Grazing Cycle Research Articles

Pasture characteristics of Italian ryegrass and milk production under different management strategies

The objective of this work was to assess the effects of the sward structure of Italian ryegrass (Lolium multiflorum), during the first grazing cycle, on its morphological and bromatological characteristics throughout the growing season, and on the performance of dairy cows. The treatments consisted of two structures obtained as a function of canopy-light interception: high-light interception (HLI) and low-light interception (LLI), with different pre-grazing heights in the first grazing cycle. Pasture was managed under rotational grazing with a herbage allowance not below 30 kg dry matter (DM) per cow per day. Three grazing cycles, with a grazing interval of 30 days, were evaluated. Pre-grazing herbage mass was greater (2,240 vs. 1,656 kg ha-1 DM), but the proportion of leaf blades was smaller (0.35 vs. 0.43) for HLI swards. Neutral detergent fiber (NDF) content and organic matter digestibility (OMD) were similar between treatments in the first grazing cycle, but in the second and third ones NDF was greater, and OMD lower, for the HLI swards. Milk yields were greater for cows grazing LLI swards (19.4 vs. 21.1 kg per day). Initial grazing with 90% of light interception promotes greater nutritional value in the subsequent cycles.

Frequencies and intensities of defoliation in Aruana Guineagrass swards: accumulation and morphological composition of forage

The objective of this study was to assess the accumulation and morphological composition of forage in Aruana Guineagrass (Panicum maximum cv. Aruana) swards subjected to intermittent stocking with sheep. Experimental treatments when grazing was introduced included low (95%) and high (98%) incident light interception and low (10 cm)- and high (15 cm)-residue height and were allocated to experimental units (196 m2 fenced areas) in a completely randomized 2 × 2 factorial arrangement with three replicates. Treatments were imposed between January and May 2009. More frequent grazing cycles (95% light interception) resulted in better control of stem elongation, lower proportion of dead plant material and invasive plants, and higher proportions of leaf blades in the grazing strata, compared with less frequent grazing cycles (98% light interception). Grass managed with 95% light interception combined with 10 and 15 cm post-grazing height and grass managed at 98% light interception combined with 15 cm post-grazing height did not show differences in forage accumulation rate. These results indicate that more frequent (30 cm pre-grazing height) and less severe (15 cm post-grazing height) grazing cycles provided animals with high leaf blade mass and low stem mass forage.

Tillering dynamics in pastures of guinea grass subjected to grazing severities under intermittent stocking

This experiment was carried out to analyze the tillering dynamics of the species Panicum maximum cv. Mombaca subjected to three post-grazing heights: residue of 30 cm (30); residue of 50 cm (50); and residue of 50 cm during spring and summer, lowered to 40 cm in the first fall season grazing and to 30 cm in the following grazing cycle, resuming to 50 cm after the first grazing of the following spring season (50-30). Grazings were initiated whenever the swards intercepted 95% of the incident light. The post-grazing heights were allocated in the experimental units in a completely randomized block design with three replications. The density of basal tillers did not vary between the residual heights evaluated. Swards managed with variable residual height (50-30) presented higher rates of appearance and mortality of basal tillers during the summer of 2007, indicating high tiller renovation. Regardless of the post-grazing height evaluated, lower rates of appearance of basal tillers were found in the spring of 2006. The stability index of guinea grass cv. Mombaca was close to 1.0 throughout the experimental period. Swards managed with variable post-grazing present structural changes able to improve the regrowth vigor, which may be important to maximize the use of the forage species in the production system.

Herbage Mass, Herbage Rejection, and Chemical Composition of Signalgrass under Different Stocking Rates and Distances from Dung Pads

