Spruce Stands Research Articles

Lower alpha, higher beta, and similar gamma diversity of saproxylic beetles in unmanaged compared to managed Norway spruce stands.

Strong anthropogenic pressures on global forests necessitate that managed forests be evaluated as habitat for biodiversity. The complex pattern of habitat types created in forestry systems is ideal for analyses through the theoretical framework of alpha (local), gamma (total) and beta (compositional) diversity. Here I use saproxylic beetles, a species-rich threatened group, to compare four Norway spruce-dominated habitats representative of the boreal forest landscape of northern Europe: unmanaged semi-natural stands, nature reserves, unthinned middle-aged production stands and commercially thinned production stands. The beetles (in total 38 085 individuals of 312 species), including red-listed ones and three feeding guilds (wood consumers, fungivores and predators) were studied in 53 stands in central-southern Sweden, in two regions with differing amounts of conservation forest. Alpha diversity of saproxylic, but not red-listed, beetles was higher in the thinned stands than in the semi-natural stands, and did not differ for the other forest types. Beta diversity of saproxylic beetles was higher in unmanaged semi-natural stands than in the other forest types, but species composition did not differ noticeably. Furthermore, red-listed saproxylic beetles had higher gamma diversity in unmanaged semi-natural stands in the region with more conservation forest, but not in the one with less such forest. The local factors dead wood volume and dead wood diversity did not influence alpha diversity of beetles, but increasing canopy openness had a minor negative influence on saproxylic and red-listed beetles. While the local scale (alpha diversity) indicates the potential for managed forests to house many saproxylic beetle species associated with spruce forests in this boreal landscape, the larger scales (beta and gamma diversity) indicate the value of unmanaged forests for the conservation of the entire saproxylic beetle fauna. These results show the importance of analyses at multiple levels of diversity (alpha, beta, gamma) for identifying patterns relevant to conservation.

ТАКСАЦИОННЫЕ ПОКАЗАТЕЛИ И ТОВАРНАЯ СТРУКТУРА СПЕЛЫХ СМЕШАННЫХ ДРЕВОСТОЕВ СОСНЫ И ЕЛИ В ЗЕЛЕНОМОШНЫХ ТИПАХ ЛЕСА

The influence of the stand composition on taxation indicators and the stock of mature pine and spruce stands have been considered. The objects of study are forest stands with different proportions of pine and spruce in the most represented green moss forests of the landscape of the Tikhvin Ridge (Boksitogorsky forestry) of the Leningrad region. Stock assessment and determination of taxation indicators were carried out on trial plots laid out according to standard forestry methods. Commodity structure of the stand and different yield of assortments by size categories depend on the share of participation of one or another species in the composition of the plantation by the age of clear-cutting in the taiga zone in different types of forests. The purpose of the study was to determine optimal plantation composition in terms of stock and large-scale marketability by the age of maturity of the stand. The most productive in terms of stock and output of large-scale coniferous wood under the studied growing conditions are pine forests with a stand composition of 8C2E, 7C1E2B, 5C4E1B and spruce forests with a composition of 7E1C2B, 4E2C4Os+B. The obtained results of the study show that with an increase in the share of pine in the plantations of the green moss group of forest types, the yield of large-sized coniferous wood assortments increases by the age of a ripe forest stand

Differential Effects of Tree Species on Soil Microbiota 45 Years after Afforestation of Former Pastures

Several decades ago, many former pastures in Central Europe were afforested or colonized by trees after being abandoned. Knowledge of the effects of tree species on soil properties is important for planning of the composition of future forests. In this regard, a research location in Vrchdobroč (Central Slovakia), which is former agricultural land used as pasture, enables the exploration of ecosystem processes and properties in stands of different tree species after afforestation. The goal of our study was to find out whether changes in soil properties, including soil microbial activity and diversity among different stands, were already observable 45 years after the afforestation, and how the effects differed among stands of different tree species. The study was conducted in the pure stands of Norway spruce (Picea abies L. Karst.), Douglas fir (Pseudotsuga menziesi (Mirb.) Franco), European beech (Fagus sylvatica L.) and sycamore maple (Acer pseudoplatanus L.). Multivariate analyses of physico-chemical soil properties indicated an overlap between the soils under the Douglas fir and the spruce, but a clear separation of beech from sycamore. In general, both microbial activity and diversity were, surprisingly, highest under the Douglas fir, followed by the sycamore, with the beech and the spruce showing mostly lower values.

