Western Scheldt Estuary Research Articles

Analytical and numerical analysis of tides and salinities in estuaries; part I: tidal wave propagation in convergent estuaries

Analytical solutions of the momentum and energy equations for tidal flow are studied. Analytical solutions are well known for prismatic channels but are less well known for converging channels. As most estuaries have a planform with converging channels, the attention in this paper is fully focused on converging tidal channels. It will be shown that the tidal range along converging channels can be described by relatively simple expressions solving the energy and momentum equations (new approaches). The semi-analytical solution of the energy equation includes quadratic (nonlinear) bottom friction. The analytical solution of the continuity and momentum equations is only possible for linearized bottom friction. The linearized analytical solution is presented for sinusoidal tidal waves with and without reflection in strongly convergent (funnel type) channels. Using these approaches, simple and powerful tools (spreadsheet models) for tidal analysis of amplified and damped tidal wave propagation in converging estuaries have been developed. The analytical solutions are compared with the results of numerical solutions and with measured data of the Western Scheldt Estuary in the Netherlands, the Hooghly Estuary in India and the Delaware Estuary in the USA. The analytical solutions show surprisingly good agreement with measured tidal ranges in these large-scale tidal systems. Convergence is found to be dominant in long and deep-converging channels resulting in an amplified tidal range, whereas bottom friction is generally dominant in shallow converging channels resulting in a damped tidal range. Reflection in closed-end channels is important in the most landward 1/3 length of the total channel length. In strongly convergent channels with a single forward propagating tidal wave, there is a phase lead of the horizontal and vertical tide close to 90o, mimicking a standing wave system (apparent standing wave).

Optimization and development of analytical methods for the determination of new brominated flame retardants and polybrominated diphenyl ethers in sediments and suspended particulate matter

With more stringent legislation on brominated flame retardants, it is expected that increasing amounts of substitutes would replace polybrominated diphenylethers (PBDEs). Therefore, the development and optimization of analytical methodologies that allow their identification and quantification are of paramount relevance. This work describes the optimization of an analytical procedure to determine pentabromochlorocyclohexane, tetrabromo-o-chlorotoluene, 2,3,5,6-tetrabromo-p-xylene, tetrabromophthalic anhydride, 2,3,4,5,6-pentabromotoluene, tris(2,3-dibromopropyl)phosphate, decabromodiphenylethane and 1,2-bis(2,4,6-tribromophenoxy)ethane together with PBDEs in sediments and in suspended particulate matter. This method comprises a pressurized liquid extraction followed by three cleanup steps (gel permeation chromatography and solid phase extraction on Oasis™ HLB and on silica cartridges). Gas chromatography–mass spectrometry, using electron capture negative chemical ionization, is used for the final analysis. The proposed method provides recoveries >85%. The method was applied to sediment and suspended particulate matter samples from different locations in the Western Scheldt estuary (the Netherlands). To the best of our knowledge, this is the first time that the occurrence of the additive flame retardants 2,3,5,6-tetrabromo-p-xylene, 3,4,5,6-tetrabromo-o-chlorotoluene and 2,3,4,5,6-pentabromochlorocyclohexane is reported in the literature. The concentrations of these new flame retardants ranged from 0.05 to 0.30 μg/kg dry weight.Electronic supplementary materialThe online version of this article (doi:10.1007/s00216-011-4807-8) contains supplementary material, which is available to authorized users.

Changes in Phytoplankton Biomass in the Western Scheldt Estuary During the Period 1978–2006

