Steppe Eagle Research Articles

Evaluation of habitat suitability and migratory paths of an endangered raptor, Steppe Eagle (Aquila nipalensis) in Iran

Understanding habitat suitability, the environmental variables limiting the distribution of species, and migratory paths are important issues for conservation of threatened bird species. Identifying areas important for birds and their overlap with conservation areas (CAs) can guide conservation managers in establishing new CAs. The Steppe Eagle (Aquila nipalensis) is a globally endangered winter visitor raptor in Iran. We used 164 occurrence records of Steppe Eagles and data on 12 environmental variables in Iran as input to ensemble modeling and electrical circuit theory models to identify, respectively, potential wintering areas and migratory paths between those wintering areas. Our results revealed that elevation, distance to rodents, mean diurnal range, distance to villages, and distance to cities were the most influential variables for habitat suitability in Iran. Potential wintering areas identified by our models were mainly located in the north and south of Iran and migratory paths connected these areas through the central plains. CAs covered about one-fifth of potential wintering areas. Conservation of the species within potential wintering areas and the migratory paths from northern to southern Iran is necessary for the survival of this endangered species in its entire distribution. Therefore, wildlife managers should pay increased attention to non-protected parts of potential wintering areas in order to establish new CAs and protect migration paths against threats. Our results pave the way for proper planning for the conservation of threatened raptors in Iran, particularly Steppe Eagle.

Birds of Shivapuri Nagarjun National Park, Nepal

Shivapuri Nagarjun National Park harbors diverse avifauna. The last checklist of birds for Shivapuri, published by SNP and BCN (2007), covered 318 species. However, it excluded the Nagarjun Forest birds as this area wasn't incorporated into the national park until 2009. Since then, many new bird sightings have been reported in Shivapuri Nagarjun National Park (SNNP). To address this gap, we've compiled a comprehensive checklist of SNNP birds in Nepal's central mid-hills, drawing on direct observations and a literature review. A total of 396 species of birds belonging to 18 orders and 71 families were known to occur in the park. Order Passeriformes (n=260, 65%) and family Muscicapidae (n=50, 12%) have the highest record of the number of species. Out of 396 bird species, 237 species were recorded during the recent survey of 2021–2022 and 159 species were considered from the past literatures. 10.60% (n=42) of the recorded bird species are classified as nationally threatened according to Nepal’s national red list and 2% (n=8) are listed as globally threatened. The globally threatened birds include Vulnerable greater spotted eagle Aquila clanga, imperial eagle Aquila heliaca, and Kashmir flycatcher Ficedula subrubra; Endangered steppe eagle Aquila nipalensis and Pallas’s fish eagle Haliaeetus leucoryphus; and Critically Endangered white-rumped vulture Gyps bengalensis, slender-billed vulture Gyps tenuirostris and red-headed vulture Sarcogyps calvus. A total of 55 species recorded are included in the CITES Appendices; one species is also listed in the National Parks and Wildlife Conservation Act (NPWCA) Schedule I, and the park includes spiny babbler Acanthoptila nipalensis the only endemic bird of Nepal. Resident species make up the majority (n=243, 61.36%) of birds recorded in the national park. The remaining are the migrants which include summer and winter visitors, passage migrants and vagrants.

Assessing the susceptibility of raptor species to electrocution: A framework for Kenya

Developing countries are witnessing rapid expansion of electrical infrastructure to meet increasing energy demands, prompting a critical need to assess the potential impact on avian biodiversity. Leveraging open access data, our study conducted a comprehensive assessment to detect electrocution and collision risk hotspots in Kenya while identifying raptor species highly susceptible to these risks. Through an integrated approach that considers morphological and behavioral traits of the species, environmental factors at the site, and technical parameters of the power lines, we developed risk maps and categorized raptors’ susceptibility into high, medium, and low-risk levels. Applying this framework, we identified three raptor species at high risk of electrocution; the long-crested eagle, the augur buzzard, and the steppe eagle. Meru and Kiambu counties emerged as high-risk areas due to substantial overlap between high-risk buffer zones and areas with high raptor species distribution. It is worth noting that this framework only accounts for thirteen raptor species, and there is scope to expand it in the future to include other avian species, such as storks, bustards, pelicans and flamingos, which are also susceptible to electrocution and collision incidents and measures to mitigate electrocution of raptors may not be sufficient for these other group of birds.