The impact of forest naturalness and tree species composition on soil organic carbon content in areas with unnatural occurrence of Norway spruce in the Czech Republic

Climate change has increased attention paid in the research to forest soils and tree species composition, in respect to the potential for carbon sequestration. It is known that forest stands are able to store soil organic carbon (SOC), but little is known about the effect of forest naturalness on SOC content. This is important in relation of dying of unnatural spruce stands. It is necessary to determine a suitable composition of tree species which will replace them. This research is based on 248 plots with oak, beech, and spruce stands and mixtures of these species, with measured values of SOC. Our results show that autochthonous and mixed stands, in terms of tree species composition, in the study area had a higher SOC content than allochthonous and pure stands. In addition, it was found that autochthonous oak and beech stands, especially in mixtures, had a higher SOC content than allochthonous spruce stands (monocultures). On the basis of the presented results, it is possible to optimize the future tree species composition of stands in the study area, which currently have an allochthonous representation of spruce, to provide better function of carbon sequestration and resistance to climate change.

Pre-commercial thinning in Norway spruce-birch mixed stands can provide abundant forage for ungulates without losing volume production

Mixed stands of Norway spruce and birch have the potential to simultaneously produce timber and provide large ungulates with a significant amount of forage during the regeneration phase. While the growth and yield of such mixtures are well studied, little is known about potential trade-offs between timber and forage production and which management techniques are suitable for meeting both goals. In this study, four different pre-commercial thinning (PCT) strategies were used to study the trade-offs between production and available forage for free-ranging ungulates in a Norway spruce-birch mixture. The four PCT strategies were: 1) retaining 2000 birch stems ha−1 with 2000 Norway spruce ha−1, 2) removing all birches within a 0.75 m radius around Norway spruce stems, 3) removing all birches and other broadleaves, and 4) no PCT (control). Growth of Norway spruce was higher in the 2000 birch ha−1 and full removal treatments compared to the untreated control, but these two treatments did not differ from one another in volume production of Norway spruce. We found a negative effect of PCT on forage availability but no effect on ungulate browsing. Therefore, PCT strategies that provide both sufficient birch forage and maximize volume production of Norway spruce can be implemented.

Pre-commercial thinning could mitigate drought stress of black spruce stands

Rising temperatures are likely to increase the risk of drought across the globe over the next century. Boreal forests are particularly vulnerable to drought because temperatures within these biomes are projected to warm the fastest. Warm and dry conditions can reduce tree growth, particularly in regions that are already moisture-limited, which may reduce forest productivity. Forest stand density management, such as pre-commercial thinning (PCT), can reduce moisture stress for residual trees by reducing canopy evaporation and increasing soil water availability. PCT is applied 15–20 years after clearcut and removes smaller stems to increase resource availability to residual trees, thereby increasing diameter growth and wood value. How PCT can mitigate climate change impact in boreal forests is still unclear, partly due to heterogeneity in moisture availability and growth response. This study tests the relative effect of PCT on radial growth of black spruce (Picea mariana (Mill.) B.S.P.) on sites with varying water availability to inform forest managers about PCT’s potential to mitigate anticipated effects of drought on tree growth. Tree discs from PCT and non-thinned plots were harvested from three contrasting site types in eastern Canada (warm-dry, warm-wet and cool wet). Using dendrochronology, the relationship between annual ring width index (RWI) and standardized-evapotranspiration index (SPEI) was explored in the time since PCT and during known climate anomalies. RWI showed a positive growth response to increasing SPEI on warm-dry sites, but a negative growth response on warm-wet and cool-wet sites. PCT provided a greater benefit to radial growth on warm-dry site types in the 15 years since treatment but provided no additional benefits during years of climate anomalies. Results suggest that PCT will remain an important forest management practice on moisture-limited sites in order to maintain black spruce productivity, whereas tree growth may benefit from future warming on relatively wet sites.

