Empirically, it was established that during the performance of official duties, in particular, during practical classes (training), fire training instructors are in "zones" of significant psychophysical stress, which in the case of low (unsatisfactory) individual physical and special physical readiness can slow down, and in some cases even to level the process of formation of practical shooting skills from various types of small arms (automatic) weapons among military personnel of various categories who are studying. This, in turn, actualizes the need for a suitable professional development program for future fire training instructors, which, in addition to specific theoretical knowledge and practical skills (according to the military accounting specialty), provides for the use of special physical training. According to the results of the monitoring of specialized Internet resources (analysis of scientific-methodical and reference) literature, it was established that theoretical and empirical studies that comprehensively reveal the peculiarities of the organization of the training system of fire training instructors with an emphasis on the development of applied motor actions are rare. In the process of research and analytical work, the following methods were used (at the theoretical level): axiomatic, idealization, instrumental, historical and logical, descent from the concrete, formalization, as well as the experience of organizing the system of fire and special physical training of servicemen of various categories of institutions of the security and defense sector of Ukraine. As a result of the research, the members of the research group developed experimental organizational and pedagogical conditions for the formation of the readiness of future fire training instructors for professional activity with an emphasis on the use of special physical training tools. Prospects for further research in the chosen direction of scientific intelligence provide for the testing of the organizational and pedagogical conditions proposed by us (on the example of future instructors in fire training, representatives of military units of the National Guard of Ukraine).