Composition and Conservation Status of Avifauna in Urban Non-protected Important Bird Area (IBA) Site of Western India

Kumbharwada wetland, an Important Bird Area (IBA) component in Bhavnagar, Gujarat, India, is a crucial wintering ground for migratory birds. Long-term monitoring of the avian community over a period of long time-frame is an excellent way to examine the health of this IBA site and thus provide an important ground to foster the conservation of birds in the region and management of this wetland. Given this consideration, field surveys were carried out from December 2020 to May 2023, following point count method to study the avian species richness in the study area. A total of 204 bird species belonging to 20 orders and 56 families are recorded, of which 85 species are migratory and 119 are resident. Anatidae is the most species-rich avian family (16 species). Highest number of species was recorded in the month of January (165 species in 2021 and 163 species in 2022). This wetland supports 107 (52.45%) wetland-associated species and 97 (47.55%) terrestrial species of birds. Twelve species are considered as Near Threatened, four species as Vulnerable and one species (Aquila nipalensis) as Endangered in IUCN Red List of Threatened Species. Four species (Sterna aurantia, Mycteria leucocephala, Phoeniconaias minor, and Threskiornis melanocephalus) with globally declining trend, are commonly seen in the study area, which shows that the wetland is a crucial habitat for bird species with high conservation priorities. Industrialization, encroachment, discharge of sewage water and chemical effluents, high-tension powerlines, stray dogs, and expansion of exotic vegetation remained the major threats to the habitat and avian community.

Characterising ecological resource utilization by the endangered Egyptian vulture in Oman to better manage and mitigate electrocution risk

Electrocution is a threat to birds and can undermine their conservation status. We tracked 15 globally endangered Egyptian vultures (Neophron percnopterus) in Oman and used Resource Selection Functions (RSFs) to identify habitats important to vultures and locate places where electrocution risk is high. During daytime vultures selected rugged areas with low vegetative cover, and habitats near roads, waste disposal sites, and both high (>50 kV, large pylons) and medium voltage electricity infrastructure. At night vultures also selected rugged habitats with scarce vegetation. At night high voltage infrastructure was selected, especially in areas of relatively low ruggedness at higher elevations; medium voltage infrastructure was avoided at night. The pattern of infrastructure use suggested that vultures were using large pylons for night-time roosting, but used both large pylons and smaller poles for perching during the day. 54.2 % of 11–33 kV powerlines in the study area were within the top six habitat suitability categories, which also contained 87.3 % of 34,778 vulture daytime test locations. Retrofitting powerlines overlapping the highest suitability zones would have the largest immediate impact. Areas surrounding landfills were also important: 10.34 % of all test locations were located within 2 km of landfills, which contained only 14.5 km of 11–33 kV lines. Targeting these areas would provide high impact at low cost. These maps are particularly useful in Oman because it is a stronghold for resident endangered large birds (i.e., Egyptian vulture, lappet-faced vulture Torgos tracheliotos) and an important winter destination for endangered steppe eagles (Aquila nipalensis).

Habitat-dependent avian diversity in the agri-industrial zone of Jhajjar, Haryana, India

The study was performed to evaluate the habitat-dependent avian diversity in the southern region of Haryana from March 2021 to March 2023. The study aimed to identify avian species diversity in three habitats (agrifields, human settlement, and industrial zone) along with diversity indices and relative diversity. A total of 5037 birds belonging to 17 orders, 41 families, and 88 species were investigated with Simpson’s diversity index of 0.973 and Shannon– Wiener index of 4.04. Analysis of threat status revealed 38 species with stable population trends, 19 increasing, 20 decreasing, and 11 species having unknown trends. Out of 88 species, the IUCN status of two species Aquila nipalensis and Psittacula eupatria was endangered and near threatened, respectively; the rest 86 belonged to the least concern category. Of the total, 81% of species were residential, 10 were winter migratory, and 7 were summer migratory. Among the three habitats, agrifields were the most diverse, followed by human settlement and industrial zone. Diversified assemblages of birds suggested the region as a suitable habitat for both resident and migratory birds. This study highlights the need for more regular investigations to assess yearly avian diversity and possible threats to design comprehensive conservation plans.