Forest Structure and Composition Diverge Following Harvesting Compared to a Spruce Budworm Choristoneura fumiferana (Clem.) Outbreak

Balsam fir (Abies balsamea) is the most vulnerable species to the spruce budworm (Choristoneura fumiferana), one of the most devastating defoliators in the world. For decades, pest managers have advocated for reducing its abundance in the landscape to minimize losses to the spruce budworm (SBW). Although reduction of fir occurred during the endemic phase of the SBW cycle, there is little information about the extent to which this general principle (reduction of fir) was applied during an outbreak and whether it occurs at both stand and landscape levels. The objective of this paper is to compare the effect of insect and harvest disturbances on forest structure during the 1970–80s outbreak in Québec. We evaluate whether, (i) forest management activities targeted fir forests and whether patch size of host species influences management or SBW disturbance, (ii) SBW outbreaks and logging have similar or divergent effects on forest composition. Although data are from an earlier outbreak, they are at a scale rarely studied and will be useful in guiding decisions made at larger scales in the current and future outbreaks. Our results show that spruce was targeted preferentially by harvesting (up to 69% of plots) during the outbreak period, while it represented less than one third of plots defoliated by the SBW. On the other hand, fir stands represented up to 75% of plots that were defoliated by the SBW but less than 35% of plots that underwent harvesting. Harvesting targeted large blocks of spruce forest more than large blocks of fir-dominated forest while the opposite was observed for the SBW. In terms of regeneration, SBW tends to reduce fir and favor spruce recruitment, along with non-host species, whereas the opposite tendency was observed following harvesting. In terms of spatial organization of stands, our results support the suggestion that small stands of fir and large stands of spruce undergo the least SBW damage. Thus, in order to attenuate SBW impacts in the future, efforts should be made to ensure that spruce recruitment is favored and that its abundance increases at both the stand and landscape scale.

Environmental Effects among Differently Located and Fertile Sites on Forest Basal-Area Increment in Temperate Zone

Environmental properties differently influence the growth of forest tree species. The antagonistic effects of variable environmental properties classify the forest response according to various tree compositions among different sites. The division of the forest response was assessed in 52 stands arranged into 26 types of 13 site management populations (MPs) in 5 areas in the Czech Republic territory. The assessment was performed using time-series multiple regressions of basal-area increment from pure immature stands of Norway spruce (Picea abies), Scots pine (Pinus sylvestris), oaks (Quercus sp.), ash (Fraxinus excelsior) and willows (Salix sp.) dependent on the interpolated average temperatures, annual precipitation, atmospheric concentrations of SO2, NOx and O3 and soil properties over the period 1971–2008 at p < 0.05. Site MPs differentiated the forest response to a greater extent than tree species. The response of the forests was significantly distributed by means of the montane, upland and waterlogged sites. The multiple determination index (r2) ≥ 0.6 indicated an adaptable tree increment but an interval of r2 between 0.80–0.92 implied forest sensitivity to variability in environmental properties on non-waterlogged sites. The index r2 < 0.6 suggested a fluctuating forest increment that reflects environmental variability inconsistently. The fluctuating increment most affected the spruce and pine stands grown from upland to submontane locations. Montane spruce stands, as well as rock pines, appeared to be one of the most sensitive ones to environmental change. Floodplain forests seemed as adaptable to variable environmental properties.

