Tortula caroliniana n. sp.-Growing in extensive mats on the bark of trees. Stems somewhat matted below with radicles, with brown leaves, above simple, with bright green leaves, which readily become brown with age. Plants very short or sometimes more lengthened, up to about I cm. Stem in section round, with central strand; outer 2-3 rows of cells gradually smaller and thickerwalled than those within. Leaves obovate, normally about 2.5 mm. long, I mm. wide in widest part, borders strongly reflexed in basal M-%, apex abruptly apiculate with costa only percurrent; costa slender, up to 50o in width in lower part of leaf, brown, smooth on back, in section with two guide-cells and two ventral cells nearly equally large adjoining them, dorsally of uniform stereid cells. Leaf-cells at base smooth, rectangular, the outer ones narrow and somewhat chlorophyllose, up to 50o x Io , in region of costa larger, hyaline with brown walls, up to 70 x 25 ;~; cells of upper part of leaf irregularly roundish quadrate, with fairly thick walls and distinct trigones, averaging about 14 t in diameter, in border region slightly smaller and with thicker walls, but not giving the impression of a distinct border, papillose on both surfaces, the papillae mostly crescent shaped,' 3 V or less from end to end of the crescent, normally 4 per cell on either side, rarely more than 6, in smaller cells of border region often 2 or 3. Papillae rather low in profile and quite independent of each other, the wall to which they are attached not at all thickened, the outer wall of cell very slightly bulging. Inflorescence and sporophyte not seen. Reproduction by means of very numerous propagula, more or less cylindrical in shape, with rounded ends, produced from ventral surface of upper half of leaf blade,2 the cells from which they originate generally slightly smaller than those surrounding them, often lacking chlorophyll and with papillae less distinct or lacking. Propagula densely chlorophyllose, somewhat roughened on outer surface, divided by parallel planes at right angles to the axis of development at intervals of 20-25 t, these divisions often further subdivided into 2 or 4 parts by planes at right angles, the subdivision of adjacent segments in this case not necessarily conformable, the end segments usually smaller and undivided. The number of primary divisions varies with the length of the propagulum all the way up to 8, which is sometimes but not often exceeded. The width of the propagulum varies up to about 45 [L, the length is entirely variable, extending up to 200 L or more.