It is generally accepted that the total power loss in gears can be decomposed into contributions of friction between the teeth, lubrication (churning and/or jet lubrication that could induce gas—oil trapping in inter-tooth spaces), and gear windage effects. For low- to medium-speed transmissions, tooth friction is recognized as the main source of dissipation for gears and can significantly influence the temperature equilibrium of a gearbox. In this article, a model to predict power losses due to tooth friction combined with a thermal network to describe the temperature distribution in a geared transmission are presented. The numerical results compare favourably with experimental measurements from a gear test rig.