Bird Diversity in Bhaludhunga: Exploring Avian Diversity in Eastern Mid-hills of Nepal

Birds are crucial to biodiversity and serve as an indicator of habitat quality, productivity and stability. This study documents the avian diversity in Bhaludhunga Community Forest and associated tea garden in Ilam, which is a part of the Mai Valley Forests Important Bird and Biodiversity Area. It addresses the limited research on avi-faunal diversity in eastern Nepal, particularly outside protected areas. MacKinnon’s Listing method was used to collect the data following the existing trail transect walk in the study area. During the study period, a total of 132 bird species from 41 families of 10 orders were recorded. Order Passeriformes were the most dominant order with 102 species followed by Accipitriformes and Columbiformes. Muscicapidae was the most commonly represented family with 19 species followed by Phylloscopidae, and Pellorneidae. One globally endangered species, Steppe Eagle along with five nationally threatened species was recorded during the survey. Two Restricted Range Species from the Eastern Himalayas Endemic Bird Area Spiny Babbler and Yellow-vented Warbler were also recorded in the area. However, extensive survey in the different seasons is required for further exploration of bird community, which might play a crucial role in developing baseline information and implementing conservation implications.

System Identification of Unmanned Air Systems at Texas A&M University

This paper presents a summary of system identification flight testing and results for a variety of large and small fixed-wing and multirotor unmanned air systems at Texas A&M University from 1999 to 2023. The six different types of vehicles range from a large powered parafoil, to a fixed-wing vehicle with synthetic jet-actuated roll control effectors, to a radially asymmetric multirotor, to large and small fixed-wing vehicles, and to a Steppe eagle. The observer/Kalman filter identification algorithm is used to generate linear time-invariant state-space models, and the results for both near-real-time online model generation and postflight offline model generation are presented. The use and efficacy of a variety of test input types and their sensitivity to exogenous inputs such as turbulence, in addition to identified model evaluation and selection criteria, are discussed. Several generations of low size, weight, power, and cost flight-test instrumentation including the Developmental Flight-Test Instrumentation data acquisition package are also presented. Challenges that arose from the flight-testing campaigns along with solutions are highlighted in the paper.

Mapping suitable habitats for globally endangered raptors in Kenya: Integrating climate factors and conservation planning.

Raptors face global threats like electrocution, collisions, and habitat fragmentation. Many species remain understudied, and their distribution patterns are unknown. Understanding their current and future distribution is crucial for conservation. Protecting these top predators requires knowledge of their spatial distribution and environmental influences. This study addresses knowledge gaps in raptor habitats and distributions in Kenya, considering current and future climate changes. Using species distribution models and occurrence data from the Global Biodiversity Information Facility, we evaluated suitable habitats for four endangered Kenyan raptor species: Martial eagle, Secretarybird, Bateleur, and Steppe Eagle. We assessed the impact of climatic predictors on their distribution, considering two climate change scenarios for 2020-2040. Our findings reveal that raptor distribution in Kenya is predominantly concentrated in the southwestern region, extending into the central region of the country. The most significant predictors of raptor species distribution varied for each species, with Steppe eagle and Secretarybird being highly influenced by precipitation during the warmest quarter, Martial eagle being influenced by mean temperature during the driest quarter, and Bateleur being primarily influenced by precipitation during the coldest quarter. When projecting our model into the climate change scenarios for 2020-2040, all species except the Bateleur exhibited a negative range shift. The results of our study suggest that climate change may have adverse impacts on the raptor species examined. In light of these findings, we recommend implementing targeted monitoring and conducting surveys in accordance with our current model predictions. Specifically, our focus should be on monitoring areas that exhibit the highest climate suitability, as these areas are likely to undergo significant shifts in the near future. By conducting regular monitoring and engaging in further research, we can enhance our understanding of these raptor species and gather valuable data to improve the accuracy and reliability of our model predictions.

Степной орёл в Алтае-Саянском регионе – результаты исследований 2019–2023 годов

Степной орёл в Алтае-Саянском регионе – результаты исследований 2019–2023 годов

Популяционно-генетическая структура степного орла: есть ли надежда на сохранение вида?

