Wing Loading Research Articles

Arrangements of intramembranous muscles of wings are influenced by body mass across the radiation of phyllostomid bats.

Extensive research into bat flight mechanisms has highlighted the complex functional and evolutionary dynamics of their wing structures, yet the anatomical details of certain wing muscles remain elusive. In particular, the intramembranous plagiopatagiales proprii muscles, located within the plagiopatagium-an area of the wing lacking direct joint connections-exhibit remarkable variation across bat families. These muscles, which extend anteroposteriorly in macroscopic bundles, play a crucial role in wing stiffening, modulating membrane tension, and reducing wing curvature during flight. Since larger bats tend to have higher wing loading (WL; the ratio of body mass [BMa] to wing area) and may therefore experience increased patagial curvature and resultant drag, we hypothesized that body size significantly influences the evolutionary development of the plagiopatagiales proprii muscles. This study investigates the relationship between BMa and the morphology of the plagiopatagiales proprii in New World leaf-nosed bats (Phyllostomidae), employing bivariate allometry, multivariate analysis, and comparative phylogenetic methods across 24 species from eight phyllostomid subfamilies. Our findings reveal a significant phylogenetic signal in muscle architecture, along with positive evolutionary allometry in muscle area. This suggests an adaptive increase in muscle size in larger species, likely to counterbalance the increased WL, reduce wing curvature, and minimize drag. This research enhances our understanding of the functional and adaptive morphological evolution of intramembranous wing muscles in phyllostomid bats, underscoring their evolutionary significance.

Paving the Way for Sustainable UAVs Using Distributed Propulsion and Solar-Powered Systems

Hybrid systems offer optimal solutions for unmanned aerial platforms, showcasing their technological development in parallel and series configurations and providing alternatives for future aircraft concepts. However, the limited energetic benefit of these configurations is primarily due to their weight, constituting one of the main constraints. Solar PV technology can provide an interesting enhancement to the autonomy of these systems. However, to create efficient propulsion architectures tailored for specific missions, a flexible framework is required. This work presents a methodology to assess hybrid solar-powered UAVs in distributed propulsion configurations through a two-level modeling scheme. The first stage consists of determining operational and design constraints through parametric models that estimate the baseline energetic requirements of flight. The second phase executes a nonlinear optimization algorithm tuned to find optimal propulsion configurations in terms of the degree of hybridization, number of propellers, different wing loadings, and the setup of electric distributed propulsion (eDP) considering fuel consumption as a key metric. The results of the study indicate that solar-hybrid configurations can theoretically achieve fuel savings of up to 80% compared to conventional configurations. This leads to a significant reduction in emissions during long-endurance flights where current battery technology is not yet capable of providing sustained flight.

Gust Response and Alleviation of Avian-Inspired In-Plane Folding Wings.

The in-plane folding wing is one of the important research directions in the field of morphing or bionic aircraft, showing the unique application value of enhancing aircraft maneuverability and gust resistance. This article provides a structural realization of an in-plane folding wing and an aeroelasticity modeling method for the folding process of the wing. By approximating the change in structural properties in each time step, a method for calculating the structural transient response expressed in recursive form is obtained. On this basis, an aeroelasticity model of the wing is developed by coupling with the aerodynamic model using the unsteady panel/viscous vortex particle hybrid method. A wind-tunnel test is implemented to demonstrate the controllable morphing capability of the wing under aerodynamic loads and to validate the reliability of the wing loads predicted by the method in this paper. The results of the gust simulation show that the gust scale has a significant effect on the response of both the open- and closed-loop systems. When the gust alleviation controller is enabled, the peak bending moment at the wing root can be reduced by 5.5%∼47.3% according to different gust scales.

