It is not yet known if the properties of molecular gas in distant protocluster galaxies are significantly affected by their environment as galaxies are in local clusters. Through a deep, 64 h of effective on-source integration with the Australian Telescope Compact Array (ATCA), we discovered a massive,Mmol= 2.0 ± 0.2× 1011M⊙, extended, ~40 kpc, CO(1–0)-emitting disk in the protocluster surrounding the radio galaxy, MRC 1138−262. The galaxy, atzCO= 2.1478, is a clumpy, massive disk galaxy,M∗~ 5 × 1011M⊙, which lies 250 kpc in projection from MRC 1138−262 and is a known Hαemitter, named HAE229. This source has a molecular gas fraction of ~30%. The CO emission has a kinematic gradient along its major axis, centered on the highest surface brightness rest-frame optical emission, consistent with HAE229 being a rotating disk. Surprisingly, a significant fraction of the CO emission lies outside of the UV/optical emission. In spite of this, HAE229 follows the same relation between star-formation rate and molecular gas mass as normal field galaxies. HAE229 is the first CO(1–0) detection of an ordinary, star-forming galaxy in a protocluster. We compare a sample of cluster members atz> 0.4 thatare detected in low-order CO transitions, with a similar sample of sources drawn from the field. We confirm findings that the CO-luminosity and full-width at half maximum are correlated in starbursts and show that this relation is valid for normal high-zgalaxies as well as for those in overdensities. We do not find a clear dichotomy in the integrated Schmidt-Kennicutt relation for protocluster and field galaxies. Our results suggest that environment does not have an impact on the “star-formation efficiency” or the molecular gas content of high-redshift galaxies. Not finding any environmental dependence in these characteristics, especially for such an extended CO disk, suggests that environmentally-specific processes such as ram pressure stripping do not operate efficiently in (proto)clusters.