For centuries, Steppe Eagle (Aquila nipalensis) had been the most numerous Palearctic Aquila species, but nowadays it has been decreasing in numbers progressively, so in 2015 the IUCN changed the species status to “endangered”. The decrease is evidently related to Steppe Eagles’ electrocution at nesting areas and during migration, poisoning by veterinarian drugs when feeding on agricultural animal carcasses at wintering grounds, steppe fires and total habitat loss combined with loss of prey like susliks. To the contrary, the sister Aquila species, the Imperial Eagle (Aquila heliaca), which exhibits similar biological traits and lives simpatrically to the Steppe Eagle at some parts of areas, shows local increase in numbers and even substitutes the decreasing Steppe Eagle in steppe biomes. In the 1960s, many Palearctic birds of prey came through population decline due to so-called “DDT crisis” – massive pesticide poisoning. To restore the species numbers, specific conservation actions were initiated in Europe to support population numbers and genetic diversity of raptors. Regarding the Steppe Eagle with its nesting area located mostly at Kazakhstan, Russian, Mongolian and Chinese territories, the implications of DDT and electrocution impacts on the population numbers stayed poorly investigated. Meanwhile, loss of genetic stability can be a reason for lasting population decline. Genetic threats like population fragmentation, genetic erosion, and inbreeding depression are believed more prolonged in comparison to anthropogenic impacts and more rarely come under increasing scrutiny of conservation practitioners. However, if exist, they can lead to species extinction regardless of removal of all other threats. The Steppe Eagle population structure stays poorly investigated. In 2018, during the 1st Steppe Eagle conservation international workshop, this problem was highlighted with recommendations for filling this gap. Our presented study was conducted following this recommendation and focused on genetic structure, fragmentation, and effective number comparison of the Steppe Eagle population at the individual samples) and sympatric populations of the Imperial Eagle (over 100 samples). For conservation criteria estimation, we used widespread genetic markers like nuclear microsatellites (9 loci) and mitochondrial control region (D-loop). Also we performed molecular individual tagging and parentage analysis to study the population structure using samples from the numerous Western Kazakhstan population. We also used GIS methods to study geographic fragmentation of populations. The genetic analysis showed that the Steppe Eagle population structure is highly similar to that of the Imperial Eagle but with less genetic diversity and has traces of repeated survival of “population bottlenecks” with huge decline in the number and genetic diversity. Fragmentation analysis showed no genetic isolation of geographically fragmented nesting groups despite the presumed natal phylopatry in accordance with our Steppe Eagle parentage data. Preliminary estimation of the Steppe Eagle population effective numbers by linkage disequilibrium method showed rather high values inspite of the lasting population decline. The molecular coancestry estimation showed more serious decline during the latest bottleneck which was also far below the ancestral Imperial Eagle population at the studied areas. The inbreeding rate for the Steppe Eagle population at the most part of the species areas turned out to outnumber one for the local population of the Imperial Eagle. Our data showed no immediate genetic threats for the Steppe Eagle population restoration and also demonstrated traces of multiple bottlenecks in the species evolution. Still the genetic structure of the species population keeps some traits of modern decline and should be monitored permanently according to the species numbers. Unique population structure patterns of the Steppe Eagle between other Aquilinae species and their conservation aspects are directions for future Aquila nipalensis genetic research.

О гнездовании степного орла в Дагестане

О гнездовании степного орла в Дагестане

Гнездовые группировки орлов на ключевой орнитологической территории «Чинк Донызтау» в 2022 году

Гнездовые группировки орлов на ключевой орнитологической территории «Чинк Донызтау» в 2022 году

Степной орел на территории резервата «Бокейорда» и Ащиозекского заказника (Западно-Казахстанская область) в 2022–2023 годы

Степной орел на территории резервата «Бокейорда» и Ащиозекского заказника (Западно-Казахстанская область) в 2022–2023 годы

Опыт использования гнездовых платформ для степного орла на юге Костанайской области, Казахстан

