Western Interior Research Articles

A Climatology of Dryline‐Related Convection on the Western Plateau of Subtropical Southern Africa

AbstractDrylines occur regularly during summer over the interior of sub‐tropical southern Africa. The highest frequency of drylines is over the western interior where it is also climatologically much drier than the eastern parts. An existing drylines data set is compared with lightning data between 2010 and 2021, to determine how often drylines were associated with convective storms. A minimum stroke threshold is applied to eliminate instances of weak convection. Seventy‐six percent of drylines during austral summer (October–March) were associated with significant convection with a standard deviation of 8.6. There is a peak frequency of 12 convective dryline days during December. Composite synoptic analyses were conducted for 591 convective and 186 non‐convective dryline days to determine the ideal formation environment for convective drylines. Composites were also created for the days leading up to convective and non‐convective drylines, showing the importance of moisture advection from the southwest Indian Ocean, through the Zambezi and Limpopo River Valleys. Early summers tend to be drier than late summers over the southern African interior and colder in the mid‐levels. These differences affect the formation environment of convective drylines. Composites show that dryline‐related convection during early summer tends to occur on days with a deep surface trough and a trough at 500 hPa further west. Convection during late summer is associated with increased surface moisture, higher instability, and a weaker trough at 500 hPa than during early summer.

Paleoenvironmental reconstruction of Late Cretaceous rivers, Dinosaur Park Formation, Alberta, Canada

Dinosaur Provincial Park is a UNESCO World Heritage Site with one of the richest vertebrate fossil accumulations globally. The “badlands”-style topography exposes Upper Cretaceous (Campanian) meander-belt deposits of the Dinosaur Park Formation, providing an excellent opportunity for sedimentological and three-dimensional stratigraphic analysis. One of the main unknowns regarding the depositional environments of the Dinosaur Park Formation, specifically the proximity to the fluvial-to-marine transition zone, has been the subject of extensive debate. Our study aims to refine paleoenvironmental reconstructions of Late Cretaceous rivers through facies and facies associations analysis, meander-belt architecture characterization, and paleohydraulic parameter calculations.We investigate a ~7 m thick single-story meander-belt interval of the Dinosaur Park Formation. Six facies, four facies associations, and two architectural elements (both point bars) were identified from 20 stratigraphic sections. In addition, paleohydraulic reconstruction of the formative river channel was completed by estimating bankfull channel depth and width, paleoslope, and backwater length.Contrasting with previous interpretations of deposition (partially or wholly) influenced by tides, these data allow us to demonstrate that the meander-belt deposits of the Dinosaur Park Formation were deposited in a fluvial environment upstream of the fluvial–marine transition zone. Inclined heterolithic strata (IHS) present in the area are of fluvial origin, adding to the growing body of evidence for heterolithic bedding in fluvial point-bar environments. The newly inferred depositional setting for the studied portion of the Dinosaur Park Formation changes our understanding of the paleogeography of the Western Interior Seaway in western Canada and will allow paleontologists to revise the reconstructions of dinosaur habitats and better constrain the taphonomy of these globally significant fossil accumulations. Additionally, this study contributes to the validation and development of a systematic approach to the paleoenvironmental reconstruction of ancient river systems based on paleohydraulic analysis.

Integrated statistical analysis of calcareous nannofossil and elemental geochemistry of an outcrop from of the eastern Cenomanian Western Interior Seaway: Novel insights of shallow marine paleoceanography and nannoplankton paleoecology