Evaluation of a Restoration Approach After One Century – Effects of Admixed European Beech on the Natural Regeneration Potential and Humus Condition in Spruce Stands

About 100 years ago, groups of European beech were reintroduced into Norway spruce stands in some parts of Germany as a restoration approach. The aim of this approach was to maintain or rehabilitate the ecological function of soil fertility and the regeneration option of beech for the next generation. The effect of beech groups on surrounding spruce stands was studied in the Ore Mountains by recording natural regeneration of beech and humus layer thickness and form. Point pattern statistics were used to analyse the spread of beech regeneration and to determine factors influencing its establishment. It was found that the density of regeneration decreases with increasing distance from the beech group. However, beech regeneration was found up to distances of 69 m. Furthermore, it becomes evident that PAR radiation (maximum regeneration densities at PAR values of 35 W/m2) and fencing (3.41 times higher regeneration density compared to unfenced areas) against deer have a positive influence on beech regeneration density. Ordered categorical models were used to model humus form and non-linear models were used to model humus layer thickness. It could be proven that the most bioactive humus forms and lowest humus layer thicknesses were found within the beech group. With increasing distance to the beech group, the total humus layer thickness and the proportion of mormoder in the spruce stand increased. The positive influence of the beech group on the humus composition extends to about 40 m from the centre of the beech group. Due to the former arrangement of the beech groups in the terrain, the effects can also extend to the spruce stands in between. The hypotheses on the restoration approach of reintroducing groups of beech into spruce stands formulated by foresters 100 years ago can thus be confirmed. For future restoration approaches of spruce stands, groupwise mixtures of beech should be established with a distance of 40–50 m.

Overstory dynamics regulate the spatial variability in forest-floor CO2 fluxes across a managed boreal forest landscape

The forest-floor represents an important interface for various carbon dioxide (CO2) fluxes, however, our knowledge of their variability and drivers across a managed boreal forest landscape is limited. Here, we used a three-year (2016−2018) data set of biometric- and chamber-based flux measurements to investigate the net forest-floor CO2 exchange (NEff) and its component fluxes across 50 forest stands spanning different soil types, tree species, and age classes within a 68 km2 boreal catchment in Sweden. We found that the forest-floor acted as a net CO2 source with the 10th–90th percentile (used hereafter for describing reported variations) ranging from 149 to 399 g C m−2 yr−1. Among the key landscape attributes, stand age strongly affected most NEff component fluxes, whereas tree species and soil type effects were weak and absent, respectively. Specifically, forest-floor net CO2 emissions increased with stand age due to declining understory gross and net primary production, ranging between 77–275 and 49–163 g C m−2 yr−1, respectively. Furthermore, we observed higher understory production rates in pine than in spruce stands. Across the 50 stands, the total forest-floor respiration ranged between 340 and 549 g C m−2 yr−1 and its spatial variation was primarily regulated by its autotrophic components, i.e., understory and tree root respiration, which displayed divergent increasing and decreasing age-related trends, respectively. Furthermore, heterotrophic soil respiration remained within a relatively narrow range (154–290 g C m−2 yr−1), possibly owing to compensating gradients in forest-floor properties. We further identified tree biomass as the major driver of the landscape-scale variations of CO2 fluxes, likely attributable to modulating effects on forest-floor resource availability and growing conditions. This implies that tree growth responses to forest management and global change will be particularly important for regulating magnitudes and spatial variations of forest-floor CO2 fluxes in boreal forests.

Forest Management under Climate Change: A Decision Analysis of Thinning Interventions for Water Services and Biomass in a Norway Spruce Stand in South Germany

Climate change is producing threats to forests’ capacity of regulating water regimes. Therefore, thinning strategies can be applied to mitigate climate change impacts more efficiently by providing more spaces for trees to utilize resources e.g., water and nutrients. This study examined the effects of different thinning intensities and intervals on water characteristics and biomass growth of a 75-year-old Norway spruce (Picea abies) stand in the Black Forest, Germany. Here we used a water and management sensitive update of the process-based forest growth model 3PG, 3PG-Hydro. We applied light (10%), moderate (30%), and heavy thinning (50% intensity) in the interval of 10, 25, and 50 years of the management period. We simulated growth with climate change scenario RCP 8.5 data from 1995 to 2065. We analyzed the effects of the different thinning regimens on biomass, evapotranspiration as well as water yield. Thinning intensity and interval as well as their interaction have significant influence on production of stand biomass and water yield for all thinning regimes applied (p < 0.05). However, there is no significant difference (p > 0.05) in accumulated biomass (thinned biomass added to the stand biomass) between the applied thinning regimes. Light thinning in a long interval (50 years) produced highest stand biomass among the applied thinning regimes. Furthermore, the prediction showed that accumulated water yield increased with increasing thinning intensity. Our study concludes that repeated moderate thinning at intermediate intervals results in a high water yield without losing biomass production.