ABSTRACTNutrients from fecal return may improve herbage growth and chemical composition, but herbage rejection may occur around dung pads. Rejected herbage increases in maturity and might be rejected in future grazing cycles. The aim of this study was to determine the influence of dung pad on soil and herbage characteristics of signalgrass [Urochloa decumbens (Stapf) R. D. Webster (syn. Brachiaria decumbens Stapf.)] pastures managed under three stocking rates (1.9, 3.2, and 4.2 animal units (AUs) ha−1; 1 AU = 450 kg of cattle live weight). Herbage and soil responses were measured at different distances from the dung pad border (0 to 20, 20 to 40, and 40 to 60 cm). A rotational stocking was applied with 3 d of grazing and 32 or 67 d of resting for the dry and rainy seasons, respectively. Herbage mass increased near the dung pad by 40% compared to the surrounding areas but was not affected by stocking rate. Herbage rejection was greater near the dung pad and rejection was greater at lower stocking rate. Plants distant from the dung pad presented lower P concentration. Regardless of stocking rate, K, N, and P fecal and herbage concentrations were affected by the grazing cycle, and there was a higher nutrient concentration during the rainy season. Soil beneath dung pad had higher pH value (5.6 vs. 5.2), Mehlich‐I P (22.4 vs. 12.5 g m−3), and K concentration (9 vs. 5 mol m−3) compared to soil from the surrounding areas. Grazing management practices should improve fecal distribution in pastures to maximize nutrient return via this pathway.

CARACTERÍSTICAS ESTRUTURAIS E BROMATOLÓGICAS DO CAPIM TANZÂNIA SOB PASTEJO ISOLADO, SIMULTÂNEO E ALTERNADO DE OVINOS COM BOVINOS

The aim of this study was to evaluate the effect of three types of rotational pasture: isolated (I – grazing only by sheep), simultaneous (S – simultaneous grazing by cattle and sheep) and alternated (A – granzing by cattle and then by sheep) on the structural and chemical traits of grass. Twelve crossbreed heifers, weighing initially 207 kg, 30 Santa Ines lambs, weighing 23 kg, and 16 adult ewes weighing 47 kg were used. The following parameters were determined on the pasture: forage mass available per grazing cycle (FMC); proportion of leaf (PF); stalk (PS); dead material (PDM), leaf:stalk ratio (L/S); levels of dry matter (DM); mineral matter (MM); crude protein (PB); ether extract (EE); neutral detergent fiber (NDF); acid detergent fiber (ADF); inorganic phosphorus (Pi) and total digestible nutrients (TDN). The S and I systems presented the smallest values for PS and NDF, while their values for L/S, CP, EE and TDN were higher than in A system. Therefore, the results showed that the grass offered to sheep in S and I systems had better quality than in the A system. There was not effect of systems and cycles of pasture on the others parameters evaluated. Tanzania grass available to sheep showed best structural and chemical traits in S and I systems while it presented worse quality in A system. The S system may be an alternative to traditional sheep rearing systems in the Brazilian savannah region. KEYWORDS: grass; pasture; ruminant.

Adjustment of Animal Growth Rate Responses in Repeat Measurement Grazing Trials

In animal-response grazing trials there are sources of uncertainty in one-period one-off measurements, which as partial factorisation over time, become traceable and quantifiable sources of variation in repeat measurement trials. This is illustrated for a trial comparing sheep and goat live-weight gains under two stocking rates on mixed species pastures established by three contrasting sowing methods. It used variable plot size to give uniform animal numbers and tracked changes in individual animal performance and pasture growth in different periods. It was repeated on the same plots and animals over 17 grazing cycles. The variation explainable was greater with growth rates expressed as percent live-weight increase per day, than as weight or metabolic weight increase per day. The base data sets were adjusted for specific weighing-day effects of estimated gut-fill using moving averages, and for calibration for individual animal effects by genotype/environment analysis. Collectively these significantly increased the proportion explainable by 3.1-3.8% in variance analyses using qualitative treatment variables, and 2.7-3.7% in response function analyses relative to measured climate, pasture, plot and collective animal covariates. Simulation studies, to allow for variability in the independent variables as well as the dependent variables, indicated that the proportion explainable could increase by a further 0-1.2% and 1.1-1.9% respectively for the variance or response function approaches.