Oxidative state is associated with migration distance, but not traits linked to flight energetics

Flight can be highly energy demanding, but its efficiency depends largely on flight style, wing shape and wing loading, and a range of morphological and lifestyle adaptations that can modify the cost of sustained flight. Such behavioural and morphological adaptations can also influence the physiological costs associated with migration. For instance, during intense flight and catabolism of reserves, lipid damage induced by pro‐oxidants increases, and to keep oxidative physiological homeostasis under control, the antioxidant machinery can be upregulated. Studies on the oxidative physiology of endurance flight have produced contradictory results, making generalization difficult, especially because multispecies studies are missing. Therefore, to explore the oxidative cost of flight and migration, we used samples collected during the breeding season from 113 European bird species and explored the associations of measures of antioxidant capacity (total antioxidant status, uric acid and glutathione concentration) and oxidative damage of lipids (malondialdehyde) with variables reflecting flight energetics (year‐round or specifically during migration) using a phylogenetic framework. We found that none of the traits predicting year‐round flight energy expenditure (flight style, wing morphology and flight muscle morphology) explained any measures of oxidative state. Our results suggest that birds endure their everyday flight exercise without or with low oxidative cost. However, oxidative damage to lipids and one component of the endogenous antioxidant system (uric acid), measured after the end of spring migration on breeding adult birds, increased with migration distance. Our results suggest that migration could have oxidative consequences that might be carried over to subsequent life‐history stages (breeding).

Flight activity and effort of breeding pied flycatchers in the wild, revealed with accelerometers and machine learning.

Flight behaviours of birds have been extensively studied from different angles such as their kinematics, aerodynamics and, more generally, their migration patterns. Nevertheless, much is still unknown about the daily foraging flight activity and behaviour of breeding birds, and potential differences among males and females. The recent development of miniaturized accelerometers allows us a glimpse into the daily life of a songbird. Here, we tagged 13 male and 13 female pied flycatchers (Ficedula hypoleuca) with accelerometers and used machine learning approaches to analyse their flight activity and effort during the chick rearing period. We found that during 2h of foraging, chick-rearing pied flycatchers were flying on average 13.7% of the time. Almost all flights (>99%) were short flights lasting less than 10s. Flight activity changed throughout the day and was highest in the morning and lowest in the early afternoon. Male pied flycatchers had lower wing loading than females, and in-flight accelerations were inversely correlated with wing loading. Despite this, we found no significant differences in flight duration and intensity between sexes. This suggests that males possess a higher potential flight performance, which they did not fully utilize during foraging flights.

Reduced palatability, fast flight, and tails: decoding the defence arsenal of Eudaminae skipper butterflies in a Neotropical locality.

Prey often rely on multiple defences against predators, such as flight speed, attack deflection from vital body parts, or unpleasant taste, but our understanding on how often and why they are co-exhibited remains limited. Eudaminae skipper butterflies use fast flight and mechanical defences (hindwing tails), but whether they use other defences like unpalatability (consumption deterrence) and how these defences interact have not been assessed. We tested the palatability of 12 abundant Eudaminae species in Peru, using training and feeding experiments with domestic chicks. Further, we approximated the difficulty of capture based on flight speed and quantified it by wing loading. We performed phylogenetic regressions to find any association between multiple defences, body size, and habitat preference. We found a broad range of palatability in Eudaminae, within and among species. Contrary to current understanding, palatability was negatively correlated with wing loading, suggesting that faster butterflies tend to have lower palatability. The relative length of hindwing tails did not explain the level of butterfly palatability, showing that attack deflection and consumption deterrence are not mutually exclusive. Habitat preference (open or forested environments) did not explain the level of palatability either, although butterflies with high wing loading tended to occupy semi-closed or closed habitats. Finally, the level of unpalatability in Eudaminae is size dependent. Larger butterflies are less palatable, perhaps because of higher detectability/preference by predators. Altogether, our findings shed light on the contexts favouring the prevalence of single versus multiple defensive strategies in prey.

Island area and diet predict diversity and distribution of bats in a Pacific Northwest archipelago

Abstract The island biogeography theory predicts that species richness in islands and island-like systems is the ultimate result of island isolation and area. Species with high dispersal capabilities are predicted to be less affected by these factors because of their capacity to move more efficiently between islands or habitats, and here we test this idea in bats, the only mammals capable of flight. We conducted mist net and acoustic surveys across 21 islands in the San Juan Archipelago (Washington State, United States) and adjacent northwest mainland to: (i) investigate the effects of island area, distance from mainland, and habitat on bat diversity; and (ii) evaluate whether differences in morphological (body mass, forearm length, wing loading) and ecological (dietary niche breadth, foraging guild) traits among species influence their prevalence across islands. We found that island size strongly influenced patterns of species richness, with larger islands having a greater number of bat species. However, neither island distance from mainland nor any measure of habitat availability was a significant predictor of species richness at the scale of this study. Additionally, we found that dietary niche breadth, as opposed to any morphological trait, best predicted the prevalence of species across the islands. Our results suggest that species with more specialized diets may be more vulnerable to habitat fragmentation, and provide insight into how geographic and ecological factors affect the diversity of insular bat communities, adding to growing knowledge about the role of species traits as mediators of their responses to large-scale landscape structure.