We investigated whether climate change results in long-term changes in phytoplankton biomass and phenology in a turbid eutrophic coastal plain estuary. Changes in annual mean chlorophyll a (chla) concentrations were studied for the period 1978–2006 in the eutrophic and turbid macro-tidal Western Scheldt estuary. Three stations were investigated: WS1, at the mouth of the estuary; station WS6, halfway up the estuary; and station WS11, near the Dutch–Belgian border near the upstream end of the estuary. No significant long-term changes in yearly averaged chla concentrations were observed in WS1 and WS6, but in WS11 the phytoplankton biomass decreased considerably. This is most likely due to an increase in grazing pressure as a result of an improvement in the dissolved oxygen concentrations. Spectral analyses revealed a possible periodicity of 7 years in the mean chla which was related to periodicity in river discharge. We also observed strong phenological responses in the timing of the spring/summer bloom which were related to a well-documented increase in the temperature in the estuary. The fulcrum, the center of gravity or the day at which 50% of the cumulative chla was reached during the year, advanced by 1–2 days/year. A similar trend was observed for the month in which the maximum bloom was observed, with the exception of station WS1. All stations showed an earlier initiation of the bloom, whereas the day at which the phytoplankton bloom was terminated also moved forward in time excepted for WS11. As a result, the bloom length decreased at station WS1, remained the same at station WS6, and increased at WS11. This complicated pattern in bloom phenology demonstrates the complex nature of ecosystem functioning in estuaries.

Presence of Connecting Channels in the Western Scheldt Estuary

To predict the effects of dredging operations in the Western Scheldt estuary, insight into the morphological behaviour of its channels is needed. The estuary features two large main channels that meander alongside each other and that are linked by smaller connecting channels. These connecting channels originate from water level differences between the two main channels. Three hydrodynamic mechanisms are investigated that are responsible for generating such water level differences: (1) differences in tidal wave propagation along two neighbouring main channels, (2) centrifugal forces, and (3) Coriolis forces. The magnitude of the three mechanisms is analytically assessed based on channel geometry, which is derived from historical depth charts. The analysis demonstrates a large temporal variability of the water level differences due to tidal wave propagation, mainly as a function of changes in the depth ratio between the two main channels. Conversely, both the centrifugal effect and the Coriolis effect are relatively constant over time. The temporal evolution of the connecting channels can therefore primarily be attributed to the wave propagation mechanism. A correlation analysis reveals a linear relationship between connecting channel dimensions and the net water level differences produced by the three processes. The relationship suggests that dredging operations may significantly influence the evolution of connecting channels by changing the depth ratio between the two main channels. The proposed future deepening of the navigation channel is therefore expected to induce a decline in the size, or even a total disappearance, of connecting channels in some parts of the estuary.

Modelling impact of dredging and dumping in ebb-flood channel systems

For a channel-shoal system in a funnel-shaped basin the impact of dredging and dumping is investigated using a complex process-based model. First, the residual flow and sediment transport circulations are analysed for the channel-shoal pattern, which has emerged after a long-term model simulation. Results are compared to the Western Scheldt estuary, which forms the inspiration for this study. Subsequently, different dredge and dump scenarios are modelled, according to a conceptual model, in which ebb-and flood-channels and enclosed shoals form morphodynamic units (cells) with their own sediment circulation. Model results show that dumping sediment in a channel further reduces the channel depth and induces erosion in the opposite channel, which enhances tilting of the cross-section of the cell and eventually can lead to the degeneration of a multiple channel system into a single channel. The impact of different dredging and dumping cases agrees with results from a stability analysis. This means that this type of model applied to a realistic geometry can potentially be used for better prediction of the impact of human interventions.

Long‐term morphodynamic evolution of a tidal embayment using a two‐dimensional, process‐based model

The research objective is to investigate long‐term evolution of estuarine morphodynamics with special emphasis on the impact of pattern formation. Use is made of a two‐dimensional (2‐D), numerical, process‐based model. The standard model configuration is a rectangular 80 km long and 2.5 km wide basin. Equilibrium conditions of the longitudinal profile are analyzed using the model in 1‐D mode after 8000 years. Two‐dimensional model results show two distinct timescales. The first timescale is related to pattern formation taking place within the first decades and followed by minor adaptation according to the second timescale of continuous deepening of the longitudinal profile during 1600 years. The resulting longitudinal profiles of the 1‐D and 2‐D runs are similar apart from small deviations near the mouth. The 2‐D results correspond well to empirically derived relationships between the tidal prism and the channel cross section and between the tidal prism and the channel volume. Also, comparison between the current model results and data from the Western Scheldt estuary (in terms of bar length, hypsometry, percentage of intertidal area and values for the ratio of shoal volume and channel volume against the ratio of tidal amplitude and water depth) shows satisfying agreement. On the basis of the model results a relationship for a characteristic morphological wavelength was derived on the basis of the tidal excursion and the basin width and an exponentially varying function was suggested for describing a dimensionless hypsometric curve for the basin. Furthermore, special attention is given to an analysis of the numerical morphodynamic update scheme applied.