Evolução de atributos físicos de solo em sistema de integração lavoura-pecuária

O objetivo deste trabalho foi avaliar o efeito das alturas de pastejo e dos sucessivos ciclos de pastejo sobre os atributos físicos do solo, em um sistema de integração lavoura-pecuária. O experimento foi implantado em 2001, na região do Planalto Médio, RS, em um Latossolo Vermelho, com o cultivo consorciado de azevém (Lolium multiflorum) e de aveia-preta (Avena strigosa), sob pastejo contínuo, no inverno, e o cultivo de soja (Glycine max) no verão. Os tratamentos consistiram de diferentes intensidades de pastejo, definidas pela altura da pastagem (10, 20, 30 e 40 cm), tendo-se utilizado uma área sem pastejo como controle. Foram avaliadas a densidade e a porosidade do solo após o ciclo de pastejo e de cultivo da soja, bem como a resistência mecânica do solo à penetração e a estabilidade de agregados no sétimo ano do experimento. Não houve alterações significativas na densidade e na porosidade do solo após sete anos em integração lavoura-pecuária. A resistência do solo à penetração é maior na camada superficial após o ciclo de pastejo. A agregação do solo aumenta nas áreas pastejadas, independentemente da intensidade de pastejo.

Performance of dairy heifers in a silvopastoral system

The characteristics of the pasture and performance of dairy heifers were evaluated in a silvopastoral system (SPS) established with Brachiaria decumbens grass in a consortium with four tree species (105trees/ha) and in a monoculture system of Brachiaria decumbens (MS). Thirty-two paddocks (0.5ha/paddock) were used, sixteen for each system (SPS and MS). The paddocks were managed under rotational stocking, with seven days occupation and 35 and 45days of rest during the rainy and dry seasons, respectively. The treatments were distributed in a randomised block design with two repetitions. The heifers used had an initial weight of 200kg and a variable stocking rate as a function of the herbage allowance of 7.0kg of DM/100kg of body weight/day. The forage mass was estimated at each grazing cycle. For each sample, the DM, crude protein and neutral detergent fibre contents were determined as well as the in vitro dry matter digestibility (IVDMD). All animals were weighed every 35days to estimate daily body weight gain and weight production per area. The forage dry mass, stocking rate and herbage allowance were similar between the systems studied, but they varied according to season. Average values of 2031 and 1100kg/ha/grazing cycle, 1.6 and 0.8AU (Animal unit=450kg of body weight)/ha and 7.2 and 5.0kg DM/100kg of body weight were estimated for the rainy and dry seasons, respectively. The values for crude protein content were higher in the SPS in the rainy season, while the NDF and IVDMD did not vary with raising system. The body weight gains per animal and per hectare in the dry season did not vary, but were higher in the SPS during the rainy season in the first and third experimental years. The annual body weight gains were also higher in SPS than in the MS. It was concluded that the SPS is more efficient for rearing dairy heifers than the MS due to the higher body weight gain per heifer and per area.

Performance and productivity of Nellore steers on rotationally stocked palisadegrass (Brachiaria brizantha) pastures in response to herbage allowance

SUMMARYHerbage allowance (HA) impacts performance of grazing cattle as well as sward structure and herbage yield. The objective of the current study was to compare average daily gain (ADG) of Nellore steers on Marandu palisadegrass (Brachiaria brizantha (Hochst ex A. Rich.) Stapf. cv. Marandu) pastures in response to HA. Treatments were applied as 5, 10, 15 and 20 kg dry matter (DM) per 100 kg live weight (LW) per day HA5, HA10, HA15 and HA20), in rotationally stocked pastures under a 35-day grazing cycle (28 days of rest and 7 days of grazing). The trial was carried out in Pirassununga, Brazil, during two summer seasons in 2003 and 2004, each divided into four grazing cycles. Individual animal (250 kg yearling steers) gain increased with HA up to HA15 (mean=0·69 kg/head per day) during each of four grazing cycles per year. The 2-year gain per area was greater for HA5 and HA10 than for HA15 and HA20 and average stocking rate was 9·6, 6·7, 5·4 and 4·6 head/ha for HA5 to HA20, respectively. In addition to the effects of HA on ADG, changes in sward structure, especially sward height, help to explain variation in daily gain in both years. The HA10 treatment provides higher gain per area without excessively reducing individual animal performance, and thus may be the level of choice for optimizing animal output in pasture-only systems.