The effect of stand characteristics on soil organic carbon content in spruce and deciduous stands

Aim of study: To determine the effects of stand characteristics, which closely relate to forest management practices, on the soil organic carbon (SOC) content in the organic (O) and surface mineral (A) soil horizons in spruce and deciduous stands, and to show SOC dynamics during the life of production stands. Area of study: Spruce and deciduous stands located throughout the Czech Republic. Material and methods: The effects of age, density of stocking and canopy of stand on SOC content in the O and A horizons, and the difference between categories of variables and the trends of SOC were evaluated in spruce and deciduous stands (401 plots) at lower and middle elevations. Main results: SOC content changed during the life of stands. In spruce stands, a decreasing trend of SOC with stand age was found in the A horizon. In deciduous stands, SOC content was higher overall in the A horizon, fluctuating slightly with stand age, but more balanced during the life of stands. Based on the results, in terms of management of dying spruce stands and carbon sequestration, felling should be carried out in the age group of 81-120 years in spruce stands, whereas in deciduous stands felling should take place in older stands (141 years and more). Density of stocking and canopy of stand had no substantial effect of SOC content. Research highlights: Deciduous stands have the potential to replace dying spruce stands at lower elevation in terms of carbon sequestration.

Heterotrophic and rhizospheric respiration in coniferous forest soils along a latitudinal gradient

Northern forest soils are a major carbon (C) reservoir of global importance. To estimate how the C balance in these soils will change, the roles of tree roots and soil microbes in C balance should first be decoupled. This study determined how the activity of heterotrophs and tree roots together with root-associated microbes in the rhizosphere varies in coniferous forest soils in boreal, hemiboreal, and temperate climates along a latitudinal gradient using a trenching approach. We created experimental plots without living tree roots, measured soil respiration (CO2 efflux) from these and from unmanipulated plots using the chamber technique, and partitioned the efflux into root-rhizosphere (RR) and heterotrophic (RH) respiration. The share of RR in ecosystem gross primary production (GPP) decreased from north to south in the Scots pine (Pinus sylvestris L.) and the Norway spruce (Picea abies (L.) Karst.) forests, with the exception of a mixed site, where the share of RR in GPP varied strongly between the years. RR per ground area and per root biomass were mainly independent of climate within the gradient. RH per ground area increased from north to south with temperature, while RH per soil C did not change with temperature. Soil moisture did not significantly affect the respiration components in the northernmost site, whereas soil moisture was positively connected with RH and negatively with RR in other Scots pine sites and positively connected with RR in pure Norway spruce stands. The dynamic ecosystem model LPJ-GUESS was able to capture the seasonal dynamics of RH and RR at the sites, but overall accuracy varied markedly between the sites, as the model underestimated RH in the southern site and RR elsewhere. Our study provides knowledge about the nature of soil respiration components. The valuable insights can be used in more accurate land-ecosystem modelling of forest ecosystems.

Uncrewed aircraft system spherical photography for the vertical characterization of canopy structural traits.