Accumulation of perfluorooctane sulfonate (PFOS) in the food chain of the Western Scheldt estuary: Comparing field measurements with kinetic modeling

The environmentally persistent perfluorooctane sulfonate (PFOS) is a perfluoroalkylated acid (PFA), which has been found to accumulate and biomagnify through food webs all over the world. In the present investigation, the accumulation kinetics of PFOS was explored using the bioaccumulation model OMEGA. As accumulation behavior of PFOS may show similarities to fatty acids as well as to neutral organic compounds, different modeling approaches were used. Accumulation kinetics of PFOS was modeled similar to (1) moderately and (2) highly hydrophobic compounds, (3) metals and (4) as a combination of hydrophobic compounds and metals. Modeled elimination and uptake rate constants were compared to empirical rate constants from literature. Subsequently, model predictions were compared to field-based biota-suspended solids accumulation ratios (BSAF) in the estuarine food chain of the Western Scheldt, The Netherlands. Results show that uptake of PFOS is comparable to moderately hydrophobic compounds and elimination is best described by elimination kinetics of metals. These observations indicate that the accumulation behavior of PFOS is comparable to that of short and medium chained fatty acids.

Effects of environmental stress on the condition of Littorina littorea along the Scheldt estuary (The Netherlands)

The condition of the periwinkle Littorina littorea, expressed in terms of its shell morphology, reproductive impairment (i.e. female sterility/intersex, male penis shedding), trematode infestation load, lipid reserves and dry/wet weight ratio, was determined in function of environmental stress along the polluted Western and relatively clean Eastern Scheldt estuary (The Netherlands). The upstream increasing pollution and decreasing salinity levels along the Western Scheldt estuary (Fig. 1) are reflected in the dry/wet weight ratio and lipid content of the periwinkles. Compared to the Eastern Scheldt, female intersex (i.e. indicator of TBT pollution) and sterility occurred more frequently in the Western Scheldt estuary, while male penis shedding was even restricted to the latter estuary. The highest population intersex and sterility incidence was found near the harbour of Vlissingen and reflects potential nautical activities. The number of trematode infested periwinkles did not differ between both estuaries, although local sampling site differences were detected within each estuary, reflecting the complex interactions that exist among parasites, hosts and the local environment. Finally, both estuaries were maximally discriminated from each other based on the shell weight of the periwinkles using a canonical discriminant analysis. Periwinkles with the heaviest shells were found in the Western Scheldt estuary and may reflect growth rate or structural population differences caused by the less favourable living conditions in the Western Scheldt estuary.

Spatio-temporal variation in energy content of the periwinkle Littorina littorea, along a pollution gradient in the western Scheldt estuary, The Netherlands

Energy reserves, expressed as lipid, glycogen and protein fractions, were measured in the periwinkle L. littorea after 4 weeks of translocation from a relatively clean, marine (i.e. Westkapelle) to an intermediate (i.e. Ellewoutsdijk) and a highly heavy metal polluted, brackish site (i.e. Hansweert) in the Western Scheldt estuary (The Netherlands). Energy reserves of resident periwinkles from all three sites were assessed over time as well (i.e. 16/03/05, 23/03/05, 30/03/05, 13/04/05 and 11/05/05). None of the energy fractions reflected the upstream decreasing salinity and increasing heavy metal gradients. Glycogen and protein fractions decreased significantly in the translocated specimens, despite no differences were detected among the resident populations, suggesting adaptation and/or acclimation effects at the upstream parts of the estuary. The temporal data revealed a consistent post-winter, pre-spawning increase of glycogen and protein levels and a subsequent drop after spawning. Compared to glycogen and proteins, lipids follow a slightly different pattern, which may be explained by sex related and site specific interactions between biotic and abiotic factors.

