Ammonite Species Research Articles

Oxyarietites boletzkyin. gen., n. sp., nouveau genre et nouvelle espèce d'ammonite dans le Sinémurien inférieur de Bourgogne (France): un rare précurseur des morphologies oxycônes pour le Jurassique

Oxyarietites boletzkyi n. gen., n. sp., a new genus and species of ammonite for the Lower Sinemurian of Burgundy (France): a rare forerunner of the oxycone morphologies for the Jurassic.One of the very first quasi-oxycone ammonites following the Triassic/Jurassic boundary crisis is described. It was collected from the fossiliferous Lower Sinemurian (Semicostatum or Turneri Chronozone) strata of Burgundy (Mavilly-Mandelot, Côte-d'Or, France). The new taxon, Oxyarietites boletzkyi n. gen., n. sp., possesses an involute, compressed and keeled shell of suboxycone morphology, a shell type previously unknown for the Lower Sinemurian. The discovery makes younger by about 2 Ma the emergence of keeled (sub)oxycone shells following the Triassic/Jurassic boundary crisis. Its obviously keeled ventral area allows a probable assignation to the Arietitoidea Hyatt, 1875 sensu Guex (1995) superfamily, but family level assignation and its evolutionary origin remain obscure. Although, it is generally accepted that involute, compressed and keeled suboxycone and oxycone ammonite shells possess the best hydrodynamical abilities and mobility, the acquisition of this probable adaptive advantage in O. boletzkyi n. gen., ç. sp. does not go together with abundancy in the fossil record.

Early Cretaceous ammonite fauna of Catanduanes Island, Philippines

Nine Aptian to early Albian ammonoid species, apparently from the Yop Formation, in Catanduanes Island, Philippines, are described herein: Goretophylloceras sp., Eogaudryceras (Eotetragonites) sp., Notosilesitoides philippinensis gen. et sp. nov., Miyakoceras tanohatense, Neosilesites nepos, Neoastieria? sp., Marshallites miyakoensis, Colombiceras cf. satowi and Colombiceras sp. This is the first systematic palaeontological report of Cretaceous ammonoids from the Philippines. Among the described species, Notosilesitoides philippinensis is proposed as a new genus and species. This fauna shows close similarity to that of the Tomochi Formation, Choshi Group, and the Miyako Group in Japan. These faunas were in a warm current regime that extended from the equator. The age and palaeogeographical significance of the assemblage is discussed.

Stratigraphically important ammonites from the Campanian–Maastrichtian boundary interval of the Middle Vistula River section, central Poland

ABSTRACT Machalski, M. 2012. Stratigraphically important ammonites from the Campanian-Maastrichtian boundary interval of the Middle Vistula River section, central Poland. Acta Geologica Polonica, 62 (1), 91-116. Warszawa. The formal definition of the Global Stratotype Section and Point (GSSP) for the base of the Maastrichtian Stage at Tercis, southwest France, is based on the first or last occurrences of twelve taxa, including three species of ammonites, Pachydiscus (Pachydiscus) neubergicus, Diplomoceras cylindraceum, and Nostoceras (Nostoceras ) hyatti. The taxonomy and stratigraphical distribution of these, and allied forms, are studied, on the basis of material from the opoka succession across the upper Campanian-lowermost Maastrichtian boundary in the Middle Vistula River section, central Poland. In view of the imprecise location of two ammonite GSSP markers in that section, a direct ammonite-based correlation with the GSSP at Tercis is impossible. However, data available indicate that in the Middle Vistula River section the first occurrence of Pachydiscus (P.) neubergicus and the last occurrence of Nostoceras (N.) hyatti are situated significantly higher than the first occurrence of Belemnellalanceolata, the traditional belemnite marker for the base of the Maastrichtian Stage in the Boreal Realm, and that Diplomoceras cylindraceum appears significantly below this level. There are no unequivocal records of Pachydiscus (P.) neubergicus from the Belemnella lanceolata Zone s.l. in the Middle Vistula River section.