Novel Mito-Nuclear Combinations Facilitate the Global Invasion of a Major Agricultural Crop Pest.

A fundamental understanding of the underlying mechanisms involved in biological invasions is crucial to developing effective risk assessment and control measures against invasive species. The fall armyworm (FAW), Spodoptera frugiperda, is a highly invasive pest that has rapidly spread from its native Americas into much of the Eastern Hemisphere, with a highly homogeneous nuclear genetic background. However, the exact mechanism behind its rapid introduction and propagation remains unclear. Here, a systematic investigation is conducted into the population dynamics of FAW in China from 2019 to 2021 and found that FAW individuals carrying "rice" mitochondria (FAW-mR) are more prevalent (>98%) than that with "corn" mitochondria (FAW-mC) at the initial stage of the invasion and in newly-occupied non-overwintering areas. Further fitness experiments show that the two hybrid-strains of FAW exhibit different adaptions in the new environment in China, and this may have been facilitated by amino acid changes in mitochondrial-encoded proteins. FAW-mR used increases energy metabolism, faster wing-beat frequencies, and lower wing loadings to drive greater flight performance and subsequent rapid colonization of new habitats. In contrast, FAW-mC individuals adapt with more relaxed mitochondria and shuttle energetics into maternal investment, observed as faster development rate and higher fecundity. The presence of two different mitochondria types within FAW has the potential to significantly expand the range of damage and enhance competitive advantage. Overall, the study describes a novel invasion mechanism displayed by the FAW population that facilitates its expansion and establishment in new environments.

A straightforward protocol to sample morphological traits of dragonflies and damselflies in the field.

Scarcity of morphological data limits the potential of functional ecology approaches, which rely on traits to elucidate ecological processes. Dragonflies and damselflies (Odonata) are a frequently used ecological model for which, however, only limited morphological data is available. Here, it is presented a field sampling protocol to collect ecologically relevant yet largely unavailable morphological traits of Odonata. The protocol enables the straightforward collection of traits from living individuals directly in the field. Those traits include body mass, wing area and wing loading as well as thorax width, hindwing length and body length. Furthermore, the protocol allows for posterior wing morphometric analyses. The protocol proved to be robust and universally applicable based on testing on roughly half (76) of all European odonate species. The use of this protocol can increase our understanding of odonatan morphology at interspecific and intraspecific levels and assist in developing mechanistic understanding of their ecology.

On the wings of dragons: Wing morphometric differences in the sexually dichromatic common whitetail skimmer dragonfly, Plathemis lydia (Odonata: Libellulidae).

Sexual dimorphism is common throughout the animal kingdom, leading to sex-specific phenotypic differences. The common whitetail skimmer dragonfly, Plathemis lydia (Drury, 1773), is sexually dichromatic, where males of this species display a conspicuous white abdomen and females display a dark brown abdomen. Differences in abdomen conspicuousness between male and female P. lydia are likely attributed to differences in selective pressure where males use their white conspicuous abdomen during male-male territorial chases. We hypothesized that male P. lydia would exhibit wing morphology adaptations to better offset the costs of predation and territoriality and that these adaptations would differ from females. We used field-collected images to quantify differences in body length, wing length, wing area, wing shape, and wing loading between male and female P. lydia. Our results show that male P. lydia have significantly shorter fore and hind wings relative to body size with a higher wing loading when compared to females. We also found that male P. lydia have narrower and pointier fore and hind wings compared to females. These results are consistent with the idea that males are adapted for faster flight, specifically higher acceleration capacity, and higher agility whereas females are adapted for higher maneuverability.