Phytoplankton assemblages in shallow marine environments are being impacted by anthropogenic climate change, but long-term outcomes of these changes are uncertain. Investigation of past neritic calcareous nannoplankton can help us understand the fate these ecosystems face. In this study, a Canonical Correspondence Analysis (CCA) of calcareous nannofossils and X-ray fluorescence geochemistry was used to determine how past planktonic ecosystems were influenced by paleoenvironmental parameters on the eastern side of the Late Cretaceous Western Interior Seaway in the Cenomanian (ca. 95–93 Ma). Samples were collected every 10 cm from the Graneros Shale Greenhorn Formation at an outcrop in northwestern Iowa to determine high resolution changes in assemblages and paleoenvironments. Nannofossil diversity outside of a few small intervals ranges is high (generally 30–60 species) with abundant small Biscutum constans, confirming other publications that show elevated diversity in Cretaceous nearshore settings. The CCA results imply assemblages were most influenced by terrigenous influence, wet vs. dry climate, and changes in water mass source. Cretaceous nannofossil paleoecology was also revised based on the CCA results. Size differentiation of nannofossil taxa may highlight more complexity in environmental preferences that have been largely overlooked. After the initial transgression of the Greenhorn Sea into the region, the climate became wetter and terrigenous influence in the area was high. The peak of terrigenous influence corresponds with elevated nannofossil diversity but a lack of microfauna, which may indicate a similar oceanographic setting to the modern Gulf of Mexico nearshore dead zone. As sea level continued to rise above this point, nannoplankton assemblages indicate a potential shift to a higher productivity, stratified water column. As the muddy Graneros Shale transitioned to a further offshore chalky Greenhorn Formation, a normal marine, cosmopolitan nannofloral assemblage became established. Nannofossil and geochemical evidence indicates high productivity from upwelling might be related to the change of deposition to chalk in the Greenhorn Formation. While only a single outcrop was investigated, the novel use of an integrated micropaleontological and geochemical analysis has shed light on the dynamics of how phytoplankton ecosystems were established and modified in shallow marine environments of the Cretaceous and could have important implications on modern shallow marine settings.

Exceptional age constraint on a fossiliferous sedimentary succession preceding the Cretaceous Thermal Maximum

Abstract Understanding the effects of climatic upheavals during the Early to Late Cretaceous transition is essential for characterizing the tempo of tectonically driven landscape modification and biological interchange; yet, current chronostratigraphic frameworks are too imprecise, even on regional scales, to address many outstanding questions. This includes the Mussentuchit Member of the uppermost Cedar Mountain Formation, central Utah (southwestern United States), which could provide crucial insights into these impacts within the Western Interior Basin of North America yet remains imprecisely constrained. Here, we present high-precision U-Pb zircon dates from four primary ash beds distributed across ~50 km in central Utah that better constrain the timing of deposition of the Mussentuchit Member and the age of entombed fossils. Ages for ash beds are interpreted through a combination of Bayesian depositional age estimation and stratigraphic age modeling, resulting in posterior ages from 99.490 + 0.057/–0.050 to 98.905 + 0.158/–0.183 Ma. The age model predicts probabilistic ages for fossil localities between the ashes, including new ages for Moros intrepidus, Siats meekerorum, and several undescribed ornithischian dinosaur species of key interest for understanding the timing of faunal turnover in western North America. This new geochronology for the Mussentuchit Member offers unprecedented temporal insights into a volatile interval in Earth’s history.

Geological Assessment of Turonian - Coniacian terrestrial sedimentation records during climatic recovery, Moreno Hill Formation, Zuni Basin

Understanding of Upper Cretaceous terrestrial sediments within the Western Interior Basin is advancing; however, the Turonian–Coniacian transition remains enigmatic. Recent chronometry of the Moreno Hill Formation indicates that sediment deposition took place during this interval of geodynamic upheaval and climatic recovery immediately following the peak of the Cretaceous Thermal Maximum (CTM). To decipher these effects, the sedimentary record of the Moreno Hill Formation was reassessed near the type section, and near Quemado, New Mexico (USA) using facies analysis and architectural reconstruction. Seven facies types (thirteen lithofacies codes), eleven architectural elements, and three facies associations were identified. Sedimentation within the floodplain of the lower Moreno Hill Formation was affected by the east-migrating forebulge of the Western Interior Basin. Furthermore, increasingly bedload-rich multi-story channel complexes and a transition from near-coastal to alluvial coals reflect gradual climatic cooling and overall regression of the Western Interior Seaway (with interruption by a regional [T2–R2] transgressive-regressive sequence). This is consistent with more subaerial conditions indicative of continued regression reflected within the floodplain sediments of the upper Moreno Hill Formation. Whilst diversion of westerly fluvial feeder systems by ongoing forebulge migration also affected sediment transport and deposition, a return to more suspended-load-rich single-story channels and thin coals are tied to an intervening (T3) transgression. Repetitive paleosol sequences throughout the Moreno Hill Formation indicate groundwater fluctuation in response to these base level changes. Together with detrital zircon-based geochronology, these slight sedimentary differences support a revised subdivision from three into two members: lower and upper. Beyond feeding the seaward Gallup Delta, the newly defined lower member correlates to the Toreva, Straight Cliffs (Smoky Hollow member), Ferron Sandstone, Funk Valley and Frontier formations (Dry Hollow Member) and the upper member to the Wepo and Straight Cliffs (John Henry Member) formations within the Kaiparowits, Notom, Last Chance and Vernal fluvio-deltaic systems. Landward sediments of the Cardium Formation (Canada) correlate with the lower and upper members of the Moreno Hill Formation.

