SW Scotland Research Articles

Anomalous facies of the Kinnesswood Formation in the Cumbraes Basin: a late Famennian playa complex in SW Scotland

Among the important geological attributes of the small Cumbraes Basin in the southwestern off-shore Midland Valley of Scotland, the 4 km-long, fragmented but well-exposed section through the Devonian–Carboniferous boundary sequence in southern Great Cumbrae island ranks highly. In a continuous short stretch of the south and west coasts, large portions of this sequence are superbly preserved in tide-washed strandline surfaces and in old post-glacial sea cliff. This enhanced exposure leaves no doubt that the outcrops belong to two, immediately successive, members of the Kinnesswood Formation – the lower the Doughend Sandstone, the upper the Foul Port Mudstone – and that they provide some of the best available exposures on which to base description and interpretation. The facies of the fluviatile Doughend Sandstone shares many characteristics with the underlying Kelly Burn Sandstone of the Upper Old Red Sandstone, whereas that of the Foul Port Mudstone is distinct and unusual, attributed to local tectonics, influenced by movements on older and more major fractures such as the Highland Boundary Fault, with sediment accumulation taking place in a playa-lake complex of considerable extent.

Equilibrium partitioning and isotopic fractionation of nitrogen between biotite, plagioclase, and K-feldspar during magmatic differentiation

A significant portion of the continental crust is composed of plutonic igneous rocks. However, little is known about the geochemical behaviour of N between the different minerals during magmatic differentiation. To provide new constraints for the behaviour of N during crust formation, we have characterised the geochemistry of nitrogen (N) in the compositionally zoned calc-alkaline pluton at Loch Doon, SW Scotland. We present N concentration and N isotope values for whole-rock data alongside biotite, plagioclase and K-feldspar mineral separates and assess the degree to which these data preserve equilibrium partitioning during magmatic differentiation. We show that whole rock likely inherited its N contents and δ15N signatures from the initial source composition and that this signature is homogenous at a pluton scale. Whilst the whole-rock data are best explained as crust-derived N in the source, the degree of homogenisation across a pluton scale is inconsistent with empirical N diffusivities, ruling out syn-emplacement crustal assimilation as the source of N. Instead, our data suggest a crustal signature inherited from depth associated with the Iapetus subduction zone. At a mineral scale, we find that N preferentially partitions into the feldspars over mica in this system in the order K-feldspar > plagioclase ≈ biotite > quartz, with average mineral–mineral distribution coefficients of DN plagioclase-biotite = 1.3 ± 0.6 and DN Kspar-biotite = 2.8 ± 0.6. Partitioning is accompanied by a large and near constant equilibrium isotope fractionation factor between biotite and both feldspars (averages are Δ15NPlag-Biotite = +7.8 ± 1.2‰ and Δ15NKspar-Biotite = +7.9 ± 1.0‰). In contrast, Δ15NKspar-Plagioclase closely approximates 0‰, where both minerals show δ15N values overlapping with the bulk rock δ15N values. These results show that mica crystallisation generates a 15N-depleted reservoir within plutonic rocks. Moreover, our dataset suggests that feldspars might be a more significant host of N in the igneous portion of Earth’s continental and oceanic crust than previously thought.

Prevalence and distribution of lice on sheep and cattle farms in Great Britain

Lice are common production-limiting ectoparasites affecting livestock. Up-to-date data on their prevalence and spatial distribution on farms in Great Britain is important given that prevalence is believed to be increasing as a result of insecticide resistance. Here the prevalence of farms reporting lice, and factors associated with louse presence, were assessed using a retrospective questionnaire. For sheep and cattle farms, 16.1 % and 15.8 % reported lice on their livestock, respectively. Beef farms were more likely to report lice than dairy farms, with a prevalence of 18.0 % and 7.8 %, respectively. For sheep farms, prevalence was highest in Wales (27.7 %) and Scotland (22.4 %). For cattle farms, prevalence was highest in Scotland (27.6 %), Wales (18.5 %) and SW England (18.5 %). For sheep farms, statistical hotspot clusters were identified in Wales, NW England and SW Scotland, with prevalence in these areas ranging from 30.7 to 40.0%. For cattle farms clustering of cases was less evident. Multivariable analysis showed that significant factors associated with lice on sheep farms were larger flock sizes and geographic location (Scotland or Wales). For beef cattle farms, significant associated factors were larger herd sizes and upland grazing. More than 90 % of farms that reported lice, also reported treating for lice.