Changes in nematode abundances and body length in response to management intensive grazing in a low-input temperate pasture

An evaluation of changes in nematode populations in response to management was performed using a field scale experiment along a gradient of land-use intensity in the context of management intensive grazing. Four levels of decreasing intensity were measured over three growing seasons. The treatments included: 1) intensive grazing management where cows were allowed to graze for each grazing cycle, followed by routine clipping and harrowing; 2) semi-intensive grazing where cows were allowed to graze for each grazing cycle, with only one clipping event per year; 3) extensive grazing where cows were allowed to graze during every 2nd grazing cycle and no clipping or harrowing was performed; 4) stockpiling treatment where cows were permitted to graze only during the final grazing cycle, followed by clipping at the end of the season. Most indices (MI, PPI, ∑MI, and ∑MI25) resulted in significant seasonal differences; however, only familial diversity (H′) and the total number of families showed significant season by treatment interaction. Bacterivores comprised most of the nematode population with Cephalobidae being the most common family. Abundance of plant feeders was affected by season and soil moisture, whereas bacterivore abundance was associated with total organic N and soil pH. Nematode body length increased slightly in most families through time, and was associated with season, bulk density, soil moisture, and total N and C. Although omnivore numbers were highest in the extensive and semi-intensive treatments, there was a significant increase in dorylaimid body length observed in the intensive treatment through time. Increases in body length were also observed in Monhysteridae and Cephalobidae in all treatments.

Característica produtiva e estrutural de pastos mistos de aveia e azevém manejados em quatro alturas sob lotação contínua

Monitoraram-se variáveis relacionadas à produção e à estrutura de pastos de aveia-preta (Avena strigosa, Schreb) e azevém (Lolium multiflorum, Lam) manejados em quatro alturas (10, 20, 30 e 40 cm), em pastejo contínuo com lotação variável (put and take). Utilizaram-se novilhos de corte mestiços de aproximadamente 10 meses de idade com peso médio inicial de 190 kg, em delineamento de blocos completos casualizados com três repetições. Investigou-se o tipo de distribuição das frequências de altura em seis datas, bem como o potencial de predição da massa seca de forragem por meio da altura do pasto. A massa de forragem aumentou de forma linear de acordo com a altura do pasto: cada 1 cm de aumento na altura acima de 10 cm correspondeu a um acréscimo de cerca de 107 kg de MS/ha na massa de forragem. Não houve efeito da altura do pasto na taxa de acúmulo (55,8 kg de MS/ha.dia) nem na produção total de matéria seca (8.210 kg de MS/ha). A distribuição das frequências ajustou ao modelo normal em apenas uma das 96 séries analisadas. A distribuição tipo Gamma foi a que mais frequentemente se ajustou aos dados de altura do pasto, porém, uma vez iniciado o pastejo, a heterogeneidade no pasto foi tão alta que a distribuição das alturas não se ajustou a nenhum dos modelos estudados. É possível estimar a massa seca de forragem existente no pasto por meio de sua altura média, assim, sugere-se estabelecer alturas variáveis ao longo do ciclo de pastejo, com o intuito de se administrar a heterogeneidade causada pelo animal.

Soil aggregation in a crop-livestock integration system under no-tillage

Grazing intensities can influence soil aggregation, which can be temporarily and permanently affected. The objective of this study was to evaluate the aggregate stability in water at the end of a soybean cycle and during pasture development in a crop-livestock integration system under no-tillage and grazing intensities. The experiment was initiated in 2001, in a dystrophic Red Latosol, after soybean harvest. Treatments consisted of pasture (black oat + Italian ryegrass) at heights of 10, 20 and 40 cm, grazed by young cattle, and a control (no grazing), followed by soybean cultivation, in a randomized block design. Soil samples were collected at the end of the soybean cycle (May/2007), during animal grazing (September/2007) and at the end of the grazing cycle (November/2007). The grazing period influences aggregate distribution, since in the September sampling (0-5 cm layer), there was a higher proportion of aggregates > 4.76 mm at all grazing intensities. Soil aggregation is higher in no-tillage crop-livestock integration systems in grazed than in ungrazed areas.