The “Judith River−Belly River problem” revisited (Montana-Alberta-Saskatchewan): New perspectives on the correlation of Campanian dinosaur-bearing strata based on a revised stratigraphic model updated with CA-ID-TIMS U-Pb geochronology

Terrestrial strata of the Judith River−Belly River wedge, widely exposed in the plains of north-central Montana, southern Alberta, and southwestern Saskatchewan, were pivotal in early stratigraphic investigations of the Western Interior of North America and are renowned to this day for their spectacular preservation of Late Cretaceous fossils, most notably dinosaurs. Correlation of the Judith River Formation in Montana with the Foremost, Oldman, and Dinosaur Park Formations (= Belly River Group) in Canada has been challenging for a variety of reasons, including lithostratigraphic complexities, legacy bentonite ages of limited comparability, and distinctly different stratigraphic models on opposite sides of the international border. An updated model calibrated with U-Pb zircon ages provides an improved framework for stratigraphic analysis. New geochronology indicates that the Oldman−Dinosaur Park discontinuity in Dinosaur Provincial Park correlates in age with the mid-Judith discontinuity in the Judith River Formation in Montana, which is interpreted as an expansion surface linked to a major pulse of accommodation and onset of the Bearpaw transgression at ca. 76.3 Ma. The regionally expressed shift in alluvial facies marking the mid-Judith discontinuity can be traced in well logs from Montana to southern Canada, where it loses distinction and transitions to a subsurface signature typical of the Oldman−Dinosaur Park discontinuity, which in turn can be traced north to Dinosaur Provincial Park and beyond. Across this expanse, both discontinuities parallel the Eagle/Milk River shoulder at approximately the same stratigraphic height, confirming their chronostratigraphic significance. These findings have clear implications for regional correlation and the evolution of alluvial depositional systems in a foreland basin setting, and they afford an opportunity to evaluate existing interpretations and advance understanding of the stratigraphy and paleontology of the Judith River−Belly River wedge. The term “Judith River−Belly River discontinuity” should be used henceforth to refer to the chronostratigraphically significant stratal discontinuity that subdivides the Judith River−Belly River wedge throughout the plains of north-central Montana, southern Alberta, and southwestern Saskatchewan.

Late Holocene Technology Words in Proto-Athabaskan: Implications for Dene-Yeniseian Culture History

This study will review previously published Proto-Athabaskan (P-A) linguistic reconstructions related to weapons and ceramics technologies present on both sides of the Bering Strait. Na-Dene (N-D) is a large family of indigenous languages of North America, consisting mostly of the Athabaskan languages of the western interior, plus the Eyak and Tlingit languages of the southern Alaska coast. Athabaskan-Eyak (A-E) constitutes a distinct branch of Na-Dene. Dene-Yeniseian (D-Y) is a proposed transpacific family comprised of Na-Dene in addition to the Yeniseian languages of Siberia. Reconstructions pertaining to several specific technologies will be discussed in relation to likely cognates within broader A-E, N-D and D-Y historical contexts. Although D-Y is sometimes assumed to have originated near the conclusion of the Pleistocene Epoch (prior to ~11,500 years BP), this study will refocus fundamental questions on the current Holocene Epoch (after ~11,500 BP), and particularly the Late Holocene (after ~3000 BP).