New records of crinoids from Trearne Quarry SSSI (Mississippian, Lower Carboniferous), north Ayrshire

The Mississippian succession of Ayrshire, SW Scotland, is rich in fossil crinoids, albeit mainly preserved as fragments. Trearne Quarry is exceptional in yielding moderately common crinoid cups and thecae from certain horizons. To the two nominal taxa that have been documented hitherto, we add a further seven, all cladids with one exception. These species all come from the Blackhall Limestone of the Lower Limestone Formation (Visean, Mississippian). Nominal crinoid species identified from Trearne Quarry include Cladida: Rhabdocrinus scotocarbonarius (Wright), Ureocrinus bockschii (Geinitz), Ureocrinus globulus (de Koninck), Tyrieocrinus laxus Wright, Fifeocrinus tielensis (Wright), Hydreionocrinus formosus Wright, Parazeacrinites konincki (Bather) and Phanocrinus ardrossensis (Wright); and Diplobathrida, Camerata: Cribanocrinus baccatus (Wright). Species have distributions limited to one or a few mudrock beds. Identifiable crinoids are uncommon in massive, coral-rich reefal facies.

Seismic and borehole-based mapping of the late Carboniferous succession in the Canonbie Coalfield, SW Scotland: evidence for a ‘broken’ Variscan foreland?

Local seismic and borehole-based mapping of the Carboniferous Pennine Coal Measures and Warwickshire Group successions in the Canonbie Coalfield (SW Scotland) provides evidence of repeated episodes of positive inversion, syn-depositional folding and unconformities. A Duckmantian (Westphalian B) episode of NE–SW transpression is recognized, based on onlapping seismic reflector geometries against NE-trending positive inversion structures and contemporaneous NNE-trending syn-depositional growth folding. The basin history thus revealed at Canonbie is at variance with generally accepted models in neighbouring northern England that imply subsidence was due to post-rift thermal subsidence during late Carboniferous times. A late Westphalian–Stephanian unconformity recognized within the Warwickshire Group succession signifies NW–SE, c . 10% local basin shortening during a time of major shortening in the late Carboniferous Variscan foreland, contradicting suggestions that maximum Variscan shortening had negligible impact on Carboniferous basins in northern Britain. Local inversion structures appear to have strongly influenced local late Westphalian–Stephanian depocentres. In this respect, the Variscan foreland at Canonbie may have resembled a ‘broken’ foreland system. Variations in crustal rheology, fault strength and orientation, and mid-crustal detachments are suggested to have played important roles in determining strain localization and the nature of Westphalian–Stephanian depocentres in the Canonbie Coalfield.

Geology and petroleum prospectivity of the Larne and Portpatrick basins, North Channel, offshore SW Scotland and Northern Ireland

The Larne and Portpatrick basins, located in the North Channel between SW Scotland and Northern Ireland, have been the target of a small programme of petroleum exploration activities since 1971. A total of five hydrocarbon exploration wells have been drilled within the two basins, although as of yet no commercial discoveries have been made. The presence of hydrocarbon shows alongside the discovery of two good-quality reservoir–seal couplets within Triassic and underlying Permian strata has encouraged exploration within the region. The focus of this study is to evaluate the geology and hydrocarbon prospectivity of the Portpatrick Basin and the offshore section of the Larne Basin. This is achieved through the use of seismic reflection data, and gravity and aeromagnetic data, alongside sedimentological, petrophysical and additional available datasets from both onshore and offshore wells, boreholes and previously published studies. The primary reservoir interval, the Lower–Middle Triassic Sherwood Sandstone Group ( c . 600–900 m gross thickness), is distributed across both basins and shows good to excellent porosity (10–25%) and permeability (10–1000 mD) within the Larne Basin. The Middle–Late Triassic Mercia Mudstone Group should provide an excellent top seal where present due to the presence of thick regionally extensive halite deposits, although differential erosion has removed this seal from the margins of the Larne and Portpatrick basins. The Carboniferous, which has been postulated to contain organic-rich source-rock horizons, as inferred from their presence in adjacent basins, has not yet been penetrated within the depocentre of either basin. There is, therefore, some degree of uncertainty regarding the quality and distribution of a potential source rock. The interpretation of seismic reflection profiles presented here, alongside the occurrence of hydrocarbon shows, indicates the presence of organic-rich pre-Permian sedimentary rocks within both basins. 1D petroleum system modelling of the Larne-2 borehole shows that the timing of hydrocarbon generation and migration within the basins is a significant risk, with many traps post-dating the primary hydrocarbon charge. Well-failure analysis has revealed that trap breach associated with kilometre-scale uplift events, and the drilling of wells off-structure due to a lack of good-quality subsurface data, have contributed to the lack of discoveries. While the Larne and Portpatrick basins have many elements required for a working petroleum system, along with supporting hydrocarbon shows, the high risks coupled with the small scale of potential discoveries makes the Portpatrick Basin and offshore section of the Larne Basin poorly prospective for oil and gas discovery. Thematic collection: This article is part of the Under-explored plays and frontier basins of the UK continental shelf collection available at: http://www.lyellcollection.org/cc/under-explored-plays-and-frontier-basins-of-the-uk-continental-shelf

