We characterized 716 sites (including 22 beds, 60 foraging and 604 movement sites) used by red deerCervus elaphus xanthopygus Milne-Edwards, 1867 in the Wandashan Mountains, northeastern China in the winter period. We used covariates for vegetation, topography, disturbances by other ungulates, and disturbance by humans to develop movement, forage and bed site resource-selection models. We used an information-theoretic approach to select the top 5 models for movement, forage and bed site occurrences respectively. The three most parsimonious autologistic models were good predictors of movement, forage and bed occurrence of this species. Vegetation covariates were important components of all models. We recorded avoidance responses by the behaviours of species to each disturbance type (ie, villages, forest roads and abandoned roads); bed site occurrences were most sensitive to all human disturbances for their lowest odds encountered. Across all parsimonious models, villages have the largest negative effect on movement, foraging and bed occurrences. Movement and bedding behaviours were also affected by other ungulate disturbances. Altitude had slightly negative effect on movement and foraging behaviours. Ridges, topographic aspect and shrub stands were all correlated with bed-site selection. Although different behaviours may be associated with different microhabitat features, behaviours of red deer responded similarly to the same human disturbance on a broad scale. Based on the observations above, we believe that red deer are avoiding human-altered habitat for movement, bedding and foraging because of disturbances during the critical late winter period. Furthermore, various behavioural resource selection models and corresponding graphs of important habitat disturbances can be used to guide and evaluate future development proposals.
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