Calibrating geologic strata, dinosaurs, and other fossils at Dinosaur Provincial Park (Alberta, Canada) using a new CA-ID-TIMS U–Pb geochronology

The 100 m thick stratigraphic section exposed at Dinosaur Provincial Park (DPP; southern Alberta) contains bentonites that have been used for more than 30 years to date DPP’s rocks and fossils using the K–Ar decay scheme. Limited reproducibility among different vintages of K–Ar and 40Ar/39Ar ages inhibited the development of a high-resolution chronostratigraphy. Here, we employ and further test a recently completed U–Pb geochronology and associated age-stratigraphy model to update temporal constraints on the Park’s bentonites, formational contacts, and other markers. In turn, we document rock accumulation rates and calibrate ages and durations of informal megaherbivore dinosaur assemblage zones and other biozones. Weighted mean 206 Pb/238 U ages from five bentonites range from 76.718 ± 0.020 to 74.289 ± 0.014 Ma (2σ internal uncertainties) through an interval of 88.75 m, indicating a duration of ∼2.43 Myr and an overall rock accumulation rate of 3.65 ± 0.04 cm/ka. An increase in rate above the Oldman–Dinosaur Park formational contact conforms to a regionally expressed pattern of increased accommodation at ∼76.3 Ma across Alberta and Montana. Palynological biozone data suggest a condensed section/hiatus in the uppermost portion of the Oldman Formation. Dinosaur assemblage zones exhibit durations of ∼700–600 kyr and are significantly shorter than those in the overlying Horseshoe Canyon Formation. A decreased rate in dinosaur assemblage turnovers in the last eight million years of the Mesozoic in western Canada may be explained by withdrawal of the Western Interior Seaway and the expansion of ecologically homogenous lowlands in its wake.

Faunal provinciality in the Late Cretaceous Western Interior Seaway using network modeling

The Western Interior Seaway (WIS) was historically divided into latitudinal faunal provinces that were taxonomically distinct from the adjacent Gulf Coastal Plain (GCP) and that shifted in space due to sea-level changes. However, no rigorous quantitative analyses using recent taxonomic updates have reassessed these provinces and their associations. We used network modeling of macroinvertebrate WIS and GCP fauna to test whether biotic provinces existed and to examine their relationships with abiotic change. Results suggest a cohesive WIS unit existed across the Campanian, and distinct WIS and GCP provinces existed in the Maastrichtian. Sea-level changes coincided with changes in network metrics. These results indicate that, while the WIS did not contain subprovinces in the Late Cretaceous, environmental factors influenced faunal associations and their communication over time.

Productivity-induced redox transition within the Niobrara formation, western Interior seaway, Colorado

Deposition of organic rich shale/marl within the Niobrara Formation during the Coniacian to Santonian Ocean anoxic event (OAE 3) within the Western Interior Seaway presents a unique record of variable organic carbon burial under a greenhouse climate. In order to understand the mechanisms behind the changes from low to high organic carbon burial, and how this high burial rate was sustained for over 3 Ma, productivity within the surface waters and subsequent preservation of organic matter within the sediments were investigated in this study. Based on the multiproxy approach adopted herein, the sedimentary record of the Niobrara Formation in northern Colorado can be separated into two distinct intervals: Interval 1 and Interval 2. Interval 1 is bioturbated, organic lean and oxic, with low to moderate trace metal enrichment except for Mn. Interval 2 is laminated, organic-rich, more enriched in trace metals U, V and Mo, and less enriched in Mn, typical of anoxia during deposition. The transition from oxic to anoxic conditions coincides with greater enrichments in TOC, Cu and Ni, suggesting enhanced productivity is associated with redox transitions within the Niobrara Formation. These local changes in productivity and redox conditions are sudden and coincide with recurring iron (Fe) and phosphorus (P)-rich bentonites, evidence of ash deposition that could have enhanced nutrient supply, productivity and accumulation of organic carbon. Consequently, perturbations of the global carbon cycle associated with OAE 3 were inferred to have occurred in Interval 2.High total Fe, total sulfur, Fe/Al, and trace metal enrichments in Interval 2 are intrinsically linked to prevalence of anoxic bottom-water conditions which enhanced burial of pyrite and organic matter. Results from productivity proxies and Fe–S–C ternary plots show that Fe limitation on pyrite formation under anoxic porewater enhanced organic carbon burial and preservation. This implies Fe enrichment within the sediments favored pyrite formation instead of organic matter sulfidization, leading to a complex relationship between S and organic carbon in the marine sediments during Niobrara deposition.