Sporadic implementation of UK familial mammographic surveillance guidelines 15 years after original publication

The National Institute of health and Care Excellence issued guidelines on familial breast cancer screening in 2004. Such guidelines should be uniformly implemented to ensure that members of the same family with the same level of risk, but living in different areas, have the same access to screening. We assessed uptake by creating a short, six question online survey designed to assess compliance in each regional area. We used this to conduct a survey of all 22 regional genetics services. There was a 100% response to the survey allowing a complete map to be created. The devolved nations had near complete compliance with the sole exception of SW Scotland, but in England the picture was fragmented with regions representing a combined population of 26.6 million (48%) not implementing the full NICE recommendations. Fifteen years after the publication of the original guidelines, major inequity in provision for screening still occurs and a postcode lottery exists for the management of women from families with a history of breast cancer. We estimate that up to 73 preventable breast cancer deaths occur each year due to the current inequity of access. It may be time to consider alternative funding and implementation models to ensure consistent access across the country.

Soil compaction effects on grassland silage yields and soil structure under different levels of compaction over three years

Soil compaction has been estimated to be responsible for 33 million ha of soil degradation in Europe, reducing crop yields, however there is limited data on grassland silage yields loss. Extended grazing periods, increased size and weight of farm vehicles and more extreme weather have fostered concern over the consequences of grassland management on reduced grass yield and soil quality. This work aimed at studying the effect of increased animal trampling and mechanical (tractor) soil compaction on grassland silage mean dry matter (DM) yields and soil structure over a three year period at two UK sites. These sites were on two established perennial ryegrass fields with contrasting soil textures; an imperfectly drained silty clay loam in SW Scotland and a well drained sandy loam from central England. Results showed trampling and tractor compaction decreased mean DM yields over three years and by the third year DM yield for the trampled area was 11.4% less on the soil with greater clay content soil and 12.0% less on the more sandier soil than the no compaction control. DM yield for the tractor compaction, by the third year, was 14.5% less than no compaction DM yield, on both soil types. Compaction treatments gave the greatest reductions for the first silage cut DM yields annually, for both soil types. The largest reductions (19.0% for trampling and 37.7% for tractor) were on the soil with the greater clay content in the second year, with the coolest start to the growing season. Compaction reduced N uptake, decreased drainage and increased water filled pore spaces (WFPS). Linear regression of visual evaluation of soil structure (VESS) scores and bulk densities provided evidence that VESS is an effective tool for detecting grassland compaction and would assist with the management of moderately compacted soils where deteriorate soil conditions may result in yield loss.

NIRS quantification of lake sediment composition by multiple regression using end-member spectra

Here we develop a novel method for quantifying sediment components, e.g. biogenic silica, organic or mineral matter, from near infrared (NIR) spectra based on fitting by multiple regression of measured spectra for end-member materials. We show that with suitable end-members our new open-source multiple regression routine gives excellent simultaneous quantification of the major components of a sediment, the concentrations comparing well with independent methods of quantification. Widely used partial least squares regression approaches rely on large environmental training data sets; our method produces comparable results, but with the advantages of negating the need for a training dataset and with greater simplicity and theoretical robustness. We demonstrate that component NIR spectra are additive, a prerequisite for use of multiple regression to un-mix the compound spectra, and show that a number of environmental materials make suitable end-members for this analysis. We show that spectral mixing is not conservative with respect to mass proportion, but rather to the relative chromatic intensity of contributing sediment components. Concentrations can be calculated using the measured spectra by correction using a chromatic intensity factor, the value of which can be measured independently. We have applied our approach to a postglacial sediment sequence from Loch Grannoch (SW Scotland) and reveal a down-core pattern of varying dominance by biogenic silica, organic and mineral content from the late glacial to present. With isolation and measurement of appropriate end-members this multivariate regression approach to interrogating NIR spectra has utility across a wide range of sedimentary environments and potentially for other spectral analytical methods.

