Abstract

The creation and maintenance of complex forest structures has become an important forestry objective. Complex forest structures, often expressed in multimodal shapes of tree size/diameter (DBH) distributions, are challenging to model. Mixture probability density functions of two- or three-component gamma, log-normal, and Weibull mixture models offer a solution and can additionally provide insights into forest dynamics. Model parameters can be efficiently estimated with the maximum likelihood (ML) approach using iterative methods such as the Newton-Raphson (NR) algorithm. However, the NR algorithm is sensitive to the choice of initial values and does not always converge. As an alternative, we explored the use of the iterative expectation-maximization (EM) algorithm for estimating parameters of the aforementioned mixture models because it always converges to ML estimators. Since forestry data frequently occur both in grouped (classified) and ungrouped (raw) forms, the EM algorithm was applied to explore the goodness-of-fit of the gamma, log-normal, and Weibull mixture distributions in three sample plots that exhibited irregular, multimodal, highly skewed, and heavy-tailed DBH distributions where some size classes were empty. The EM-based goodness-of-fit was further compared against a nonparametric kernel-based density estimation (NK) model and the recently popularized gamma-shaped mixture (GSM) models using the ungrouped data. In this example application, the EM algorithm provided well-fitting two- or three-component mixture models for all three model families. The number of components of the best-fitting models differed among the three sample plots (but not among model families) and the mixture models of the log-normal and gamma families provided a better fit than the Weibull distribution for grouped and ungrouped data. For ungrouped data, both log-normal and gamma mixture distributions outperformed the GSM model and, with the exception of the multimodal diameter distribution, also the NK model. The EM algorithm appears to be a promising tool for modeling complex forest structures.

Highlights

  • In forestry, the contemporary paradigms of ecological forestry and close-to-nature silviculture look toward natural disturbance regimes to inform management approaches [1,2]

  • The model family that produced the closest fit with lowest Akaike Information Criterion (AIC) and largest log likelihood (LL) values differed among the three samples, with the log-normal and gamma families typically producing better fits than the Weibull family

  • The twocomponent mixture model was the superior model for grouped-5 and ungrouped, whereas the three-component model was superior for grouped-2.5, grouped-5, and ungrouped

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Summary

Introduction

In forestry, the contemporary paradigms of ecological forestry and close-to-nature silviculture look toward natural disturbance regimes to inform management approaches [1,2]. Due to tremendous variabilities in size, intensity, severity, and frequency of disturbances [4,8], large live legacy trees often survive natural disturbances and are placed in the immediate proximity of the new regeneration [7,9,10,11]. The outcomes of such partial disturbances are highly heterogeneous vertical forest structures in time and space that range from single to two or more cohorts or single to two or multiple canopy layers formed by pure or mixed tree species [12]. Relatively even-aged stands are often characterized by unimodal and near normal distributions, uneven-aged stands typically exhibit rotated sigmoid or reverse-J diameter distributions and multi-aged stands such as many old-growth forests show multimodal and/or irregularly descending distributions characterized by asymmetry, skewness, interruptions (i.e., gaps in the distribution), multimodality, and heavy tails [14]

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