The diversity and ubiquity of insects on the planet Earth is a commonplace premise of every entomology textbook. From the tropics to Antarctica, from deserts and hot pools to high arctic islands, and from open ocean to mountain peaks, insects have found unorthodox niches where other arthropods and microorganisms are commonly their sole companions. Their great capacity for dispersal and their relatively small size, which together at least partially explain insect diversity (71), enable them to find and exploit habitats in which they are' protected from macroscale environmental factors. In addition to the benefits of scale, various insects have the physiological capacity to survive ambient temperatures that exclude most Metazoa, salinities that defy most regulatory systems, and/or ability to tolerate deep-freezing and desiccation. Of all the eccentric habitats colonized by insects, perhaps the most extreme are those where the nonnal pattern of energy flow from photosynthetic plants is denied, as in areas of extreme aridity, e.g. deserts (64) and lava fields (47); cold, e.g. alpine (65, 66) and polar regions (39); or dark, e.g. caves (20, 49, 69). These environments, like ocean abysses where dwellers depend on organic material filtering down from the photic zone, are all examples of what Hutchinson (53) termed the allobiosphere, where resident organisms depend for their subsistence on transported energy and nutrients. Swan (92) drew attention to the widespread distribution of such environments in alpine regions and coined the term aeolian for them, thereby explicitly recognizing the role of wind in the transport of nutrients. The emphasis of this review is on alpine communities, but it is not only in
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