Abstract
The palaeoenvironmental significance of trace fossil assemblages in the shallow marine deposits of the Late Triassic Nayband Formation of the Tabas Block, Central Iran has been assessed for the first time. The siliciclastic successions of the Nayband Formation, are highly bioturbated and allow relationships between changes in ichnoassemblages within a depositional system to be documented and placed in a palaeoenvironmental framework. The main purposes of this paper are: (1) to describe and illustrate the trace fossils assemblages of the Nayband Formation; (2) integration of ichnologic and sedimentologic data to evaluate different stress factors on the infaunal communities; and (3) to briefly discuss paleoenvironmental distribution of various trace fossil assemblages in order to differentiate between deltaic and non-deltaic shoreface successions. The Upper Triassic Nayband Formation has been studied and measured in one section, where the Nayband Formation is well accessible and continuously exposed. This section was logged and evaluated using physical sedimentology and ichnology in order to establish a basis for the interpretation of sedimentary processes (facies) and depositional systems. In this study, trace fossils (diversity, abundances, and ethologies) and the spatial arrangement of sedimentary structures are used to further refining the interpretation of environmental parameters such as hydrodynamic energy, water turbidity, substrate properties, food supply, temperature, oxygenation, salinity, and sedimentation rates. Based on facies characteristics and stratal geometries, the siliciclastic succession is divided into two facies associations, fluvial-dominated delta (FA), and open marine (FB). Twenty-eight ichnogenera have been identified in delta and open marine successions: The assemblage is dominated by Rhizocorallium, Thalassinoides, Ophiomorpha, Helminthopsis, Palaeophycus and Skolithos. Their distribution is clearly linked with lithofacies and depositional environments. High rates of fluvial discharge, high water turbidity, seasonally high rates of deposition and phytodetrital pulses in river-dominated deltas may cause marked variations in the temperature, oxygenation and salinity of the fluvial-dominated delta successions. All of these factors in combination lead to impoverishment in trace diversity and reduction in burrow size but to rapid colonization of the substrate by single-layer colonizers of opportunistic deposit feeders. Low ichnodiversity, low bioturbation intensities with highly bioturbated clusters shown by single colonization events by opportunistic, simple feeding strategies of trophic generalists and a paucity of suspension-feeding structures elements represent a “stressed”, non-archetypal Cruziana ichnofacies. The changes of behavioural complexity and the tiering profiles in the offshore-shoreface complex of the Nayband Formation when compared with the river-dominated delta successions represent the transition from epifaunal life habits with simple tiering structures to infaunal life habits with complex tiering pattern. The occurrence of diverse trace fossil suites attributable to the Zoophycos ichnofacies, the archetypal Cruziana ichnofacies and the Skolithos ichnofacies in wave-dominated shoreface-offshore complex point to stress-free environmental conditions in open waters due to persistent wave agitation, and hence, a wide colonization window.
Published Version
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