Review and meta-analysis of correlates of home range size in bats

Abstract Home range is an ecological concept that affects many aspects of the life of vertebrates and hence understanding how it varies between species is crucial. Mammalian home range size has been linked to body size and diet, but these studies were based predominantly on terrestrial species and most specifically excluded bats. As the only group of flying mammals, bats experience distinctly different constraints on movement, and hence home range. However, despite their diversity, relatively little is known about the home ranges of bats, and this is the first global review assessing correlates of their home range size. Our hypothesis is that home range will be impacted by different ecological conditions experienced by the bats and by the biological traits of the bats themselves. We performed a meta-analysis based on published data for 81 bat species to identify variables that contribute most to home range size. Sex, wing loading, functional group, colony size, dietary class, distance from the equator (latitudinal region), habitat type, and the interaction between habitat type and latitudinal region were all important explanatory variables. Wing loading was positively correlated with home range size—while females, open-air foragers, large colony sizes, and bats in temperate regions consistently had large home ranges. Understanding the correlates of home range has important implications, for example, for bat conservation and for assessing the risk of spillover of zoonotic pathogens from bats to humans and livestock.

Simplified Coupled Calculation Method for Aeroelasticity of Large Flexible Aircraft

Large flexible aircraft have the characteristics of large wingspan, low wing load, slow flight speed, and significant impact from gust loads. Therefore, the load and response issues of such aircraft have received widespread attention from scholars at home and abroad. This article establishes a simplified coupled calculation method for gust loads on large flexible aircraft, achieving coupled calculations of flight motion, structural deformation, and aerodynamic changes at different time steps. Due to the simple strip method used for aerodynamic modeling, this method is relatively fast compared to the CFD-CSD calculation method. In order to verify the accuracy of the method, the calculation results were compared with those of the standard calculation method. The comparative calculation results show that the simplified coupling calculation method is relatively accurate for this calculation example and can play an important role in engineering experiments with low requirements for calculation results.

Fluid–Structure Coupled Analysis of Maneuver Load Alleviation on a Large Transport Aircraft

Fluid–structure coupled simulations are performed for quasi-steady pitching maneuvers of a large transport aircraft with a load alleviation system consisting of trailing edge flaps and droop noses at the leading edge. For the fluid part of the simulation, a RANS approach is used, whereas the structural simulation is based on a linear modal model. The selected maneuvers are located on the lift boundary of the maneuver envelope at maximum load factor, as not only are the structural loads high here, but also redistributing lift for load alleviation is especially challenging as the wing is close to stall. For a target value of 33% reduction in wing bending loads, which is derived from the CS-25, the effects of applying load alleviation on wing loads, deformations, and aircraft maneuverability are investigated. It is found that the desired reduction of wing bending loads can be achieved for all maneuvers considered, while controlling the torsional moment turns out to be more difficult. A loss of maneuverability due to load alleviation is observed for only one of the maneuvers, while for the other maneuvers, maneuverability could even be increased. To assess the influence of the elasticity of the wing, additional simulations of the rigid wing are carried out and compared with the elastic results. Finally, other potential applications of the considered flap system are also explored. Here, for example, a cruise drag reduction of up to 1.8% is possible by adjusting the spanwise lift distribution.

Research on an optimization design method for a TBCC propulsion scheme

Abstract An optimization methodology for a TBCC propulsion schemes was established for hypersonic vehicles, focusing on the integration of aircraft and engine performance. Altitude-velocity characteristics of TBCC propulsion were obtained through engine performance calculations. By analyzing mission requirements, lift-drag characteristics, and flight constraints, the take-off thrust-weight ratio and wing load of the vehicle were optimized to meet the flight conditions. In addition, the fuel ratio was calculated. To determine the vehicle’s gross take-off weight and the engine’s take-off thrust, a model considering the weight of the vehicle and the turbine engine were used. The optimization process selects four thermodynamic cycle parameters for the turbine engine as the independent variables. An improved particle swarm optimization-back propagation neural network was used to establish the relationship between the four parameters and the gross take-off weight, aiming to minimize the vehicle’s weight. The results of the optimization process show that the total take-off weight of the optimized vehicle has decreased from 112363.41 kg to 102218.98 kg. The required uninstalled take-off thrust of TBCC has also been reduced from 162.86 kN to 147.49 kN, resulting in a decrease in mass flow from 156.50 kg/s to 133.30 kg/s.

