Observations of foraging patterns are essential to understanding the energetic ecology of marine animals. However, direct observations are not often feasible in the field because most foraging takes place below the sea surface or at great distances from land. Attachment of data-recording devices to animals can greatly facilitate the collecton of meaningful data. Devices are being used to an increasing extent on marine mammals (Kooyman et al. 1976, Ray et al. 1978), birds (Kooyman et al. 1971, 1982, Adams and Brown 1983, Lishman and Croxall 1983, Wilson and Bain 1984a, b), reptiles (Stoneburner 1982), and fish (Voegeli and Piucock 1980, Priede 1983a). These devices may supply data upon recovery (e.g., Kooyman et al. 1971, 1982) or through telemetry to local receivers (e.g., Voegeli and Piucock 1980, Priede 1983a, b) or to orbiting satellites (Stoneburner 1982). The effects of instrument mass and harness attachment on land animals are considered to have the potential of affecting the outcomes of experiments and observations (Dumke and Pils 1973, Gilmeretal. 1974, Perry 1981, Perryetal. 1981). However, the effects of devices on marine animals, which live in a much more viscous medium than air, have not been evaluated. In this study we investigated the effects of attached instruments on the foraging activity of the African Penguin (Spheniscus demersus). We show that instrument size may adversely affect foraging behavior, but that data obtained using devices of varying sizes can be used to back-calculate true foraging parameters of free-swimming penguins without devices. Data-recording devices are especially important in studies of free-swimming penguins because these birds are inconspicuous on the surface of the water and are not visible during their frequent traveling and foraging dives. Instruments have been constructed to record speed (Wilson and Bain 1 984b), foraging range (Wilson and Bain 1984b, Wilson and Achleitner 1985), dive time (Trivelpiece et al., in press), dive frequency and depth (Kooyman et al. 1982, Wilson and Bain 1984a). Results show that African Penguins are capable of sustained speeds of at least 7.5 km/h (Nagy et al. 1984) and that other penguin species forage at depths exceeding 200 m (Kooyman et al. 1982). Measurements of foraging distance have also been used to estimate the energy cost of swimming and to construct energy budgets (Nagy et al. 1984). Devices may affect performance in two ways. The mere presence of a device may modify behavior. However, in our experience with penguins this does not appear to be important; after a short period of adjustment, African Penguins with devices continued their normal nesting activity and entered the sea to forage with frequencies similar to control birds (Wilson and Bain 1984a, b). Second, the mass or drag of a device may retard swimming speed, thus reducing dive depth, foraging range, and the number of prey encountered. Instrument mass in itself probably does not adversely influence foraging behavior because attached devices are usually .05) which suggests that the smallest meters had virtually no effect on foraging behavior. On the other hand, two penguins fitted with an electronic distance meter 14.6% of penguin
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