Summary The plant area index (PAI) is a structural trait that succinctly parametrizes the foliage distribution of a canopy and is usually estimated using indirect optical techniques such as digital hemispherical photography. Critically, on‐the‐ground photographic measurements forgo the vertical variation of canopy structure which regulates the local light environment. Hence new approaches are sought for vertical sampling of traits.We present an uncrewed aircraft system (UAS) spherical photographic method to obtain structural traits throughout the depth of tree canopies. Our method explained 89% of the variation in PAI when compared with ground‐based hemispherical photography.When comparing UAS vertical trait profiles with airborne laser scanning data, we found highest agreement in an open birch (Betula pendula/pubescens) canopy. Minor disagreement was found in dense spruce (Picea abies) stands, especially in the lower canopy.Our new method enables easy estimation of the vertical dimension of canopy structural traits in previously inaccessible spaces. The method is affordable and safe and therefore readily usable by plant scientists.

Fungi Occurring in Norway Spruce Wood Decayed by Heterobasidion parviporum in Puszcza Borecka Stands (Northeastern Poland)

In many spruce stands, trees are frequently attacked by the pathogen Heterobasidion parviporum, albeit without visible symptoms in the crown. In the present work, the results of the presence of stem rot, assessed by PICUS Sonic Tomography, and the fungal biota on trees and stumps in eight plots in the Puszcza Borecka Forest are described. The plots were located in stands on original forest soil (4) and on post-agricultural soil (4), where around a stump with H. parviporum symptoms (signs of internal rot and basidiocarps), 30 trees were selected and examined for internal rot. Wood samples were collected from two selected trees for fungal molecular analysis. A total of 79 fungal taxa were found, including 57 taxa in plots on post-agricultural soil and 45 on forest soil. There were 395 fungal records on stumps and 22 records on trees, therein, from the inner parts of felled trunks. Significant differences in the Chao-1 diversity index indicate that the origin of the soil—post-agricultural or forest soil—influenced the alpha diversity of the fungal communities in the forests studied. The values of the Shannon and Simpson indices show that the two communities were similar in terms of species numbers. The presence of basidiomata of H. parviporum and two species of Armillaria (mainly A. cepistipes) in samples on all plots is striking, although Armillaria spp. was detected more frequently. Most of the species identified were typical saprotrophs, although rare species were also found, such as Entoloma byssisedum, Onnia tomentosa, Physisporinus vitreus, Postia ptychogaster, and Ramaria apiculata. The presence of H. parviporum in the inner woody parts was confirmed by PCR analysis, and decay was detected even up to a stem height of 6 m. Armillaria was the dominant genus in the studied stands and plays a significant and underestimated role in heartwood decay of old spruce trees in Puszcza Borecka Forest.

Analysis of the association Galio rotundifolii-Abietetum albae Wraber (1955) 1959 on Manjača Mt (NW Bosnia and Herzegovina)

This research aimed to study phytocoenological characteristics of a fir-dominated community on silicate substrate at the locality of Sitnica on Manjača Mt, northwestern Bosnia and Herzegovina. This community originated from artificially forested mixed stands of fir and spruce planted in the habitat of the mountain Dinaric fir-beech forest, over a hundred years ago. With regular selective cuttings and natural regeneration, the structure shifted from even-aged to uneven-aged. In total, 18 phytocoenological relevés were made using the Braun-Blanquet method during the vegetation seasons of 2021 and 2022. Relevés were stored in Turboveg database. Ecological factors were calculated based on Ellenberg bioindicator values. Species composition was analyzed in terms of life form, geoelements and chorotypes. Our analysis indicates that collected relevés can be classified as the association Galio rotundifolii- Abietetum albae belonging to the alliance Fagion sylvaticae.

A First Assessment of Canopy Cover Loss in Germany’s Forests after the 2018–2020 Drought Years

Central Europe was hit by several unusually strong periods of drought and heat between 2018 and 2020. These droughts affected forest ecosystems. Cascading effects with bark beetle infestations in spruce stands were fatal to vast forest areas in Germany. We present the first assessment of canopy cover loss in Germany for the period of January 2018–April 2021. Our approach makes use of dense Sentinel-2 and Landsat-8 time-series data. We computed the disturbance index (DI) from the tasseled cap components brightness, greenness, and wetness. Using quantiles, we generated monthly DI composites and calculated anomalies in a reference period (2017). From the resulting map, we calculated the canopy cover loss statistics for administrative entities. Our results show a canopy cover loss of 501,000 ha for Germany, with large regional differences. The losses were largest in central Germany and reached up to two-thirds of coniferous forest loss in some districts. Our map has high spatial (10 m) and temporal (monthly) resolution and can be updated at any time.