Flight dynamics of aircraft incorporating the semi-aeroelastic hinge

Aircraft like the Boeing 777-X use on-ground folding wingtips to meet the airport gate size restriction while increasing the aspect ratio during flight to reduce induced drag. A recent concept of aircraft design is to utilise in-flight floating wingtips as a means of load alleviation, which is known as semi-aeroelastic hinge (SAH). This device allows wingtips to be released during manoeuvre and severe gusts to alleviate wing loads, while locking the wingtips during cruise to maintain an optimum aerodynamic shape. This paper develops a flight mechanics model incorporating flexible wings to study the influence of the SAH device on multiple aspects of aircraft flight dynamics. The dynamic responses of the aircraft to gusts and control surface inputs are computed and compared for various hinge and wingtip configurations and release times. It was found that the gust load measured from the wing with free hinge configuration was approximately 40% lower compared to that of the fixed hinge case. It is also shown that when the SAH is released, it can significantly reduce an aircraft's roll damping and the frequency of the short-period mode, leading to higher roll and pitch rates. Furthermore, it shows that the transient responses following the wingtip release will exacerbate the responses induced by gusts, and greater load alleviation can be achieved by manipulating the hinge release time for each gust length. Finally, a step input of the elevator during the hinge release was found to be beneficial for enhancing the gust load alleviation.

Effects of wing loading on the aerodynamic characteristics of autorotating maple seeds

Natural maple seeds can fall into stable autorotation when descending. Their excellent aerodynamic performance can be applied to biomimetic aircraft design. Wing loading plays an important role in flight performance. To make clear the effects of wing loading on the aerodynamic characteristics of autorotating maple seeds, experiments integrated with numerical simulation were performed. In the experiment, the free fall test and the wind tunnel test were conducted on maple seeds with variable wing loadings. During the free fall process, four typical stages can be divided for the maple seed according to different kinematic characteristics. In the numerical simulation, the Moving Reference Frame method was used to simulate the fluid of autorotating maple seeds. Both experiment and simulation results show that descending velocities and spinning rates rise almost linearly with the increase in wing loadings, and descending velocity is closely related to spinning rate. Obvious leading-edge vortexes were observed on seed wing, which are the flow mechanism of high lift. The pressure differences between the top and lower surfaces of seeds grow with increasing wing loadings. Larger wing loadings lead to more serious separation of leading-edge vortex. The parameter influence analysis demonstrates that smaller conning angles or larger spinning rates are beneficial for high-lift flight. The spinning rate has a more significant effect on the autorotating maple seeds. However, when the spinning rate is larger than 200 rad/s, the lift does not increase anymore. The spinning rate should be controlled smaller than 200 rad/s when applied to biomimetic aircraft.

Heat stress reduced survival but sped up development in Heliconius erato butterflies

AbstractAnthropogenic climate change is thought to present a significant threat to biodiversity, in particular to tropical ectotherms, and the effects of long‐term developmental heat stress on this group have received relatively little research attention. Here, we studied the effects of experimentally raising developmental temperatures on a tropical butterfly. We measured survival, development time, adult body mass and wing size of Heliconius erato demophoon (Linnaeus) (Lepidoptera: Nymphalidae) across three temperature treatments. Egg survival was lower in the hotter treatments, with 84%, 73% and 49% of eggs hatching in the 20–30°C (fluctuating temperature with 12 h at 20°C followed by 12 h at 30°C), 23–33°C and 26–36°C treatments, respectively. Larval survival was three times lower in the 26–36°C treatment (8%) compared with the 20–30°C treatment (26%), and we did not detect differences in pupal survival across treatments due to high mortality in earlier stages. Under a moderately increased temperature at 23–33°C, larvae developed faster and adults had a higher body mass and wing loading, but this was not seen in the hottest treatment (26–36°C). Females were also heavier than males in the 23–33°C treatment, but there was no associated increase in wing size. This may suggest a different developmental response to moderately elevated temperatures between the sexes. In summary, high developmental temperatures are particularly lethal for eggs and less so for larvae and also affect adult morphology. This highlights the importance of understanding the effects of temperature variation across ontogeny in tropical ectotherms.

