Nel, A., Roques, P., Prokop, J., & Garrouste, R., 11 September 2018. A new, extraordinary ‘damselfly-like’ Odonatoptera from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: ‘Exopterygota’). Alcheringa 43, 241–245. ISSN 0311-5518.Enigmaptera magnifica gen. et sp. nov., type genus and species of the new odonatopteran family Enigmapteridae, is described from the Moscovian of Avion (northern France). It is the sister group of the major clade Neodonatoptera, placed together in the new clade Paneodonatoptera. Its wing venation has characters never found in other Odonatoptera. It is a further case of convergent wing petiolation in this superorder. Enigmaptera magnifica, like the protozygopteran Jacquesoudardia magnifica from the same outcrop, probably lived like the extant damselflies along the shores of lakes and rivers, hunting the small insects found in the same deposits. These discoveries show that very small insects were significant elements of the entomofaunal diversity and trophic chains of the Late Carboniferous ecosystems.Romain Garrouste [garroust@mnhn.fr] Institut Systématique Evolution Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, 57 rue Cuvier, CP 50, 75005 Paris, France; Patrick Roques [patrick.roques93@wanadoo.fr] Allée des Myosotis, Neuilly sur Marne, F-93330, France.; Jakub Prokop [jprokop@natur.cuni.cz] Charles University, Faculty of Science, Department of Zoology, Viničná 7, CZ-128 44, Praha 2, Czech Republic.