BENTHIC FORAMINIFERAL MORPHOGROUPS AND TRACE FOSSILS FROM ALBIAN SHALLOW-MARINE MUDSTONE FACIES OF THE VIKING FORMATION, WESTERN INTERIOR SEAWAY, CANADA

ABSTRACT This study assesses the foraminiferal morphogroup concept as a quantitative tool for interpreting environments of deposition in the geological record. This was achieved by first establishing a solid paleoenvironmental framework based on assessment of trace fossils (Zoophycos, Cruziana, and Phycosiphon ichnofacies, and the unnamed brackish-water association), sedimentology, and facies analysis of shallow-marine siliciclastic units of the upper Albian Viking Formation of the Western Interior Seaway, Canada. Foraminiferal morphogroups were integrated into a framework of six well-defined marine mudstone-dominated facies—estuarine, embayment, prodelta, upper offshore, lower offshore, and marine shelf. Seven morphogroups were recognized, quantified, and labeled using alphanumeric codes—A (tubular), B1 (globular), B2 (coiled, flat), B3 (lenticular), C1 (elongate), C2 (quinqueloculine), and D (trochospiral). Tubular forms (A, Bathysiphon) occur rarely in these shallow-marine facies. Globular forms (B1, Lagenammina) are common in estuarine (33%) and embayment (23%) facies, but also occur elsewhere in low numbers (7–14%). Coiled planar forms (B2, Ammodiscus) occur only rarely (1–8%), but have a minor presence in embayment facies (8%). Lenticular forms (B3, Haplophragmoides) are dominant in estuarine facies (43%) and occur commonly in other environments (9–29%) as well. Elongate forms (C1, Ammobaculites, Reophax, Verneuilinoides) dominate all shallow-marine facies (43–80%), except estuarine (10%). Quinqueloculine forms (C2, Miliammina) have a minor presence in prodelta and marine facies (5%) but are generally rare. Trochoid forms (D, Trochammina) occur rarely, but are relatively common in the prodeltaic facies (15%). Albian foraminiferal morphogroups of the Viking Formation are comparable to those of modern-day offshore environments from water depths of 13 to 160 meters. Foraminiferal morphogroups are objective entities independent of taxonomic determinations. Analysis of their form and function is applicable not only in the shallow-marine facies of the Albian Western Interior Seaway, but in similar facies throughout the geologic record.

Sand Dragons: Species of the Ctenophorus maculatus Complex (Squamata: Agamidae) of Australia's Southern and Western Interior

Sand Dragons: Species of the Ctenophorus maculatus Complex (Squamata: Agamidae) of Australia's Southern and Western Interior

A new lower Turonian mosasaurid from the Western Interior Seaway and the antiquity of the unique basicranial circulation pattern in Plioplatecarpinae