A new species of Andiceras Krantz (Cephalopoda: Ammonoidea) from the Late Jurassic-Early Cretaceous of the Neuquen Basin, Mendoza, Argentina. Systematics and Biostratigraphy

A new ammonoid species of the Andean endemic genus Andiceras Krantz is here described from upper Tithonian-lower Berriasian levels of the Vaca Muerta and Chachao Formations of the Mendoza Group, in north-western Mendoza Province, Argentina. Andiceras planulatus sp. nov. lies between the last beds with representatives of the Substeueroceras koeneni Assemblage Zone of late Tithonian age and the first bed yielding elements of the Argentiniceras noduliferum Assemblage Zone of early Berriasian age, according to the traditional Andean ammonoid zonation scheme. The particular stratigraphic position of Andiceras planulatus together with its easily recognizable morphology makes this species a good potential biostratigraphic marker for the Jurassic-Cretaceous boundary beds in the Andean region.

Cephalopod and brachiopod fossils from the Pacific: Evidence from the Upper Cretaceous of the Magellan Seamounts

Maastrichtian cephalopods and a brachiopod were dredged from the Butakov, Fedorov, Kotsebu, Il’ichev, Govorov, Gelendzhik, and Ita-Mai-Tai guyots in the Magellan Seamounts. The ammonoids Hypophylloceras sp., Phyllopachiceras sp., Anagaudryceras? sp. A, Anagaudryceras? sp. B, Gaudryceras aff. propemite Marshall, Gaudryceras sp., and Pseudophyllites cf. indra (Forbes), and the single brachiopod Basiliolidae gen. and sp. indet. are a first discovery in this oceanic region, following earlier finds of belemnites (Dimitobelus? sp., Dimitobelidae gen. and sp. nov., and Belemnitella? sp.), and two ammonoid species (Zelandites aff. japonicus Matsumoto and Tetragonitidae gen. and sp. indet.). The Late Cretaceous Magellan Seamounts dimitobelid belemnite fauna shows affinities with southern high latitude forms (New Zealand) and the ammonoid fauna with northern, middle and high latitude ones (Hokkaido-Sakhalin and/or Kamchatka). This suggests by the end of the Cretaceous major surface palaeo-currents from S and N sides in direction of the central Palaeo-Pacific, a position coinciding with the plate tectonic reconstruction of the Magellan Seamounts.

Detailed stratigraphy of the Jurassic-Cretaceous boundary beds of the Lena River lower reached based on ammonites and Buchiids

The Jurassic-Cretaceous (Volgian-Lower Valanginian) boundary deposits in the lower reaches of the Lena River (near the village of Chekurovka and on the Cape Chucha, North Yakutia) are described bed-by-bed. The taxonomic composition of ammonites and bivalves (mainly Buchia) is determined. The species assemblages are biostratigraphically analyzed and biostratigraphic units of the bed and zone ranks are recognized based on by Buchia and ammonites. Analogues of the Buchia Zones (buchiazones) Unschensis, Okensis, Tolmatschewi, Inflata and ammonite zones Exoticus, Okensis, Sibiricus, Analogus, Mesezhnikowi and ? Klimovskiensis are established in this region for the first time. A correlation of the distinguished buchiazones with the buchiazones of the Nordvik Peninsula, New Siberian Islands (Stolbovoy Island), the basin of the Anyuy River, and Northern California is proposed. The ammonite assemblages constantly contain phylloceratids; some stratigraphic intervals also contain lytoceratids; this is probably connected with a Paleo-Pacific influence. Unlike the ammonite assemblages of the same age of the Khatanga depression, the subzones cannot be distinguished in the Sibiricus and Kochi Zones of the Lower Lena due to differences in the stratigraphic ranges of some ammonite species. Hectoroceras in the Kochi zone is very rare and Praetollia predominates here.