Produção de forragem e carga animal de pastagens de Coastcross sobressemeadas com forrageiras de inverno

O objetivo nesta pesquisa foi avaliar a produção de forragem e a carga animal de quatro pastagens: Coastcross (Cynodon dactylon L. Pers.) sobressemeada com azevém (Lolium multiflorum Lam. cv. Comum); Coastcross sobressemeada com azevém e trevo-branco (Trifolium repens L., cv. Yi); azevém e trevo branco; e azevém. O experimento foi realizado no período entre 15 de maio e 24 de outubro de 2006, quando foram realizados cinco ciclos de pastejo. Utilizaram-se vacas da raça Holandesa recebendo concentrado (3,5 kg/dia) como suplemento alimentar. Nos períodos pré e pós-pastejo, foram avaliadas a massa de forragem, as composições botânica e estrutural da pastagem e a carga animal. O delineamento experimental foi inteiramente casualizado com quatro tratamentos (pastagens), duas repetições (piquetes) e cinco períodos de avaliação (pastejos). Não foram detectadas diferenças entre pastagens para as médias de massa de forragem e carga animal. As pastagens exclusivamente de azevém tiveram maiores produções de lâminas foliares de azevém. As pastagens sobressemeadas apresentaram maior produção de forragem. A sobressemeadura do azevém e do trevo-branco em Coastcross permite estender o período de utilização da pastagem anual e aumenta a produção de forragem.

Repetibilidade e respostas de características morfofisiológicas e produtivas de capim-elefante de porte baixo sob pastejo

O objetivo deste trabalho foi estimar os coeficientes de repetibilidade, de determinação (R²) e o número mínimo de medições necessárias para predizer o valor real de variáveis morfofisiológicas e produtivas, bem como as respostas do capim-elefante de porte baixo ao pastejo com ovinos. O experimento foi realizadoem blocos ao acaso, com cinco tratamentos - Taiwan A-146 2.27, Taiwan A-146 2.37, Taiwan A-146 2.114, Merker México MX 6.31 e Mott - e quatro repetições, ao longo de cinco ciclos de pastejo entre setembro de2008 e abril de 2009. Foram avaliados a altura média de plantas, o índice de área foliar (IAF), a interceptação luminosa (IL), os ângulos foliares médios (AFM), o acúmulo de forragem (AF) e a taxa de acúmulo de forragem(TAF). Para as variáveis IAF, IL e altura média de plantas, nos ciclos longos, foi necessária apenas uma medição (R² = 80%) para classificá-las como de mais fácil predição em clones de capim-elefante. Avaliações de AF e de TAF necessitam de períodos experimentais mais longos, para maior confiabilidade na estimativa dessas variáveis resposta. Dosséis de capim-elefante de porte baixo interceptam maior quantidade de radiação incidente, quanto maior for a altura média das plantas e o IAF.

Milk production potential of two ryegrass cultivars with different total non-structural carbohydrate contents

The aim of the study was to compare a new Italian ryegrass (Lolium multiflorum) cultivar (Enhancer), bred to contain a high total non-structural carbohydrate content, with the cultivar, Dargle, in terms of dry matter (DM) production, nutritional value, carrying capacity and milk production. The ryegrass cultivars were sown (25 kg/ha) under supplementary irrigation in a randomized block design (7 blocks with 2 paddocks, 4.5 ha/treatment) on the 2nd May 2001 (year 1) and the 15th March 2002 (year 2) on an Estcourt soil type. Nitrogen was applied at 56 kg N/ha after each grazing. Grazing started on 26 June 2001 and the grazing cycle varied from 24 to 28 days. Pasture yield was estimated with a rising plate pasture meter. Forty Jersey cows from 50 to 150 days in milk were randomly allocated to one of the two treatments. The experimental period started on 22 August 2001 and consisted of an adaptation period of 15 days followed by a measurement period of 75 days. Pasture was allocated at 10 kg DM/cow/day above 30 mm. Cows were weighed and condition scored on two consecutive days at 14:00 at the start and the end of the experimental period. Milk production was recorded daily and milk composition was determined every 14 days. All cows were fed a flat rate of 3.6 kg DM of a dairy concentrate (120 g crude protein (CP)/kg DM, 11.5 MJ ME/kg DM) per day. The concentrate was fed in two equal portions during milking at 06:00 and 15:00. The total production of Enhancer was higher at 8438 and 9084 kg DM/ha in 2001 and 2002, respectively, compared to 7570 and 7694 kg DM/ha of Dargle. The DM and total non-structural carbohydrate content of Enhancer was higher than Dargle in 2001 but not in 2002. In 2002 the CP content of Enhancer was lower than that of Dargle. Enhancer increased 4% fat corrected milk production by 1.3 and 1.4 kg/cow/day in 2001 and 2002, respectively, and DM intake by 1.1 and 0.88 kg/cow/day compared to Dargle. The total milk production per hectare of Enhancer was 1499 kg and 2277 kg higher during 2001 and 2002, respectively, compared to Dargle. Enhancer, a high sugar Italian ryegrass, demonstrated good potential to increase milk production.