Burnswark Hill: the opening shot of the Antonine reconquest of Scotland?

This paper represents an interim report of the Burnswark Project that has utilised the techniques of battlefield archaeology to locate and accurately identify missiles across a substantial field of conflict in SW Scotland. The distribution, characteristics, dating and proposed historical context of these missiles are argued to support their use in a dramatic episode of ‘exemplary force’ by the Roman army. The magnitude of the event might be seen to be compatible with an emperor establishing his legitimacy as a military leader.

On the age of the Ballantrae Complex, SW Scotland

The Ballantrae Complex, SW Scotland, is an ophiolitic assemblage of mostly Early and early Middle Ordovician age (Tremadoc–Arenig in terms of the British Ordovician Series). Its varied components were generated and assembled in the Iapetus Ocean, then obducted on to the Laurentian continental margin by the earliest Llanvirn. The timing of obduction is constrained by biostratigraphic and radiometric data. It was most probably a polyphase process initiated at about the beginning of the Arenig, at around 478 Ma. However, parts of the Complex are significantly younger, with some recent evidence taken to suggest an earliest Llanvirn age of about 464 Ma for the emplacement of some of the volcanic and pelagic sedimentary rocks. The oldest strata in the succession that now unconformably overlies the Ballantrae Complex were deposited at about 463 Ma. Hence there may have been as little as one million years available for the final stages of the Complex's tectonic assembly, obduction, uplift, erosion and downfaulting. Obduction of the Complex has been invoked as a factor in the initiation of the Grampian Orogeny and, whilst there is a broad correlation in timing, the detail from Ballantrae militates against a causal relationship.

Nasty, Brutish and Short?; The Life Cycle of an Iron Age Roundhouse at Black Loch of Myrton, SW Scotland

ABSTRACTExcavations at Black Loch of Myrton, Dumfries & Galloway are revealing the very well-preserved remains of an Iron Age settlement, the wetland context ensuring that the timber structures have remained intact and that the detritus of daily occupation survives for us to pick apart and understand. One of the structures in this settlement is an exceptionally well-preserved roundhouse, the material remains of which have been subjected to a barrage of analyses encompassing the insect, macroplant, bone and wood assemblages, soil micromorphology, faecal steroids, radiocarbon-dating and dendrochronology. These will enable us to address some of the key issues regarding the life cycles of Iron Age roundhouses, from conception and construction, use of internal space, nature of occupation and likely function, through to abandonment. Critically, we are now able to view that life cycle through the lens of a tightly-defined chronology bringing us close to the ‘ … short-term timescales of lived reality’ [Foxhall, L. 2000. “The Running Sands of Time: Archaeology and the Short-Term.” World Archaeology 31 (3): 484–498].

Climate fluctuations recorded in phreatic and vadose calcretes of the Lower Carboniferous Clyde Sandstone Formation of Machrihanish, Kintyre Peninsula, SW Scotland

The Galdrings cliffs of Machrihanish, in the Kintyre Peninsula of Scotland, expose a Lower Carboniferous clastic succession that hosts a wide variety of calcretes, including thick and massive host-replacing phreatic calcrete hardpans (HRPCHs), which are geologically rare, and which are the products of a thorough replacement of host minerals by calcite in the mixing zone between fresh and evaporitic groundwaters. Isotopic values of 53 samples from various calcretes distributed in a c. 45 m thick succession provided well-defined trends of covariance between carbon and oxygen isotopes, thus delineating trends of aridification and humidification from the interplay of precipitation and evaporation rates. Although values from HRPCHs are more constrained, all other forms of calcrete in the succession (pedogenic nodules, pillar calcrete, laminar calcrete and invasive phreatic calcrete cement in sandstone) follow similar stable isotopic trends. An aridification trend preceded the formation of each HRPCH occurrence, corroborating studies on modern equivalents that suggest that they develop in the more arid range of calcrete formation. The better constrained isotopic values of HRPCHs compared with those of more common forms of calcrete are interpreted to be indicative of more specific environmental requirements, which may in part explain their rarity in the geological record.