Phlebiopsis gigantea strains from Estonia show potential as native biocontrol agents against Heterobasidion root rot and contain diverse dsRNA and ssRNA viruses

Coniferous forests, especially Norway spruce (Picea abies L.) stands, are threatened by root rot disease caused by Heterobasidion annosum sensu lato (Fr.) Bref. The basidiomycetous fungus, Phlebiopsis gigantea (Fr.) Donk, is used as a biocontrol agent against this pathogen in several countries. In Estonia, stump treatments are currently conducted using the Rotstop preparation, which consists of a P. gigantea strain isolated in Finland. However, for ecological reasons it would be preferable to use an indigenous biocontrol strain. In this study, we tested new strains of P. gigantea isolated from Norway spruce stumps in Estonia for their growth rate and competitive ability against H. parviporum and H. annosum, and identified P. gigantea strains promising for biocontrol purposes. As viruses may cause phenotypic debilitation in fungi, we analyzed the virome of these strains by high-throughput sequencing (RNA-Seq) and assessed whether the presence of mycoviruses affects their performance as compared to virus-free strains of P. gigantea. Viruses related to members of Partitiviridae and Curvulaviridae, as well as unclassified ambi-like, aspi-like and “fusariviruses” were identified for the first time in P. gigantea.

Using hybrid modelling to predict basal area and evaluate effects of climate change on growth of Norway spruce and Scots pine stands

ABSTRACT When modelling forest growth, capturing the effects of climate change is needed for reliable long-term predictions and management choices. This remains a challenge because commonly used mensurational forest growth and yield models, relying on inventory data, cannot account for climate change effects. We developed hybrid physiological/mensurational basal area growth and yield models, which combine physiological response to climatic conditions and empirical relations. We included climate and site effects by replacing time with light sums of photosynthetically active radiation and modifying the latter with monthly soil water, vapour pressure deficit, temperature, and frost days. When parameterised with permanent sample plot data for Scots pine and Norway spruce across Sweden, the hybrid models could reproduce observations well, although with no increase in precision compared with time-based mensurational models. When considering different climate scenarios, a significant impact on productivity from climate change emerged. For example, a 2 °C warming enhanced Scots pine production by up to 14% in regions where temperatures were originally cooler and soil water deficit was low (i.e. northwest Sweden), but depressed it, up to 9%, elsewhere. Hence, climate-sensitive models that take local variations into account are necessary for accurate predictions and sustainable forest management.

Diversity of click beetles in managed nonnative coniferous and native beech stands: Consequences of changes in the structural and species composition of tree stands in Central Europe

The natural composition of forests has undergone significant changes over recent centuries. A closer-to-natural tree species composition has long been perceived as key to a high biodiversity. We investigated the impact on communities of click beetles (Elateridae) caused by changes in the tree species composition of spruce monocultures compared to reference sites of recently unmanaged natural beech forests. To collect data, passive interception traps were distributed within managed spruce stands of different age classes and natural beech forests of various developmental stages. The beetle species richness was slightly but not significantly higher in the beech forests. The saproxylic species group was significantly more common in the spruce stands, whereas the group of nonsaproxylic species was significantly more abundant in the beech stands. In the commercial stands, the significantly highest species richness was in the clearings (0–10-year-old stands), and at this forest age class, the vast majority of the beetle species occurred in the spruce stands. In the developmental stages of the natural forest, a slightly higher beetle richness was found at the disintegration stage. The study results suggested that different tree species compositions and stand structures affect the communities of click beetles and substantially change their species composition and thus their response to external influences. Therefore, management of stands using diverse silvicultural systems is recommended for creating diverse ecological niches in forests.