Atributos estruturais e produtivos de capim-marandu em resposta à suplementação alimentar de bovinos e a ciclos de pastejo

O objetivo deste trabalho foi avaliar as alterações estruturais e produtivas de capim-marandu (Brachiaria brizantha cv. Marandu) em resposta à suplementação alimentar de bovinos e a ciclos de pastejo. Os níveis de suplementação com concentrado à base de polpa cítrica, milho, farelo de soja e ureia foram 0,2, 0,6 e 1% do peso vivo. O delineamento experimental foi o de blocos ao acaso, com arranjo em parcelas subdivididas no tempo, com os níveis de suplementação nas parcelas e os ciclos de pastejo nas subparcelas. Os efeitos da suplementação alimentar dos animais sobre o pasto foram detectados sobre o índice de área foliar, interceptação luminosa e relação folha/colmo, em pré-pastejo. Os menores valores foram observados na suplementação de 0,2% do peso vivo. No pós-pastejo, a massa de folhas e a relação folha/colmo foram menores com a menor suplementação (0,2%). Os ciclos de pastejo afetaram as características produtivas e estruturais do pasto, bem como os índices morfogênicos do capim-marandu. Apenas a massa de material morto aumentou, enquanto o número e o peso de perfilhos não foi afetado pelos ciclos de pastejo. O pasto de capim-marandu foi influenciado pela suplementação alimentar dos bovinos. Com o suceder dos ciclos de pastejo, do verão para o outono, há redução do crescimento do pasto, com prejuízos à sua estrutura.

Desempenho de bezerros da raça Nelore e cruzados desmamados recebendo concentrado em pastagem adubada de Cynodon dactylon cv. Coastcross

Em três anos de pesquisa, avaliou-se o desempenho de bezerros desmamados de quatro grupos genéticos (Nelore; Canchim × Nelore; Angus × Nelore; e Simental × Nelore) em pastagem de Cynodon dactylon, cv. Coastcross, adubada, sob manejo rotacionado, recebendo mistura mineral ou concentrado. O concentrado foi fornecido na quantidade de 3 kg/animal/dia e continha 18,8% de proteína bruta e 81,5% de nutrientes digestíveis totais na matéria seca. A quantidade de forragem disponível diferiu com a utilização de mistura mineral e concentrado (2.961 kg e 3.383 kg de matéria seca (MS) por hectare, respectivamente). A oferta de MS/animal/dia foi 9,9 kg/dia (3,9% PV) para mistura mineral e 9,0 kg/dia (3,3% PV) para concentrado. A forragem disponível possuía, na matéria seca, 13,6% de proteína bruta, 79,8% de fibra em detergente neutro; 62,3% de digestibilidade in vitro da matéria seca, 3,9 g/kg de cálcio, 2,0 g/kg magnésio, 2,5 g/kg de fósforo, 22,7 g/kg de potássio, 2,8 g/kg de enxofre, 9,9 mg/kg de cobre, 22,5 mg/kg de zinco, 98 mg/kg de manganês e 188 mg/kg de ferro. Foram observadas interações grupo genético × suplementação e ano × suplementação para ganho diário de peso e taxa de lotação das pastagens e ciclo de pastejo × suplementação para a taxa de lotação. O ganho diário de peso nos animais Nelore, Canchim × Nelore, Angus × Nelore e Simental × Nelore criados com suplementação do pasto com mistura mineral foi de 0,48; 0,63; 0,68 e 0,50 kg, enquanto naqueles que receberam concentrado foi de 0,87; 0,95; 0,99 e 0,95 kg, respectivamente. As médias estimadas das taxas de lotação das pastagens foram 6,1 para todos os grupos genéticos que receberam mistura mineral e 7,5; 7,6; 8,8 e 9,0 unidades animais/ha para os grupos genéticos Nelore, Canchim × Nelore, Angus × Nelore e Simental × Nelore que receberam concentrado, respectivamente. O grupo genético influencia o desempenho de bovinos em pastagem de capim-coastcross e interage com a suplementação com concentrado.

