Pat's research has addressed an impressively broad range of questions emanating from her core interest in understanding the causes and consequences of life-history trade-offs and resulting life-history variation. She has provided ground-breaking and much needed insights into key physiological and genetic mechanisms that underlie life-history variation, and is unusual in the degree to which she has retained a clear-sighted view and commitment to injecting her research into applied ecology and conservation policy. Although Pat's studies have involved insects, amphibians, fish and mammals, birds have proved an ideal model for addressing her questions of interest, and so ornithology is her natural home. After graduating in Zoology at the University of Glasgow in 1973, Pat moved to Durham undertake a PhD on Herring Gull Larus argentatus ecology, supervised by John Coulson (awarded the Goldman Salvin Prize in 1992). She then returned to a Lectureship in Glasgow, and through the rest of her career has played a pivotal role in the development of ornithological science in Glasgow, the UK and beyond. Pat's early research continued to focus on seabird ecology, with a developing emphasis on interdisciplinary studies into issues such as disease transmission by wintering gulls. These studies continued through the 1980s, including research on emerging conservation issues that showed that seabird declines were being driven by reductions in the local availability of their sandeel prey. Those studies in Shetland illustrate a theme that has underpinned much of Pat's subsequent work, where she has successfully demonstrated fitness consequences of the varied responses that different animals have to environmental change. Building upon those field studies, in which environmental variation provided her with natural experiments, Pat identified a series of systems where experimental manipulations could be used on wild populations. This led to investigations into the constraints on maternal investment, and how maternal condition influenced egg quality, sex ratios and offspring fitness. Her interest in the longer-term consequences of variation in early condition led Pat to develop increasingly elegant experiments with laboratory species such as Zebra Finches Taeniopygia guttata. Collaborating with a diverse group of modellers, physiologists and molecular biologists, these laboratory studies have revealed how early growth conditions, relating both to nutrition and social environments, can influence a range of biological characteristics in later life. Alongside this work on captive species, she has successfully identified opportunities to explore these questions in wild populations, collaborating with colleagues such as previous Goldman Salvin medallists Mike Harris and Sarah Wanless, and further demonstrating the value of the long-term individual-based studies of seabirds on the Isle of May. Together, these studies highlight how Pat has used ornithology to address fundamental questions of broad biological interest. Yet at the same time, she retains an active research interest in avian conservation, particularly in the way that she has driven the science underpinning the Scottish Chough Study Group's major impact on current conservation strategies for this species. Pat's research has resulted in over 200 scientific papers and contributions to many books and popular science articles. Her work is highly cited, and has had an important influence across ornithology and other disciplines. Arguably, however, her greatest legacy is the enormous impact she has made as a research leader through her absolute commitment to scientific rigour and excellence, and her unquestionable integrity and commitment to fairness in society. She has contributed these attributes through her supervision and mentoring of students and early-career scientists, and her contributions to numerous bodies and panels, including the Royal Society and the Leverhulme Trust. Your career will have been directly or indirectly influenced by Pat, whether you know it or not. Pat has supervised over 40 PhD students, almost all of whom have carved successful scientific careers in roles that range from independent academics to senior conservation managers. Pat contributes to all the key programmes that now promote women in science. More critically, she has been an exceptional role model and mentor for young female scientists since the very beginning of her career, culminating in her becoming the first woman to be appointed to Glasgow's Regius Chair in Zoology. Directly and indirectly, she has undoubtedly contributed to the high numbers of women now working in ornithology within the UK. Pat's professional services have also involved extensive work within government and learned societies, where her achievements have been recognized through her appointment as President of both the International Society for Behavioural Ecology and the Association for the Study of Animal Behaviour. These activities have resulted in a number of awards, notably her election as a Fellow to the Royal Society of Edinburgh in 1997, and the Association for the Study of Animal Behaviour Scientific Medal in 2015. The strength and breadth of her work within ornithology now makes her an outstandingly deserving recipient of the BOU Goldman Salvin Prize.