Intercanopy bare patches are small, relatively circular communities with an exposed central bedrock, shallow soil, low plant cover, and dominated by non-vascular plants, annual grass- es, and forbs. Although these patches are relatively common in the Edwards Plateau of central Texas, their characteristics have never been described. Twelve circular, intercanopy bare patches in the Hill Country State Natural Area in Bandera and Medina counties, were examined using the line-point procedure. Mean cover (?SD) of plant, litter, rock, and bare soil was 23 - 3%, 23 ? 5%, 44 ? 9%, and 10 ? 4%, respectively. Approximately half of the total plant relative cover consisted of forbs with 25% grasses and 25% non-vascular plants. Fifty-seven species of flowering plants from 30 families were identified, including 27 annuals and 30 perennials. The most com- mon families were the Poaceae (9 species), Asteraceae (5 species), Euphorbiaceae (5 species) and Labiatae (4 species). Flowering plant species encountered in 10 to 12 of the intercanopy bare patches sampled were Chaetopappa bellidifolia, Evax prolifera, Croton monanthogynus, Spermolepis inter- mis, Centaurium texense, Galium virgatum, Sida abutifolia, and Schizachyrium scoparium. Nostoc commune, a common soil blue-green algae, was found in every intercanopy bare patch. Non-vascular plant cover was highest near the edge of the central bedrock and decreased with distance. Total plant and litter cover increased with distance from the central bedrock. Total annual cover was five times higher than total perennial cover and both increased with distance from the central bedrock. Mean number of species found per intercanopy patch was 31 ? 4 with a mean Simpson's diversity of 9.2 ?- 2.8. Soil depth increased with distance from the central bedrock, whereas rock cover decreased. Total plant, annual and perennial cover, and litter cover were positively related to soil depth; rock cover was negatively related to soil depth. Nostoc commune relative cover decreased with distance, but Chaetopappa bellidifolia relative cover increased with distance from the patch center, and then decreased near the outer edge of the patch. Heliotropium tenellum and Bothrichloa ischaemum relative cover increased with distance from the center of the patch, with no indication of a decrease near the outer edge. Nostoc commune relative cover was inversely related to soil depth. Relative cover of C. bellidifolia, H. tenellum and B. ischaemum was positively related to soil depth. Almost all soil chemical characteristics were the same from near the central bedrock area to the outer edge of the patch. Soils were slightly basic, high in calcium, low in nitrogen and organic matter. Shallow soil depth and intermittent water availability are probably the main factors that determine composition, including low cover and the high proportion of annuals, in these com- munities.
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