We describe and name a new mosasaur taxon, Sarabosaurus dahli gen. et sp. nov., from the lower Turonian part of the Tropic Shale in Utah, USA. The holotype specimen preserves significant portions of the skull and axial postcranial skeleton. It was found in the upper part of the Watinoceras devonense Ammonite Zone, bounded by radioisotopic dates above and below, and is thus about 93.7 Ma, the oldest mosasaurid taxon known from the Western Interior Seaway. The new taxon possesses a vascular pattern of the basisphenoid heretofore only seen in late diverging plioplatecarpine mosasaurids. Reevaluation of the morphology of the basisphenoid of previously described Turonian mosasaurs using μCT techniques reveals the derived condition is also present in Yaguasaurus and the incipient condition in Tethysaurus and Russellosaurus. In these two taxa, the canals enter the basisphenoid, but do not pass into the basioccipital. Instead, they exit only high on the posterior wall of the sella turcica, in a position similar to the basilar artery of other lizards. This vascular pattern, both in its incipient and derived states, is unique among squamates and supports inclusion of the aforementioned taxa in a monophyletic Plioplatecarpinae, for which we provide an emended diagnosis. Phylogenetic analysis recovers Sarabosaurus dahli gen. et sp. nov. as the sister taxon to Yaguarasaurus and all other later diverging plioplatecarpines, with Russellosaurus and Tethysaurus as successive sister taxa. Tylosaurine mosasaurids retain the primitive condition of the basisphenoid vascularization pattern and implies a tylosaurine-plioplatecarpine divergence in the late Cenomanian or earliest Turonian.

An early-diverging iguanodontian (Dinosauria: Rhabdodontomorpha) from the Late Cretaceous of North America.

Intensifying macrovertebrate reconnaissance together with refined age-dating of mid-Cretaceous assemblages in recent decades is producing a more nuanced understanding of the impact of the Cretaceous Thermal Maximum on terrestrial ecosystems. Here we report discovery of a new early-diverging ornithopod, Iani smithi gen. et sp. nov., from the Cenomanian-age lower Mussentuchit Member, Cedar Mountain Formation of Utah, USA. The single known specimen of this species (NCSM 29373) includes a well-preserved, disarticulated skull, partial axial column, and portions of the appendicular skeleton. Apomorphic traits are concentrated on the frontal, squamosal, braincase, and premaxilla, including the presence of three premaxillary teeth. Phylogenetic analyses using parsimony and Bayesian inference posit Iani as a North American rhabdodontomorph based on the presence of enlarged, spatulate teeth bearing up to 12 secondary ridges, maxillary teeth lacking a primary ridge, a laterally depressed maxillary process of the jugal, and a posttemporal foramen restricted to the squamosal, among other features. Prior to this discovery, neornithischian paleobiodiversity in the Mussentuchit Member was based primarily on isolated teeth, with only the hadrosauroid Eolambia caroljonesa named from macrovertebrate remains. Documentation of a possible rhabdodontomorph in this assemblage, along with published reports of an as-of-yet undescribed thescelosaurid, and fragmentary remains of ankylosaurians and ceratopsians confirms a minimum of five, cohabiting neornithischian clades in earliest Late Cretaceous terrestrial ecosystems of North America. Due to poor preservation and exploration of Turonian-Santonian assemblages, the timing of rhabdodontomorph extirpation in the Western Interior Basin is, as of yet, unclear. However, Iani documents survival of all three major clades of Early Cretaceous neornithischians (Thescelosauridae, Rhabdodontomorpha, and Ankylopollexia) into the dawn of the Late Cretaceous of North America.

First Cretaceous teleostean otolith assemblage (Arkadelphia Formation, upper Maastrichtian) from Arkansas, USA, early Gadiformes, and the Western Interior Seaway