Metal Content of Marine Mussels from Western Scheldt Estuary and Nearby Protected Marine Bay, The Netherlands: Impact of Past and Present Contamination

Metal Content of Marine Mussels from Western Scheldt Estuary and Nearby Protected Marine Bay, The Netherlands: Impact of Past and Present Contamination

Biological influences on morphology and bed composition of an intertidal flat

Biological activity is known to influence sediment strength at bed–water interfaces. However, its precise effect on geomorphology and on bed composition is not known. This paper proposes a parameterization of sediment destabilizing and stabilizing organisms on three parameters that describe the erosion and mixing processes of the sediment bed, namely the critical bed shear stress, the erosion coefficient and the bioturbation coefficient. This parameterization is included in a 3D sand–mud morphodynamic model to form the sand–mud–bio model. The performance of the sand–mud–bio model is demonstrated by testing it on the Paulinapolder intertidal flat in the Western Scheldt estuary of The Netherlands. Model results show that biological influences on sediment strength result in significant morphological change and bed composition variations. Destabilizing organisms always cause a significant decrease in mud content in the bed and an increase of erosion. On the other hand, stabilizing organisms can, but do not necessarily, cause an increase of mud content and additional sedimentation.

Temporal and spatial trends in heavy metal concentrations in the marine mussel Mytilus edulis from the Western Scheldt estuary (The Netherlands)

Since the first North Sea Conference (1984, Bremen), all countries bordering the North Sea made commitments to reduce discharge of hazardous substances into the North Sea. From Belgium and The Netherlands, large reductions (upto 90) in heavy metal emissions from land-based sources have been reported between 1985 and 2000. Recently, some studies in the Western Scheldt estuary (WS) have shown that total metal concentration in the water, sediments and suspended particles have decreased compared to levels in the 70s. However, data on aquatic organisms is still very limited and it is therefore difficult to confirm whether the reductions in pollution input and generally improving water quality in the WS have a corresponding impact on the levels of heavy metals in aquatic organisms. The current study measured metal concentrations in the soft tissues of mussels, Mytilus edulis (known to be good indicators of environmental metal pollution) during the period 1996–2002. Spatial (salinity and pollution gradients), temporal and seasonal variations were also studied. Results showed a down-stream decreasing trend for the metals studied (Cd, Co, Cr, Cu, Fe, Mn, Ni, Pb and Zn) during all sampling campaigns. There was also a significant seasonal effect on tissue metal concentrations, with a peak observed around spring in both WS and the nearby less polluted Eastern Scheldt (ES). On temporal trends, a clear drop of metals in mussels was observed in the early 80s coinciding with the start of the efforts to reduce chemical pollution input into the North Sea. Since those early reductions, metal concentrations in mussels generally remained unchanged upto mid 90s. However, in recent times metal concentration in mussels have increased significantly, for example Cd in 2002 was almost 10 times the values in 1983 and similar to levels seen during the peak in the 70s. Other metals also increased in the 90s also reaching levels seen in the 70s. As there is no indication of recent increase in metal input into the estuary, we suggest that increased metal concentrations observed in mussels in recent years especially in the upper estuary are most likely a result of changes in physical and chemical speciation and metal bioavailability. Such changes may be caused by changes in some water quality parameters in the estuary (i.e. increased dissolved oxygen, concentration of organic matter), resulting in conditions that favour releases of sediment-bound metals into the water column. The relationship between metal content and season showed very similar annual profiles in the polluted WS and less polluted ES. Thus, seasonal variations in metal concentrations appear to be largely controlled by biological processes, while total body burdens are dependent on environmental levels and bioavailability.