Flight Dispersal in Supratidal Rockpool Beetles.

Flight dispersal is ecologically relevant for the survival of supratidal rockpool insects. Dispersal has important consequences for colonisation, gene flow, and evolutionary divergence. Here, we compared the flight dispersal capacity of two congeneric beetle species (Ochthebius quadricollis and Ochthebius lejolisii) that exclusively inhabit these temporary, fragmented, and extreme habitats. We estimated flight capacity and inferred dispersal in both species using different approaches: experimental flying assays, examination of wing morphology, and comparison of microsatellite markers between species. Our findings revealed that both species exhibited similar flight behaviour, with 60 to 80% of the individuals flying under water heating conditions. Notably, females of both species had larger body sizes and wing areas, along with lower wing loading, than males in O. quadricollis. These morphological traits are related to higher dispersal capacity and more energetically efficient flight, which could indicate a female-biassed dispersal pattern. The wing shapes of both species are characterised by relatively larger and narrower wings in relation to other species of the genus, suggesting high flight capacity at short distances. Molecular data revealed in both species low genetic divergences between neighbouring populations, non-significant differences between species, and no isolation by distance effect at the study scale (<100 km). These results point to passive dispersal assisted by wind.

Single-component Multi-rate Real Time Hybrid Simulation Pilot Test on a Composite Structure

This paper represents a single component multi-rate Real-Time Hybrid Simulation (mrRTHS) strategy for structural assessment of a cantilever Glass Fiber Reinforced Polymer (GFRP) beam loaded at the tip by a sinusoidal point load. This emulated structure is implemented as a simplified wind turbine blade in terms of geometry, scale and load – here with special attention paid to the root and max-chord section. Thus, the experimental substructure comprises the clamped end of the GFRP beam while the free end makes up the numerical substructure. The partitioning between the numerical and experimental substructure – referred to here as the shared boundary – Includes a discrete point with 3 degrees-of-freedom (dof). The numerical substructure generates a displacement signal through a Taylor basis with a coarse time step to optimize computational resources. Using the previous displacement data points, a finer control signal is generated to ensure accurate actuator control in the transfer system. A DIC and inertia compensator is implemented to account for the compliance and dynamics imposed by the load train in the transfer system. The structural response is investigated by mrRTHS for an execution frequency in the range: 0.074Hz – 2.96Hz for the sinusoidal point load. The system performance is evaluated against an experimental test setup of the emulated structure – referred to here as the experimental reference. With a root-mean-square (RMS) error in the order of 8-20% between the mrRTHS and reference, the system proved successful in terms of stability and overall correlation at the shared boundary, which is considered an important milestone towards single component mrRTHS on a structure like e.g. a wind turbine blade, aircraft wing or similar cantilever-shaped large load carrying structure.

Modeling advanced air mobility aircraft in data-driven reduced order realistic urban winds

The concept of Advanced Air Mobility involves utilizing cutting-edge transportation platforms to transport passengers and cargo efficiently over short distances in urban and suburban areas. However, using simplified atmospheric models for aircraft simulations can prove insufficient for modeling large disturbances impacting low-altitude flight regimes. Due to the complexities of operating in urban environments, realistic wind modeling is necessary to ensure trajectory planning and control design can maintain high levels of safety. In this study, we simulate the dynamic response of a representative advanced air mobility platform operating in wing-borne flight through an urban wind field generated using Large Eddy Simulations (LES) and a wind field created using reduced-order models based on full-order computational solutions. Our findings show that the longitudinal response of the aircraft was not greatly affected by the fidelity of the LES models or if the spatial variation was considered while evaluating the full-order wind model. This is encouraging as it indicates that the full LES generation of the wind field may not be necessary, which decreases the complexity and time needed in this analysis. Differences are present when comparing the lateral response, owing to the differences in the asymmetric loading of the planform in the full and reduced order models. These differences seen in the lateral responses are expected to increase for planforms with smaller wing loadings, which could pose challenges. Additionally, the response of the aircraft to the mean wind field, the temporal average of the full order model, was misrepresentative in the longitudinal response and greatly under-predicted control surface activity, particularly in the lateral response.