Coffee ringspot virus (CoRSV) and Citrus leprosis virus C (CiLV-C) are two emergent pathogens that can cause significant losses to coffee and citrus, respectively. Recently, it was verified that Brevipalpus phoenicis, originally reported as vector of these viruses, is indeed a complex group reclassified into eight mite species (Beard et al. 2015). Among those, B. papayensis Baker (Acari: Tenuipalpidae) occurs in citrus, but seems to be prevalent in coffee plantations. However, it is still unclear the relationship (if any) of B. papayensis with CiLV-C or CoRSV. In order to determine whether or not this mite is capable of transmitting these viruses, one single female specimen was collected from Coffea arabica cv. Catuai in Atibaia, SP-Brazil, and multiplied in laboratory as an isoline. Specimens were mounted in Hoyer’s medium and morphologically confirmed by phase contrast microscopy as B. papayensis, according to the classification of Beard et al. (2015). This isoline population was reared onto healthy coffee leaves, in 25 ± 1°C, 14 h light/10 h dark and 60 ± 10% humidity. RT-PCR assays using primers for the detection of each of the viruses were performed in order to confirm the virus-free colony status. B. papayensis mites were transferred to either coffee leaves exhibiting ringspots or sweet orange fruits symptomatic for leprosis, which served as sources of inocula for 7 days. Viral acquisition was confirmed by RT-PCR of viruliferous mites using specific primers for the detection of CoRSV (Kitajima et al. 2011) or CiLV-C (Locali et al. 2003), according to Kubo et al. (2011). After the acquisition period, 60 mites from each source of inoculum were transferred to 20 Arabidopsis thaliana plants (6 mites per plant), a plant species that can host both. Fifteen days after infestation, typical pinpoint lesions were observed and virus presence was confirmed by RT-PCR in 60% of the CiLV-C- and 30% of the CoRSV-inoculated plants. Additionally, common bean plants (Phaseolus vulgaris cv. Una) were infested with 15 B. papayensis mites from the isoline population reared onto CiLV-C inoculum (one mite per leaf, isolated by entomological glue). Perhaps due to the low efficiency in single mite transmission, only 27% of the leaves became symptomatic. The presence of CiLV-C was confirmed by RT-PCR and sequencing (98% of similarity to GenBank accession no. KP3367461). Fifteen females were collected directly from coffee leaves with CoRSV symptoms in Cordeiropolis, SP-Brazil, and transferred to isolated leaves of common bean plants. Symptoms of necrotic local lesions appeared 10 days after inoculation in 50% of the plants. The presence of CoRSV was confirmed by RT-PCR in both coffee and bean leaves. Amplicons obtained from the latter were sequenced (QG979998) and found 98% identical to CoRSV sequences available in GenBank. This is the first report of common bean as an experimental host of CoRSV. B. yothersi has been shown to transmit cileviruses. B. californicus seems to be the only vector of the dichorhavirus Orchid fleck virus, while B. phoenicis sensu stricto is the vector of the tentative dichorhavirus Citrus leprosis virus N. This is the first confirmation that B. papayensis can transmit CiLV-C and CoRSV. This is also the first time, following the reclassification of the B. phoenicis group, that a transmission was demonstrated by a single species of Brevipalpus for both a dichorha- and a cilevirus. Since this mite species is prevalent in coffee plantations in Brazil, it is likely the main vector of CoRSV under natural conditions.