Hexabromocyclododecane in Marine Species from the Western Scheldt Estuary: Diastereoisomer- and Enantiomer-Specific Accumulation

Hexabromocyclododecane (HBCD) is a widely used brominated flame retardant, which is increasingly reported in the environment. Here, we report on the diastereomeric and, for the first time, on the enantiomeric composition of HBCD in muscle and liver of several fish species caught in the Western Scheldt Estuary (The Netherlands). The total HBCD content (sum of alpha-, beta-, and gamma-diastereoisomers), as well as the distribution of diastereoisomers and enantiomers, varied between the species. The levels of total HBCD (9-1110 ng/g lipid weight) found in fish tissues were higher than those measured in fish from European rivers with no known point sources of HBCD but lower than in fish samples collected near factories producing or using HBCD. The concentrations of total HBCD expressed on a lipid weight basis were higher in liver than in muscle for bib and whiting, while in sole, HBCD had no preferential distribution between the tissues. A similar pattern for liver and muscle distribution was already observed for polybrominated diphenyl ethers (PBDEs) in these species. The alpha-HBCD diastereoisomer was most abundant in all fish samples with a higher contribution to the total HBCD levels in liver compared to muscle for bib and whiting. The gamma-HBCD diastereoisomer accumulated less in liver than in muscle of sole, bib, and whiting. For the first time, enantiomer fractions were determined for HBCD diastereoisomers in liver of three fish species and in muscle of two fish species. A significant enrichment of the (+) alpha-HBCD enantiomer was found in whiting and bib liver samples. A high enantioselectivity has also been seen for the gamma-HBCD diastereoisomer in whiting liver.

Levels and profiles of PCBs and OCPs in marine benthic species from the Belgian North Sea and the Western Scheldt Estuary

Various benthic invertebrates (flying crab, common shrimp, and red starfish), small fish (sand goby), benthic flatfish (dab, plaice, and sole) and gadoids (bib and whiting) were collected in the Belgian North Sea and along the Scheldt Estuary, both representing areas impacted by various contaminants to different degrees. The levels of 25 polychlorinated biphenyls (PCBs) and 15 organochlorine pesticides (OCPs), which included penta- and hexachlorobenzene, α-, β-, and γ-hexachlorocyclohexane isomers, chlordanes, and DDT and metabolites, were determined. Sum of PCBs and OCPs in benthic invertebrates and goby ranged from 1.5 to 280 ng/g wet weight (ww) and from 0.27 to 23 ng/g ww, respectively. The fish livers revealed total PCB and OCP levels ranging from 20 to 3200 ng/g ww and from 6.0 to 410 ng/g ww, respectively. Levels of both contaminant groups were significantly higher in samples from the Scheldt Estuary compared to the Belgian North Sea. For most species a highly inverse correlation was found between the concentration of contaminants and the distance to Antwerp ( r between 0.812 and 0.901, p<0.05), pointing to a higher degree of exposure further upstream. PCB and OCP exposures are highly correlated ( r between 0.836 and 1.000, p<0.05), which suggests that the pollution can be classified as historical. However, because urban and industrial centres may still be emitting these compounds, more recent point and non-point sources cannot be ruled out.

PBDEs in marine and freshwater sediments from Belgium: levels, profiles and relations with biota

Sediments from the Belgian North Sea (BNS), the Western Scheldt Estuary (SE) and freshwater watercourses from the Scheldt basin were analysed for eight PBDE congeners, namely BDEs 28, 47, 99, 100, 153, 154, 183 and 209. Previously analysed biological samples from the same locations in the BNS and the SE have been shown to contain large amounts of PBDEs. Surprisingly, PBDE concentrations in the sediments were below the LOQ for samples from the BNS (except BDE 209), while in those from the SE the sum of PBDEs (not including BDE 209) were higher and ranged from 0.20 to 0.41 ng g(-1) dw. BDE 209 could be detected in 83% of the samples from the BNS and in all the samples from the SE. Concentrations up to 1200 ng g(-1) were hereby measured in the SE. Compared to the marine and estuarine locations, the sediments from the freshwater watercourses were relatively more polluted with the lower brominated PBDEs (<0.20-19 ng g(-1) dw). BDE 209 concentrations up to 320 ng g(-1) dw were measured in those sediments. However, the contribution of BDE 209 to the total amount of PBDEs varied much more at the freshwater locations than in the SE, which suggests a different input of pollutants. PBDE profiles observed in biological samples do not match the profiles of the sediments. BDE 183 and 209 could not be quantified in biota, although these congeners were undoubtedly present in the sediments. This raises questions about the bioavailability of these congeners in the environment.