Steppe Eagle (Aquila nipalensis) population currently continues to decline in Kazakhstan, the south of the Kostanay region is no exception. The reasons for the decline are of different nature, both abiotic and biotic, and include the following: destruction of nests by land predators (most often Steppe Eagle nests are located on the ground); steppe fires are of great threat, and the fact that some nests are located on trees and bushes does not help, but rather aggravates the situation; destruction of nests by livestock; developing new areas for farms, etc., as a result, Steppe Eagles are forced to leave favorable nesting territories; electrocution on power poles as Steppe Eagles use traverses to build nests. Artificial structures, such as nesting platforms, are widely used to attract raptors. During the implementation of the project “Steppe Eagle Conservation in Kazakhstan” within the international campaign “Flyway Champions”, 35 platforms were installed in favorable Steppe Eagle nesting biotopes in “Saga” and “Altybay” hunting grounds (managed by ACBK – Association for the Conservation of Biodiversity of Kazakhstan) in the South of the Kostanay region in the summer of 2021. The design is made entirely of metal, consists of a support with a cross section of 100×100 mm and a height of 3 m, where 1 m is buried in the ground, and a “basket” with dimensions of 1×1 m installed at the top. Various shrub and tree branches are placed and attached to the “basket”, designed to imitate an active nest. This design is protected from terrestrial predators, and most importantly, from steppe fires. In the breeding season of 2022, Steppe Eagles occupied three platforms, in 2023 – two. It is noteworthy that a pair of eagles used the same nest in both seasons. As an experiment, in 2021 an old nest of Long-Legged Buzzard (Buteo rufinus) was moved to one of the platforms, and in 2023 it was occupied. In 2022, one of three pairs left the nest for unknown reasons after completing it; in the remaining two, a clutch of two eggs was observed; later we observed down feathers, but only one nestling remained in the nest by the time of the departure. In 2023, clutches on two platforms consisted of two eggs each, but later two and one nestling were observed there, having left the nest in the same number. Nesting platforms also serve as a “perch” for a number of birds, and after the use by Steppe Eagles, Indian Sparrow (Passer indicus) occupies them; the number of such a colony reached 13–16 nests. It is also worth noting that sometimes the distance between occupied nesting platform and occupied natural nest is less than 2 km, which suggests that there is a need for this biotechnical activity.

Electrocution of Eagles in Mongolia

Steppe Eagles (Aquila nipalensis) and Golden Eagles (Aquila chrysaetos) are species of conservation and cultural importance in Mongolia. Electrocution at electricity distribution lines is a threat faced by both species across Central Asia. We present the results of power line surveys conducted in the Mongolian steppe region to elucidate temporal patterns of electrocution and specific danger points on power poles for eagles. There was spatial and temporal variation in electrocution rates for Steppe Eagles and Golden Eagles. Autumn surveys across Mongolia recorded the electrocuted carcasses of 18 Steppe Eagles and 9 Golden Eagles. The distribution of electrocutions revealed that Steppe Eagles are mainly killed at power lines in the central steppe zone; in contrast, only Golden Eagles were found at power lines in the southern ‘gobi’ desert zone. Daily surveys conducted for a year at a single power line in the eastern steppe detected 8 Golden Eagles and 6 Steppe Eagles. All Steppe Eagle electrocutions occurred between April and September as they are predominantly summer visitors in Mongolia, whereas six of the eight Golden Eagle electrocutions occurred in winter indicating that Golden Eagles can range over predominantly flat, open steppe landscapes at this time. Significantly more electrocutions occurred at poles with no crossarm mitigation compared to poles with deflectors, deterrents or covers, indicating that mitigation on the crossarm can reduce eagle electrocution rates. Furthermore, the absence of eagle carcasses at poles with crossarm mitigation suggests that crossarms are the main site of electrocution for eagles in Mongolia. Country-scale retrofitting of insulation to crossarms and pole tops significantly reduced eagle electrocution events.

