ABSTRACT The Peace-Athabasca Delta (PAD) is a complex, dynamic, and misunderstood ecosystem. This study uses landscape, ecological, climatic, hydrologic, historical, stratigraphic, and wildlife data that document how and why the PAD has changed and varied from ~1900 to 2023. The ecological, climatic, and hydrologic components of the ecosystem vary in concert. Flood-drawdown cycles influenced by physical and biological factors have driven a host of landscape changes over the past 120 years. Prior to regulation, net drying of the PAD took place over the period ~1900 to the mid-1940s. Since mid-20th century, there has been no multidecadal drying trend but rather decadal-scale wet and dry episodes. Levels of the confluent Lakes Athabasca, Claire, and Mamawi, and their adjacent restricted basins, are strongly correlated with the combined flows of the Peace, Athabasca, Fond du Lac, and Birch Rivers. River regulation has changed the seasonal distribution of flows and suppressed summer flows. There is, however, no evidence that ‘dramatic landscape change is underway, devastating local fauna’ or that river regulation has caused delta desiccation, declines in spring flooding, declines in lake levels, or declines in wildlife, waterfowl, or their habitat. The great areal extent of the PAD watershed imparts hydrologic and ecological resilience.