Sensitivity to cadmium along a salinity gradient in populations of the periwinkle, Littorina littorea, using time-to-death analysis

In this study, we assessed the combined effect of Cd concentration and salinity, on Cd uptake and mortality rate of Littorina littorea, collected along a salinity and pollution gradient in the Western Scheldt estuary (The Netherlands). Animals kept at their field salinity levels were exposed to three Cd concentrations (i.e. 10, 40 and 320 μM), while animals kept in 10 μM of Cd were subjected to five salinity treatments (i.e. 15, 20, 25, 30 and 35‰). Mortality was recorded every 24 h and Cd body burdens were measured with ICP-AES. Time-to-death data were analysed via Cox proportional hazard models, including the co-variates “site-Cd treatment” in the Cd experiment and “site-salinity treatment” in the salinity experiment. “Cd-treatment” and “field-salinity” affected mortality rates significantly in the Cd experiment, such that the mortality risk increased by 2.3 times when salinity was lowered from 35 to 15‰, while it decreased by 19.7 times when Cd dropped from 320 to 10 μM. “Site” did not significantly affect the mortality risk in the salinity experiment but affected time-to-death via its interaction with the “salinity-treatment”. Generally, mortality did not occur at a given threshold Cd tissue level, but changed over time and treatments, in function of the site. The results demonstrate the importance of the animals’ environmental history and illustrate the usefulness of time-to-death analyses in ecotoxicological experiments.

Polybrominated diphenyl ethers in marine species from the Belgian North Sea and the Western Scheldt Estuary: levels, profiles, and distribution.

The Western Scheldt Estuary (SE) is subjected to a variety of suspected PBDE sources, such as a brominated flame retardant manufacturing plant, the Antwerp harbor, and the textile industry located further upstream the river. The Belgian North Sea (BNS) was included in this study to analyze the influence of the SE on the levels found in biota from the BNS locations. Benthic invertebrates, such as shrimp, crab, and starfish, benthic fish, such as goby, dab, plaice, and sole, and gadoid fish, such as bib and whiting, were sampled in the BNS (nonpolluted area) and the SE (polluted area) and analyzed to determine the concentrations and spatial variation of eight polybrominated diphenyl ethers (PBDEs 28, 47, 99, 100, 153, 154, 183, and 209). Levels found in the SE samples were up to 30 times higher than those found in BNS samples, with a gradient increasing toward Antwerp. Levels in BNS ranged from 0.02 to 1.5 ng/g ww in benthic invertebrates and goby, from 0.06 to 0.94 ng/g ww in fish muscle, and from 0.84 to 128 ng/g ww in fish liver. For the SE samples, levels ranged from 0.20 to 29.9 ng/g ww in benthic invertebrates and goby, from 0.08 to 6.9 ng/g ww in fish muscle, and from 15.0 to 984 ng/g ww in fish liver. BDE 209 could only be detected in eight liver samples from the SE and levels ranged between 3.4 and 37.2 ng/g ww. PBDE profiles of the various species at the different locations were compared. Differences in profile were attributed to different exposure and to differences in metabolism among species. Ratios between BDE 99 and 100 were found to be highly location and species dependent, which could be related to differences in metabolism. Some species, such as dab, plaice bib, and whiting, showed preferential accumulation of PBDEs in the liver. Higher brominated congeners in general showed higher affinity for liver than for muscle tissue.