The fortuitous discovery of Cretaceous (late Maastrichtian) teleostean otoliths in boring samples (17–31 m below ground level) from the Arkadelphia Formation near Cabot, Arkansas, USA, has consequential and overarching ramifications. The otolith assemblage, which is relatively large with 2,109 specimens, represents the first Mesozoic otolith assemblage described from Arkansas and one of the largest Cretaceous assemblages from a single USA site. The diversity of the assemblage is fairly large with a richness of 19 species with three additional taxa in open nomenclature and one unknown lapillus, which more than doubles the known actinopterygians from the Arkadelphia Formation. The otolith assemblage is extremely uneven in its diversity with one species, a putative siluriform Vorhisia vulpes Frizzell (1965b), accounting for approximately 73% of the total. The most unique feature of the otolith assemblage is the presence of cool-water gadiforms,  which represent approximately 7.6% of the total assemblage. The presence of the gadiforms is related to the effects of the Western Interior Seaway and paleogeography during the Late Cretaceous in the western Gulf Coastal Plain. The gadiforms may represent relicts of a greater population and distribution in the early Maastrichtian. Percentage similarity measurements of the otolith assemblage indicate that the Arkadelphia Formation is more closely related to the Severn Formation in eastern Maryland (57.86%) and the  Kemp Clay Formation in northeast Texas (35.77%) than to the Ripley Formation in northeastern Mississippi (5.34%). The similarity measurements and other factors indicate that the Arkadelphia Formation otolith assemblage belongs to the Western Interior Seaway community (bioprovince). The Arkadelphia Formation otolith assemblage also contains several taxa that become extinct, such as the ubiquitous V. vulpes, at the K-Pg extinction event. The otoliths point to a very shallow marine environment (possibly inner shelf; less than 20 m in depth) with estuarine and freshwater input nearby and may be utilized for refinement of paleoshorelines for the southern reaches of the Western Interior Seaway during the Late Cretaceous.

A large, new dercetid fish (Teleostei: Aulopiformes) from the Campanian Bearpaw Formation of Alberta, Canada

Two new fossil fish specimens were recovered from deposits of the Bearpaw Formation of southern Alberta, Canada. These deposits represent the Western Interior Seaway in the Campanian. The remains can be confidently assigned to the aulopiform genus Dercetis in the extinct family Dercetidae. The two specimens are here described as Dercetis magnificus sp. nov. Based on a phylogenetic analysis, this new species is placed as the sister-group to the only other valid species of the genus, D. elongatus and D. triqueter. The genus Dercetis was previously documented from the western area of the Tethys and the Western Interior Seaway by only a small individual from Mexico and an incomplete braincase from Alberta, Canada, neither of which was assigned to a particular species. The new species is represented by specimens that are larger than the any other previously reported Dercetis, documenting the potential to attain a very large body size in the genus Dercetis and Dercetidae.

Selective extinction at the end-Cretaceous and appearance of the modern Decapoda

Abstract Extinction and origination of genera and families of marine Decapoda at the end of the Cretaceous established the modern fauna. Podotrematous crabs suffered high extinction levels, whereas heterotrematous crabs experienced both extinction and radiation. Anomuran decapods exhibited high extinction levels at the generic but not family level. In general, family extinctions predominantly occurred among podotrematous and heterotrematous crabs, which also exhibit overall shorter family ranges. Possible refugia promoted survival of genera and families into the Paleocene, whereas habitation of the Western Interior Seaway ensured high levels of extinction. Areas proximal to the Chicxulub impact site experienced moderate levels of extinction. Habitat specialists preferentially survived the end-Cretaceous extinction in some clades. Heterotrematous crabs recovered rapidly in the Paleocene, especially in the siliciclastic environments of the Americas. Because decapod crustaceans are highly variable in morphology, habitat, and ecology, no one pattern of extinction and recovery can explain the end-Cretaceous decapod extinction event. Rather, a mosaic of responses in various decapod groups led to their differential survival.

Paleoclimate and the origin of two 1000 km Lower Mississippian facies tracts in southeastern Laurentia (USA): Cool-humid Famennian and Kinderhookian – warm-arid Osagean