Relative sea-level data from southwest Scotland constrain meltwater-driven sea-level jumps prior to the 8.2 kyr BP event

The most significant climate cooling of the Holocene is centred on 8.2 kyr BP (the ‘8.2 event’). Its cause is widely attributed to an abrupt slowdown of the Atlantic Meridional Overturning Circulation (AMOC) associated with the sudden drainage of Laurentide proglacial Lakes Agassiz and Ojibway, but model simulations have difficulty reproducing the event with a single-pulse scenario of freshwater input. Several lines of evidence point to multiple episodes of freshwater release from the decaying Laurentide Ice Sheet (LIS) between ∼8900 and ∼8200 cal yr BP, yet the precise number, timing and magnitude of these events – critical constraints for AMOC simulations – are far from resolved. Here we present a high-resolution relative sea level (RSL) record for the period 8800 to 7800 cal yr BP developed from estuarine and salt-marsh deposits in SW Scotland. We find that RSL rose abruptly in three steps by 0.35 m, 0.7 m and 0.4 m (mean) at 8760–8640, 8595–8465, 8323–8218 cal yr BP respectively. The timing of these RSL steps correlate closely with short-lived events expressed in North Atlantic proxy climate and oceanographic records, providing evidence of at least three distinct episodes of enhanced meltwater discharge from the decaying LIS prior to the 8.2 event. Our observations can be used to test the fidelity of both climate and ice-sheet models in simulating abrupt change during the early Holocene.

Strain estimation in 3D by fitting linear and planar data to the March model

The probability density function associated with the March model is derived and used in a maximum likelihood method to estimate the best fit distribution and 3D strain parameters for a given set of linear or planar data. Typically it is assumed that in the initial state (pre-strain) linear or planar data are uniformly distributed on the sphere which means the number of strain parameters estimated needs to be reduced so that the numerical technique succeeds. Essentially this requires that the data are rotated into a suitable reference frame prior to analysis. The method has been applied to a suitable example from the Dalradian of SW Scotland and results obtained are consistent with those from an independent method of strain analysis. Despite March theory having been incorporated deep into the fabric of geological strain analysis, its full potential as a simple direct 3D strain analytical tool has not been achieved. The method developed here may help remedy this situation.

Intra-Carboniferous deformation and unconformity at Gilnockie Bridge, SW Scotland, reinterpreted as the result of multiple channel-bank collapse

The unconformity and deformation in Pennsylvanian strata seen at Gilnockie Bridge have previously been taken as being illustrative of the erosion and break in sedimentation between the Pennine Lower Coal Measures and Stainmore formations on nearby syndepositional anticlinal crestal areas in the Canonbie Coalfield, SW Scotland. However, Gilnockie Bridge is located in a synclinal position, and re-examination of the exposures suggests that the complex deformation seen there is a localized synsedimentary feature, resulting from multiple phases of river channel-bank collapse. This bank failure was possibly associated with contemporary movement on the nearby Gilnockie Fault, but not necessarily related to the unconformity on the nearby anticlines.

Production and management of cattle manure in the UK and implications for land application practice

AbstractA review of livestock and manure management across the UK was undertaken as part of a wider study of important factors impacting on greenhouse and other gaseous emissions from agriculture. All major livestock sectors were covered by the review, but only data on cattle manure are reported in this study. Cattle manure comprises 67 Mt (80%) of the total 83 Mt of UK annual livestock manure production during the housing period. Overall, about 47% of total UK cattle manure production was estimated to be as undiluted slurry and 53% as solid, mainly straw‐based farmyard manure (FYM). Survey data suggested important regional differences in cattle manure management across the UK; examples include the predominance of slurry in SW Scotland, N Ireland and to a lesser extent, in Wales, compared to the continued importance of straw‐based FYM systems in England. Survey data on land application of manures from the British Survey of Fertiliser Practice (BSFP) generally showed the expected distribution of cattle manure application timings, dominated by crop sowing/establishment or growth patterns. However, in recent years, an apparent increase in land spreading in February–April, with less application in November–January is encouraging, the trend towards spring/early summer spreading of cattle slurry suggesting greater appreciation of its value as a nutrient source. BSFP returns on manure application techniques have also shown increasing uptake of shallow injection of cattle slurry, from <1% (2004) to 10% (2013) and for increasing rapid (<6 h) incorporation of FYM, from 3% (2007) to 14% (2013).