Robustness of Grazing-sliding Bifurcations in Hybrid Systems

In the paper we consider how orbits characterised by the presence of so-called sliding motion, which orbits typically occur in hybrid systems of Filippov type, are affected by stable singular perturbations. To be able to pursuit our analysis we consider a planar minimal system and we tune a system parameter such that a stable periodic orbit of the system touches the discontinuity surface: this is the so-called grazing-sliding bifurcation. We then check the effect of stable singular perturbation on the grazing cycle. In the unperturbed system the periodic orbit remains stable, and its local return map becomes piecewise linear. The effect of an arbitrarily small stable singular perturbation is that the local return map changes qualitatively, giving rise to, for example, period-adding cascades or small-scale chaos.

Comportamento ingestivo de cordeiras em pastagem consorciada de inverno sob diferentes intensidades de desfolha

Avaliou-se a relação entre a estrutura do pasto, as variáveis ambientais e o comportamento ingestivo de cordeiras Ile de France × Texel em pastagem consorciada de aveia-preta (Avena strigosa Schreb.), azevém (Lolium multiflorum Lam.) e trevo-vermelho (Trifolium pratense L.) sob quatro intensidades de desfolha (muito alta, alta, média e baixa). Utilizaram-se pastejos intermitentes considerando a soma térmica de 300 graus-dia como critério para determinar os intervalos de pastejo. As avaliações do comportamento ingestivo foram feitas por meio de observação visual em quatro períodos contínuos de 24 horas realizados no período de maio a outubro de 2006. O maior tempo de pastejo ocorreu na intensidade baixa. Altas intensidades de desfolha ocasionaram aumento da densidade populacional de perfilhos de azevém e não alteraram o peso desses perfilhos. A massa de bocados e a qualidade da dieta selecionada pelas cordeiras foram semelhantes entre as intensidades testadas. Em pastagem de aveia+azevém+trevo-vermelho, quando o intervalo de pastejo é determinado pela soma térmica de 300 graus-dia, a utilização de intensidades de desfolha que variam de baixa a muito alta não altera o comportamento ingestivo de cordeiras. O comportamento ingestivo é afetado pelo ciclo do pasto e pelo fotoperíodo e a seletividade dos animais reduz ao final do período de utilização da pastagem, quando ocorre aumento no tempo de pastejo e na distância percorrida em busca de locais de alimentação.

Influence supplementary feeding on bark browsing by red deer (Cervus elaphus) under experimental conditions

The effect of various supplementary feeds on bark browsing was studied in red deer under controlled conditions. Fifteen female red deer were randomly assigned to five dietary treatments (n = 3 animals each): meadow hay as a control (C), combinations of meadow hay with grass silage (GS), maize silage (MS), maize silage with oats (MSO) or a pelleted concentrate (PC). The deer were kept in enclosures where we offered fresh spruce stems daily. The experiment consisted of two periods. In the first period (A), the animals had free access to food during 24 h, while in the second period (B), the access was restricted to night hours only. In both periods, the supplementary feeds were available ad libitum. Mean daily consumption of browsed bark in the control group in period A was 1,053 g while in the remaining groups it ranged from six to 20 g. In period B, simulating the conditions in disturbed hunting areas, bark consumption in all groups increased considerably, the highest increase being found in groups GS (430 vs. 19 g/day) and MS (101 vs. 6 g/day). In period A, the highest dry matter (DM) intake including browse was found in group MS (2,816 g/day) and the lowest one in group GS (2,307 g/day). In period B, the highest (2,763 g/day) and the lowest (2,153 g/day) DM intake was observed in groups PC and MSO, respectively. If red deer is fed supplementary, meadow hay should be combined with maize silage, grass silage or concentrate to reduce bark browsing in forests. The bark browsing rate may increase considerably if red deer are disturbed or their grazing cycle is impaired.