Abstract
Intracellular microcolonies (IMCs) of bacteria that reflect Chlamydia-like organisms (CLOs) or Rickettsia-like organisms (RLOs) cause disease and mortalities in mollusks globally including the Pacific oyster (previously Crassostrea gigas but now Magallana gigas), the clam (Meretrix lusoria), and the abalone (Haliotis rufescens). However, we will focus here on the disease impacts for scallops species such as Patinopecten yessoensis and Pecten maximus. RLOs are obligate parasites, being polymorphic shape, 0.4–2μm, and behave like bacteria during their binary fission or budding multiplication phase. These IMCs include members of the Alphaproteobacteria and Gammaproteobacteria classes of the Proteobacteria phylum, which means that they are not all RLOs (order Rickettsiales).
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