Abstract On 28 April 2019, hourly forecasts from the operational High-Resolution Rapid Refresh (HRRR) model consistently predicted an isolated supercell storm late in the day near Dodge City, Kansas, that subsequently was not observed. Two convection-allowing model (CAM) ensemble runs are created to explore the reasons for this forecast error and implications for severe weather forecasting. The 40-member CAM ensembles are run using the HRRR configuration of the WRF-ARW Model at 3-km horizontal grid spacing. The Gridpoint Statistical Interpolation (GSI)-based ensemble Kalman filter is used to assimilate observations every 15 min from 1500 to 1900 UTC, with resulting ensemble forecasts run out to 0000 UTC. One ensemble only assimilates conventional observations, and its forecasts strongly resemble the operational HRRR with all ensemble members predicting a supercell storm near Dodge City. In the second ensemble, conventional observations plus observations of WSR-88D radar clear-air radial velocities, WSR-88D diagnosed convective boundary layer height, and GOES-16 all-sky infrared brightness temperatures are assimilated to improve forecasts of the preconvective environment, and its forecasts have half of the members predicting supercells. Results further show that the magnitude of the low-level meridional water vapor flux in the moist tongue largely separates members with and without supercells, with water vapor flux differences of 12% leading to these different outcomes. Additional experiments that assimilate only radar or satellite observations show that both are important to predictions of the meridional water vapor flux. This analysis suggests that mesoscale environmental uncertainty remains a challenge that is difficult to overcome. Significance Statement Forecasts from operational numerical models are the foundation of weather forecasting. There are times when these models make forecasts that do not come true, such as 28 April 2019 when successive forecasts from the operational High-Resolution Rapid Refresh (HRRR) model predicted a supercell storm late in the day near Dodge City, Kansas, that subsequently was not observed. Reasons for this forecast error are explored using numerical experiments. Results suggest that relatively small changes to the prestorm environment led to large differences in the evolution of storms on this day. This result emphasizes the challenges to operational severe weather forecasting and the continued need for improved use of all available observations to better define the atmospheric state given to forecast models.