The structural history of the Hercynian Pyrenees is considered with respect to the relationships between the metamorphism and the successive deformations, the characteristics and mode of emplacement of the gneissic and anatectic cores of the metamorphic domes, the migmatization and the intrusions of plutonic and basic and ultrabasic rocks. The metamorphism can be shown to have progressed upwards with time, attaining the presently exposed levels just before, during and a short time after the major penetrative regional deformation. The intrusion of major plutonic, anatectic and gneissic massifs, which has determined the metamorphic domes in country rocks, was roughly synchronous with the progression of the metamorphism, but seems to have continued later in the uppermost levels. The basic and ultrabasic rocks uplifted by the intrusion of the anatectic cores of the metamorphic domes have been deformed and foliated by the major regional penetrative deformation, thus showing that their intrusion predated this deformation in the lowermost levels. These results, together with the presently available sedimentological data, lead us to propose a dynamic model of the tectonic evolution of the Hercynian Pyrenees. The first step was a crustal extension and thinning, attaining its maximum during Frasnian time, marked in the upper levels by a “ horst and graben” controlled sedimentation and in the lower crust by intrusions of basic magmas and the onset of the very low-pressure metamorphism and anatexis characteristic of the Hercynian Pyrenees. The second step was a crustal shortening marked by large-scale crustal thrusting followed by widespread regional penetrative deformation. The flysch troughs formed in front of the progressing and emerging thrusts. In lower levels, the thrusts are believed to have controlled the progressive ascent of the plutonic and anatectic magmas which continued their uplift and emplacement during major penetrative regional deformation in the actual mesozone and probably after it in the uppermost levels.