Rapid fluid flow along fractures at greenschist facies conditions on Syros, Greece

This thesis aims to provide a better understanding of the role of mountain building in the carbon cycle. The amount of CO2 released into the atmosphere due to metamorphic processes is largely unknown. To constrain the quantity of CO2 released, fluid-driven reactions in metamorphic rocks can be studied by tracking fluid-rock interactions along ancient fluid flow pathways. The thesis is divided into two parts: 1) modeling of fluid flow rates and durations within shear zones and fractures during greenschist- and blueschist-facies metamorphism and 2) the assessment of possible mechanisms of fluid infiltration into rocks during greenschist- to epidote-amphibolite-facies metamorphism and controlling chemical and mineralogical factors of reaction front propagation.On the island Syros, Greece, fluid-rock interaction was examined along a shear zone and within brittle fractures to calculate fluid flux rates, flow velocities and durations. Petrological, geochemical and thermodynamic evidence show that the flux of CO2-bearing fluids along the shear zone was 100-2000 times larger than the fluid flux in the surrounding rocks. The time-averaged fluid flow velocity and flow duration along brittle fractures was calculated by using a governing equation for one-dimensional transport (advection and diffusion) and field-based parameterization. This study shows that fluid flow along fractures on Syros was rapid and short lived.Mechanisms and controlling factors of fluid infiltration were studied in greenschist- to epidote-amphibolite-facies metabasalts in SW Scotland. Fluid infiltration into metabasaltic sills was unassisted by deformation and occurred along grain boundaries of hydrous minerals (e.g. amphibole) while other minerals (e.g. quartz) prevent fluid infiltration. Petrological, mineralogical and chemical studies of the sills show that the availability of reactant minerals and mechanical factors, e.g. volume change in epidote, are primary controls of reaction front propagation.

Regional variation in the biogeochemical and physical characteristics of natural peatland pools

Natural open-water pools are a common feature of northern peatlands and are known to be an important source of atmospheric methane (CH4). Pool environmental variables, particularly water chemistry, vegetation community and physical characteristics, have the potential to exert strong controls on carbon cycling in pools. A total of 66 peatland pools were studied across three regions of the UK (northern Scotland, south-west Scotland, and Northern Ireland). We found that within-region variability of pool water chemistry was low; however, for many pool variables measured there were significant differences between regions. PCA analysis showed that pools in SW Scotland were strongly associated with greater vegetative cover and shallower water depth which is likely to increase dissolved organic carbon (DOC) mineralisation rates, whereas pools in N Scotland were more open and deeper. Pool water DOC, particulate organic carbon and dissolved CH4 concentrations were significantly different between regions. Pools in Northern Ireland had the highest concentrations of DOC (mean=14.5mgL−1) and CH4 (mean=20.6μgCL−1). Chloride and sulphate concentrations were significantly higher in the pools in N Scotland (mean values 26.3 and 2.40mgL−1, respectively) than elsewhere, due to a stronger marine influence. The ratio of UV absorbance at 465nm to absorbance at 665nm for pools in Northern Ireland indicated that DOC was sourced from poorly humified peat, potentially increasing the bioavailability and mineralisation of organic carbon in pools compared to the pools elsewhere. This study, which specifically aims to address a lack of basic biogeochemical knowledge about pool water chemistry, clearly shows that peatland pools are highly regionally variable. This is likely to be a reflection of significant regional-scale differences in peatland C cycling.

A collection of pelmatozoans (Echinodermata) from the Lady Burn Starfish Beds (Upper Ordovician, Katian), SW Scotland

A new collection of crinoids and rhombiferans from the Upper Ordovician (Katian) Lady Burn Starfish Beds of SW Scotland is small, yet contains several significant specimens. Two species hitherto known only from single specimens, the dendrocrinid cladid Cupulocrinus drummuckensis Kolata and the pisocrinid disparid Eocicerocrinus sevastopuloi Donovan, are represented by crowns retaining the proximal column. The latter species is the earliest member of an order that is otherwise known only from the Silurian and Devonian. Unusual preservation of the monobathrid camerate Macrostylocrinus cirrifer Ramsbottom, in which the radices (roots) of the proximal column are lost, but the delicate pinnules of the arms are preserved, is problematic. A long, contorted and gracile pluricolumnal is probably derived from a disparid, possibly E. sevastopuloi .