable for its rich taxonomic diversity, and is primarily represented by members of the clade Mekosuchinae. Reports of crocodylian fossils from Australia date back to the late nineteenth century. In 1886, Charles Walter de Vis proposed the name Pallimnarchus pollens for crocodylian fossils from southeast Queensland—the first binomen given to an extinct crocodylian taxon from Australia. Pallimnarchus has come to be regarded as a large, broad-snouted crocodylian from Australia’s Plio-Pleistocene, and numerous specimens, few of which are sufficiently complete, have been assigned to it by several authors throughout the twentieth century. In the late 1990s, the genus was expanded to include a second species, Pallimnarchus gracilis. Unfortunately, the original syntype series described as Pallimnarchus pollens is very fragmentary and derives from more than one taxon, while a large part of the subsequently selected lectotype specimen is missing. Because descriptions and illustrations of the complete lectotype do not reveal any autapomorphic features, we propose that Pallimnarchus pollens should be regarded as a nomen dubium. Following this decision, the fossil material previously referred to Pallimnarchus is of uncertain taxonomic placement. A partial skull, formerly assigned to Pallimnarchus pollens and known as ‘Geoff Vincent’s specimen’, possesses many features of diagnostic value and is therefore used as basis to erect a new genus and species—Paludirex vincenti gen. et sp. nov. A comprehensive description is given for the osteology of ‘Geoff Vincent’s specimen’ as well as aspects of its palaeoneurology, the latter being a first for an extinct Australian crocodyliform. The newly named genus is characterized by a unique combination of premaxillary features such as a distinctive arching of the anterior alveolar processes of the premaxillae, a peculiar arrangement of the first two premaxillary alveoli and a large size disparity between the 3rd and 4th premaxillary alveoli. These features presently allow formal recognition of two species within the genus, Paludirex vincenti and Paludirex gracilis comb. nov., with the former having comparatively more robust rostral proportions than the latter. The Paludirex vincenti holotype comes from the Pliocene Chinchilla Sand of the Darling Downs, south-eastern Queensland, whereas the material assigned to Paludirex gracilis is from the Pleistocene of Terrace Site Local Fauna, Riversleigh, northwest Queensland. Phylogenetic analyses recover Paludirex vincenti as a mekosuchine, although further cladistic assessments are needed to better understand the relationships within the clade.

The ‘red-tailed’ Lasioglossum (Dialictus) (Hymenoptera: Halictidae) of the western Nearctic

Sweat bees in the subgenus Lasioglossum (Dialictus) are one of the most diverse and abundant bee taxa, and a critically important component of bee biodiversity. Yet, the most basic taxonomic knowledge of these bees is lacking in many regions. As a step towards a better understanding of the L. (Dialictus) of the western Nearctic region, a revision of the ‘red-tailed’ L. (Dialictus) species was completed. Thirty-six species were revised, 20 of which are described as new, and two names are treated as junior subjective synonyms. Descriptions, figures, distribution maps, floral hosts, and keys to species for females and males are provided. The following 20 species are described as new: Lasioglossum (Dialictus) arenisaltans sp. nov., L. (D.) argammon sp. nov., L. (D.) austerum sp. nov., L. (D.) cactorum sp. nov., L. (D.) cembrilacus sp. nov., L. (D.) clastipedion sp. nov., L. (D.) clavicorne sp. nov., L. (D.) decorum sp. nov., L. (D.) festinum sp. nov., L. (D.) imbriumbrae sp. nov., L. (D.) julipile sp. nov., L. (D.) lilianae sp. nov., L. (D.) meteorum sp. nov., L. (D.) miltolepoides sp. nov., L. (D.) minckleyi sp. nov., L. (D.) perditum sp. nov., L. (D.) rufornatum sp. nov., L. (D. ) spivakae sp. nov., L. (D.) tessellatosum sp. nov., and L. (D.) torrens sp. nov. Previously unknown males of L. (D.) clematisellum (Cockerell, 1904), L. (D.) droegei Gibbs, 2009, L. (D.) kunzei (Cockerell, 1898), and L. (D.) pallidellum (Ellis, 1914) are described and figured for the first time. Lasioglossum (Dialictus) clarissimum (Ellis, 1914) (= Halictus clarissimus Ellis, 1914) and L. (D.) perexiguum (Sandhouse, 1924) (= Halictus (Chloralictus) perexiguus Sandhouse, 1924) are new subjective junior synonyms of L. mesillense (Cockerell, 1898) (= Halictus nymphalis var. mesillensis Cockerell, 1898). A lectotype specimen is newly designated for L. mesillense, for which the location of the type material has not previously been known. The following five new records for Mexico are reported: L. clematisellum, L. droegei, L. eophilus (Ellis, 1914), L. kunzei, and L. pallidellum.

A new large-bodied Pliocene seal with unusual cutting teeth.

Today, monachine seals display the largest body sizes in pinnipeds. However, the evolution of larger body sizes has been difficult to assess due to the murky taxonomic status of fossil seals, including fossils referred to Callophoca obscura, a species thought to be present on both sides of the North Atlantic during the Neogene. Several studies have recently called into question the taxonomic validity of these fossils, especially those from the USA, as the fragmentary lectotype specimen from Belgium is of dubious diagnostic value. We find that the lectotype isolated humerus of C. obscura is too uninformative; thus, we designate C. obscura as a nomen dubium. More complete cranial and postcranial specimens from the Pliocene Yorktown Formation are described as a new taxon, Sarcodectes magnus. The cranial specimens display adaptations towards an enhanced ability to cut or chew prey that are unique within Phocidae, and estimates indicate S. magnus to be around 2.83 m in length. A parsimony phylogenetic analysis found S. magnus is a crown monachine. An ancestral state estimation of body length indicates that monachines did not have a remarkable size increase until the evolution of the lobodontins and miroungins.

Studi Literasi Taksonomi dan Penelusuran Spesimen Lektotipe Cengkih (Syzygium aromaticum (L.) Merr. &amp; Perry)

Old scientific names of clove, Caryophyllus aromaticus, was published in 1753 by Carrolus Linnaeus and up to now it has many synonyms. Now, the old name is still recognized as basionim of clove. Merrill &amp; Perry, based on the character, subsequently revised the genus of clove from the Caryophyllus to Syzygium, so the accepted scientific name is Syzygium aromaticum. The lectotype specimen was designed by McVaugh in 1989 and stored in BM from the Clifford collection.