Normal seafloor entering the Puerto Rico and northern Lesser Antillean trenches in the northeastern Caribbean is interrupted by a series of aseismic ridges on the North and South American plates. These topographic features lie close to the expected trend of fracture zones created about 80–110 m.y. ago when this seafloor was formed at the Mid‐Atlantic Ridge. The northernmost of the ridges that interact with the Lesser Antillean subduction zone, the Barracuda Ridge, intersects the arc in a region of high seismic activity. Some of this seismicity including a large shock in 1974, occurs within the overthrust plate and may be related to the deformation of the Caribbean plate as it overrides the ridge. A large bathymetric high, the Main Ridge, is oriented obliquely to the Puerto Rico trench and intersects the subduction zone north of the Virgin Islands in another cluster of seismic activity along the inner wall of the trench. Data from a seismic network in the northeastern Caribbean indicate that this intersection is also characterized by both interpolate and intraplate seismic activity. Magnetic anomalies, bathymetric trends, and the pattern of deformed sediments on the inner wall of the trench strongly suggest that the Main and Barracuda ridges are parts of a formerly continuous aseismic ridge, a segment of which has recently been overridden by the Caribbean plate. Reconstruction of mid‐Miocene to Recent plate motions also suggest that at least two aseismic ridges, and possibly fragments of the Bahama Platform, have interacted with the subduction zone in the northeastern Caribbean. The introduction of these narrow segments of anomalous seafloor into the subduction zone has segmented the arc into elements about 200 km long. These ridges may act as tectonic barriers or asperities during the rupture processes involved in large earthquakes. They also leave a geologic imprint on segments of the arc with which they have interacted. A 50‐km landward jump of the locus of island arc volcanism occurred in Late Miocene time along the northern half of the Lesser Antilles. We postulate that the subduction of a segment of seafloor of anomolously thick crust, being more buoyant than adjacent seafloor, resulted in a marked shoaling in the dip of the descending slab and, therefore, a shift of the locus of volcanism. In the region near western Puerto Rico and eastern Hispanolia, Plio‐Pleistocene interaction with a similar feature, in this case a part of the Bahama Platform, about 3–4 m.y. ago led to a jump in the locus of subduction as evidenced by a gap in the downgoing seismic zone. That segment of the Bahama Platform interferred with the subduction process and was subsequently sutured onto the Caribbean plate when the boundary jumped about 60 km to the northeast. The maximum size of historic shallow earthquakes along the Lesser Antillean arc varies from about 7.0–7.5 in the center of the arc where the dip of the shallow part of the plate boundary is steep to 8.0–8.5 along the northern part of the arc where the dip is shallow. The interaction of anomalous seafloor, as along the northern Lesser Antilles, can lead to the development of a wider than normal zone of interplate contact and hence to earthquakes that are larger than those associated with more typical seafloor entering subduction zones. Major seismic gaps and regions of high seismic potential currently exist along the northern Lesser Antilles and to the north of Puerto Rico. Both gaps are bounded by anomalous features on the downgoing plate. The intersection of these features with the plate boundary created large asperities that may be good places to search for precursors to future large earthquakes. A great shock in 1787 may have ruptured an existing seismic gap north of Puerto Rico between 65° and 67°W. Thus that gap can be expected to eventually rupture again in a great shock and not to accommodate plate motion by totally aseismic processes.
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