Steppe Eagle breeding population surveys in Turkey between 2015 and 2022

The status of the Steppe Eagle (Aquila nipalensis) was unclear in Turkey, which forms the south-western border of the species’ breeding distribution. The status of the species in Turkey has never been clear, as historically only two breeding territories were found and documented in the wider area of Tuz Gölü, which were proved to be active only in some years between 1969 and 1980. One nest in Tuz Gölü was found in some years between 1969 and 1975, while another nest was found in Bolluk Gölü in 1975, and was active also in 1980. There were no published breeding records since 1980 until 2003, when a Steppe Eagle brood was found accidentally in central Anatolia by Hungarian birdwatchers. There were no other published breeding records of Steppe Eagles in Turkey in the twentieth century, although adult birds have been observed in a few occasions in Central and Eastern Anatolia during the breeding season between 1998 and 2015. A short species-specific survey was undertaken by Hungarian and Russian researchers in 2015, which revealed eight potential breeding territories in Kırşehir and Konya provinces, three of them with incubating birds, one with a pair and an active nest, and four further potential territories where mature individuals were observed. Based on these results further surveys have been organized in cooperation of Hungarian and Turkish researchers between 2017 and 2022. Alltogether 38 nests were found of which active breeding attempts were proved in 23 cases. During the surveys 23 potential territories have been identified: 10 territories with detected breeding attempts (incubation or nestling rearing), 5 territories with detected occupancy (territorial pairs or active nests), 6 potential territories (single adult birdsobserved in suitable habitats), and 2 abandoned territories (no activity was observed). The controlled clutches contained 2.00 eggs (n=6), 1.65 nestlings (n=18), and 1.37 fledglings (n=16) on average. Hybridization of Steppe Eagles with Eastern Imperial Eagles (Aquila heliaca) was proved genetically in one case, where a hybrid male bird paired with an imperial eagle female and successfully raised hybrid nestlings in several years. The dispersal and migration of six young steppe eagles and two hybrid juveniles have been tracked with the help of GPS transmitters. The tracked steppe and hybrid eagles moved to the Sahel region for wintering. The surveys also revealed significant potential threats to eagle populations within Turkey, including electrocution, shooting, massive habitat alteration, and infrastructural developments. Therefore, intensive further studies, including active species and habitat conservation measures, are inevitable for the maintenance of the Anatolian Steppe Eagle population, which is most probably in critically endangered status.

Species action plan for conservation of an isolated Steppe Eagle population in Turkey

Steppe Eagle (Aquila nipalensis) is a globally Endangered (EN) species according to IUCN Red List, with a breeding range extending through the arid zones of southern Russia and Kazakhstan, from Kalmykia in the west to Dauria in the east. There is a small, isolated breeding population in Turkey, discovered in 2003, which lost contact with the main breeding range. Although there are studies carried out between 2015 and 2021, the size of the isolated population is not exactly known. To protect this small population on the brink of extinction, the General Directorate of Nature Conservation and National Parks (GDNCNP), which is officially responsible for wildlife conservation, programmed to prepare a species action plan in 2021. As part of the “Preparation, Implementation and Monitoring of Species Action Plans for Endangered Species in Turkey within the Concept of a New Methodology”, shortly “Endangered Species” project, Steppe Eagle was selected, based on a prioritization study, to be one of the pilot species for which species actions to be prepared. Field studies were conducted in the 2022 breeding season to identify threats causing stress to the Steppe Eagle. During the breeding studies in 2022, it was determined that 4 pairs of Steppe Eagles were confirmed to be breeding in Central Anatolia, and 4 pairs attempted to breed. During the field studies, threat factors were listed based on direct observations, indirect observations (nest records, mortality records, etc.), and bilateral discussions with local communities. Using the threat analysis adopted by IUCN and adapted to Turkey within the framework of the same project, a national-level threat analysis was conducted. Direct threat factors causing stress to the species were listed as conversion of grasslands into agricultural areas, excessive use of groundwater and surface water leading to the drying of wetlands, power transmission lines, human activities and grazing pressure on breeding islands in wetlands, illegal hunting, predation, researcher pressure, hybridization with Eastern Imperial Eagles (Aquila heliaca), and artificial materials in nests. Additionally, the use of rodent poisons was also identified as a potential threat. As a result of the threat analysis, the threats with the highest level of impact on the species were identified as the conversion of grasslands into agricultural areas and excessive water usage. These threats result in the transformation of steppe habitat and decrease in the species' food sources. Another high-level threat is illegal hunting. The identified threats were transferred to a problem tree and, following the method generated in the “Endangered Species” project, problem tree was transformed to a result chain. In the result chain, the selected results for eliminating/reducing threats were defined by identifying intervention points and transforming them into objectives. Activities were defined to achieve the identified objectives. Within the scope of the 10-year action plan a total of 34 activities were designed under 9 objectives to be achieved. Eight out of the 9 objectives involve conservation, and one involves research-related activities. The aim of the 10-year plan is to conserve and support the breeding population of the species in Turkey.

Современный статус степного орла в Казахстане

Современный статус степного орла в Казахстане

Мониторинг гнездовой группировки степного орла в Актюбинской области (Казахстан) в 2018–2023 годах

Мониторинг гнездовой группировки степного орла в Актюбинской области (Казахстан) в 2018–2023 годах