Two lithologically distinctive Mississippian facies tracts (MFTs) are described from Lower and Middle Mississippian strata (Kinderhookian and Osagean North American Stages) in southeast Laurentia (SEL). The MFTs extend from the central Appalachian Basin westward across the Cincinnati Arch, through the Eastern (Illinois) Basin, onto the Burlington shelf in central Missouri and into the Western Interior (Forest City) Basin, USA. The MFTs developed under radically different climatic regimes. Kinderhookian climate cycles ranged from 3rd order humid to 4th and 5th order humid-subhumid alternation. These climate fluctuations controlled Kinderhookian sediment flux, evidenced by fluvial-deltaic sandstones, coal beds, and pro-deltaic terrestrial-organic-matter-enriched marine black shales. A dramatic climate shift coincided with the Kinderhook-Osage boundary, from 3rd order cool-humid to 3rd order warm-arid conditions, causing cessation of deltaic sedimentation and the onset of eolian sedimentation at that boundary. This abrupt climate reorganization is reflected in 3rd, 4th, and 5th order continental and eolo-marine loessites that replaced fluvial-deltaic facies. Eolianites, evaporites, and calcareous protosols indicate Osagean aridity. Consequently, we reject the deltaic depositional paradigm for Osagean siliciclastic facies. Osagean eolian sediment consisted mainly of quartz silt with significant pristine (unaltered) feldspar silt (∼ ≤ 10%). Pristine feldspars are consistent with an arid sedimentary source that lacked significant chemical weathering under an arid paleoclimate. Dissolution of the chemically reactive disordered lattice of eolian-abraded quartz in subarkosic loess served as the predominant source of silica for massive amounts of biotic and abiotic chert in Osagean eolo-marine sediments. The ≤20 μm fraction of quartz dust is particularly susceptible to dissolution and re-precipitation as microcrystalline quartz (chert). We conclude that tectonic, eustatic, and climatic (allocyclic) processes all exerted some control on deposition; tectonics and eustasy controlled accommodation space, whereas paleoclimate changes (cycles), driven predominantly by orbital forcing, were the principal control on sediment supply and lithostratigraphy.

Redox conditions and ecological resilience during Oceanic Anoxic Event 2 in the Western Interior Seaway

Oceanic Anoxic Events (OAEs) are important geological events that may be analogues to future climate-driven deoxygenation of our oceans. Much of the global ocean experienced anoxic conditions during the Cenomanian–Turonian OAE (OAE2; ∼94 Ma), whereas the Western Interior Seaway (WIS) experienced oxygenation at this time. Here, organic geochemical and palynological data generated from Cenomanian–Turonian age sediments from five sites in the WIS are used to investigate changing redox and ecological conditions across differing palaeoenvironments and palaeolatitudes. Heterogeneity across the sites is apparent, but important relationships and trends among oceanographic variables are recognised: 1) Increasing total organic carbon (TOC) and CaCO3 percentages indicate the onset of a sea-level maximum towards the end of OAE2; 2) C28 sterane is shown to be a useful marker for prasinophyte abundance, and concurrent increases in this marker and overall sterane abundance indicate prasinophyte-driven increase in algal productivity in a stratified water column; and 3) sterane ratios can be a more reliable geochemical proxy than redox proxies for assessing the Benthic Oxic Zone. Our redox data do not always follow established trends for the WIS overall, particularly for proximal settings. We therefore surmise that local effects, such as nutrient-driven expansion of the oxygen minimum zone and/or sedimentation-driven anoxia just below the sediment-water interface, have overprinted regional trends.

The first small-bodied ornithopod dinosaur from the Lewisville Formation (middle Cenomanian) of Texas

ABSTRACT Sediments of the Woodbine Group exposed in northeastern Texas were deposited along the southwestern margin of Appalachia as a series of near-shore, shoreline, distal lowland swamp, lake, and fluvial deposits during a regression of the Western Interior Seaway in early and middle Cenomanian time. The Lewisville Formation (upper Woodbine Group) of north Texas preserves the most diverse terrestrial fossil assemblage known from Appalachia, but remains of small ornithischian dinosaurs have been conspicuously absent from it. An almost complete left dentary from the Lewisville Formation represents a new, small-bodied ornithopod taxon, Ampelognathus coheni gen. et sp. nov. The dentary is generally similar to those in non-iguanodontian ornithopods such as Hypsilophodon, Changchunsaurus, Haya, and Convolosaurus. Ampelognathus occupied an expected but previously missing component of the ‘mid’ Cretaceous terrestrial fauna of southwestern Appalachia. The growing diversity of fossil vertebrates and renewed paleobotanical study in the Lewisville Formation reinforces the importance of the unit’s fossil record for understanding eastern North American terrestrial ecosystems during an important transitional period in the earliest Late Cretaceous.http://zoobank.org/urn:lsid:zoobank.org:pub:02642649-ED6F-483A-994A-EC5F3FDF2AC7