Abstract

This is the first integrated ichnologic-sedimentologic study of the Khussak Formation at the Salt Range, Pakistan. This integrated approach led to identifying thirteen lithofacies and six recurring facies associations. Facies association 1 (FA-1) is made up of massive to parallel laminated mudstone with rare interbeds of sandstone. FA-1 is moderately bioturbated with trace fossils belonging to the Cruziana ichnofacies (Cruziana, Planolites, Rusophycus). FA-1 is interpreted to record deposition in the upper intertidal mud flat setting. Facies association 2 (FA-2) consists of thinly interbedded sandstone (wavy and current rippled) and mudstone (massive to parallel-laminated). The heterolithic wavy bedding is the dominant bedding type. FA-2 is variably bioturbated (BI = 0–5). Trace fossils of moderate diversity of the Cruziana ichnofacies are present. FA-2 represents deposition in the middle intertidal mixed flat setting. Facies association 3 (FA-3) comprises gradational-based, laterally extensive, crudely bedded to massive, medium to lower fine-grained sandstone with thin (cm-scale) mudstone beds. FA-3 is intensely bioturbated (BI = 3–5). Robust and diverse trace fossils belonging to the Skolithos ichnofacies are present (Arenicolites, Cylindrichnus, Diplocraterion, Phycodes, Planolites, Siphonichnus, Skolithos, and Treptichnus). FA-3 records deposition in the lower intertidal sand flat setting. Facies association 4 (FA-4) consists of muddy to silty lower-upper fine-grained sandstone with rare mudstone drapes. FA-4 displays inclined heterolithic stratification (IHS). Tidal couplets (mud and mica drapes) and mud clasts are common. Bioturbation is absent to sparse (BI = 0–2) with low-diversity trace fossils of Skolithos ichnofacies (Cylindrichnus, Diplocraterion, Palaeophycus, Planolites, Skolithos, and rare fugichnia). FA-4 is interpreted to represent deposition in an intertidal creek point bar. Facies association 5 (FA-5) consists of well sorted sandstone (upper medium to fine-grained). The sandstone is erosionally based, and laterally extensive. Planar parallel lamination, trough cross-stratification, reactivation surfaces, and mud drapes are present only locally. The degree of biogenic reworking is low to moderate (BI = 0–4), and traces are sporadically distributed. Trace fossils present are of moderate diversity and abundance, including Arenicolites, Cylindrichnus, Diplocraterion, Monocraterion, Palaeophycus, Planolites, Siphonichnus, and Skolithos. FA-5 is interpreted as subtidal bar deposits. Facies association 6 (FA-6) is characterized by bioturbated muddy sandstone, heterolithic sandstone, and massive mudstone. Physical bedforms are commonly obliterated by bioturbation. Planar horizontal laminae, wavy laminae, small ripples, and synaeresis cracks may occur locally. FA-6 is moderate to intensely bioturbated (BI = 3–6). The distribution of traces (within beds and across the FA) is heterogeneous (sporadically) to homogeneous (locally). Robust Bergaueria, Cylindrichnus, Planolites, Rhizocorallium, Siphonichnus, Teichichnus and Zoophycos are present. FA-6 records deposition in the subtidal bay region. The tidal flat deposits of the Khussak Formation show a distinctive ichnofacies trend. The Cruziana ichnofacies characterizes the lower energy proximal deposits (intertidal mixed and mud flat). In contrast, the high-energy distal deposits (lower intertidal sand flat and shallow subtidal) are characterized by Skolithos ichnofacies. The ichnologic variation between the mud flat, mixed flat, sand flat, and subtidal deposits are gradational, but they can be used as facies boundary indicators. The bioturbation intensity and diversity of trace fossils within the lower Cambrian (Series 2, Stage 4) tidal flat deposits of the Khussak Formation suggest elevated levels of bioturbation intensity during stable environmental conditions in the Cambrian. Furthermore, it implies that the Agronomic Revolution was already emplaced in lower Cambrian (Series 2, Stage 4) tidal flat settings.

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