Exposure patterns of perfluorooctane sulfonate in aquatic invertebrates from the Western Scheldt estuary and the southern North Sea.

Over the past decades little research has been conducted on the environmental behavior and effects of fluorinated organochemicals (FOCs). Recently it has been reported that perfluorooctane sulfonic acid (PFOS) is occurring worldwide. Little is known about the PFOS levels in organisms originating from the southern North Sea and the Western Scheldt estuary. In this study, we determined, for the first time, the PFOS-exposure levels in Crangon crangon, Carcinus maenas, and Asterias rubens from these ecosystems. Concentrations on a wet-weight basis in soft tissues of shrimp, crab, and starfish ranged from 19 to 520 ng/g, from 24 to 877 ng/g, and from 9 to 176 ng/g, respectively. These results show the existence of a PFOS pollution gradient in organisms along the Western Scheldt estuary, with the highest concentrations near Antwerp. The range of PFOS levels in shrimp and crab are slightly higher in coastal regions compared with sampling sites in open water. This study shows widespread distribution of PFOS in the Belgian and Dutch marine and estuarine environment at rather high concentrations.

The North-American amphipods, Melita nitida Smith, 1873 and Incisocalliope aestuarius (Watling and Maurer, 1973) (Crustacea: Amphipoda: Gammaridea), introduced to the Western Scheldt estuary (The Netherlands)

The American amphipod species Melita nitida andIncisocalliope aestuarius have been found in the WesternScheldt estuary (the Netherlands). This is the first record of these species inthe north-east Atlantic. Shipping is the most likely vector of introduction.Thedistribution of both species is investigated and compared with the distributionand the microhabitat of co-occurring amphipod species. Melitanitida is known from both the east and west coast of North Americaand I. aestuarius originates from the east coast of NorthAmerica. Until now neither has been reported from other parts of the world. Inthe Netherlands both species are restricted to the mesohaline part of theWestern Scheldt. Melita nitida occurs predominantly underPacific oysters at the underside of boulders, mainly sublittorally.Incisocalliope aestuarius is associated to hydrozoans.Bothmicrohabitats are hardly utilized by other amphipod species. Therefore, thetheory that the existence of many empty niches in north-western Europeanbrackish waters make this environment particularly susceptible to invasions ofalien species is corroborated. The application of hard substrates in a regionoriginally predominated by soft bottoms moreover facilitates the introductionofexotic species. The species community on hard substrates in the mesohaline partof the Western Scheldt contains a high proportion of introduced species:approximately one third of the macrofauna species is of allochthonous origin.

Morphodynamic equilibrium in straight tidal channels: Combined effects of the Coriolis force and external overtides

A new physical mechanism that is potentially relevant for the equilibrium morphodynamics of tide‐dominated estuaries and in understanding the occurrence of lateral shoals in these systems is identified. The mechanism acts in relatively straight and wide channels (width of the order of the horizontal tidal excursion length) in which both external overtides and the Coriolis force affect sediment transport. This is investigated by analyzing an idealized model, which consists of the 3D shallow water equations, mass conservation for suspended load, and a bed evolution equation. The model is forced by a prescribed depth‐averaged tidal current. It is demonstrated that, when viewed in the direction of the flood flow, a flood (ebb)‐dominant current generates a net cross‐sectional sediment transport to the left (right) in the Northern Hemisphere. A morphodynamic equilibrium is established by a counteracting dispersive sediment flux, generated by shear stresses that increase toward shallower water. This dispersive flux is much larger than the flux due gravitational downslope effects. The equilibrium bed profile has a constant slope in the lateral direction that varies as cos(φ), where φ is the phase difference between the M2 and M4 external horizontal tide. Hence, the smallest depths are found on the left (right) in case of a flood (ebb)‐dominant current. Typical cross‐channel depth differences may be as large as several meters. Velocity data collected in the Dutch Western Scheldt estuary are used to tune the hydrodynamic parameters in the model. Analysis of the bathymetric data seems to confirm the qualitative results of the model.