Simoceras szentei n. sp., a new ammonite species from the lowermost Tithonian of the Gerecse Mountains (Hungary) - the earliest record of the genus

Simoceras szentei n. sp., a new ammonite species from the lowermost Tithonian of the Gerecse Mountains (Hungary) - the earliest record of the genus

First record of the leptonectid ichthyosaur Eurhinosaurus longirostris from the Early Jurassic of Switzerland and its stratigraphic framework

An incomplete skull of the leptonectid ichthyosaur Eurhinosaurus longirostris found in the Rietheim Member (previously “Posidonienschiefer”; Toarcian, Early Jurassic) of Staffelegg, Canton Aargau, is the first record from Switzerland of this taxon and supports the status of Eurhinosaurus longirostris as a palaeobiogeographic very widespread ichthyosaur species in the Early Toarcian of Western Europe. Being from either the Bifrons or Variabilis zone, it is one of the youngest records of Eurhinosaurus and one of the few diagnostic ichthyosaur finds from this time interval. The partial skull is well articulated and preserved three-dimensionally in a carbonate concretion. Both the mode of preservation of the ichthyosaur and an associated ammonoid (Catacoeloceras raquinianum) provided the age of the concretion, which had been collected from scree. Taphocoenosis and taphonomy show the C. raquinianum to be one of few non re-worked fossils recorded from the Early to Late Toarcian boundary (Bifrons/Variabilis zone) of northern Switzerland in general and of this ammonite species in particular. The Toarcian section at Staffelegg differs from other localities where strata of the same age are exposed with respect to facies variations of the Rietheim Member (previously “Posidonienschiefer”, Early Toarcian) and the extraordinarily high thickness of the Gross Wolf Member (previously “Jurensis-Mergel”, Late Toarcian).

A new early Smithian ammonoid fauna from the Salt Range (Pakistan)

Recent extensive investigations in the Salt Range (Pakistan) yielded abundant, well-preserved ammonoid faunas of earliest to latest Smithian age that provided the basis for a major revision of Smithian ammonoid taxonomy and for the establishment of a high-resolution biostratigraphic sequence. Here, an additional new ammonoid fauna of typical early Smithian affinity from the uppermost part of the Ceratite Sandstone of the Nammal Gorge section is described. The new fauna, termed Euflemingites cirratus beds, is bracketed between the underlying early Smithian “Flemingites flemingianus beds”, here renamed Clypeoceras superbum beds, and the overlying middle Smithian Brayardites compressus beds. Comparison with a recently published high-resolution biochronological scheme for the Smithian of the NIM (northern Indian Margin) based on the Salt Range, Spiti (Himachal Pradesh, northern India) and Tulong (South Tibet) basins shows that the Euflemigites cirratus fauna correlates with the Dieneroceras beds from Spiti based on the common occurrence of the ammonoid species Kraffticeras pseudoplanulatum. The trans-panthalassic biogeographical distribution of Euflemingites cirratus allows correlating the new ammonoid fauna with part of the Meekoceras gracilitatis ammonoid zone of western USA. Three new species (Kashmirites weisserti, Arctoceras schalteggeri and Vercherites wyleri) are described.

Ammonite paleobiogeography during the Pliensbachian–Toarcian crisis (Early Jurassic) reflecting paleoclimate, eustasy, and extinctions

The Pliensbachian–Toarcian crisis (Early Jurassic) is one of the major Mesozoic paleoecological disturbances when ca. 20% of marine and continental families went extinct. Contemporaneously, profound paleobiogeographical changes occurred in most oceanic domains including a disruption of ammonite provincialism during the Early Toarcian. Here, we quantitatively reappraise the structure and evolution of paleobiogeographical patterns displayed by ammonite faunas before, during, and after the biological crisis, over a time-interval including 13 biochronozones. The high-resolution study presented here involves the use of hierarchical Cluster Analyses, non-metric Multi-Dimensional Scaling methods, and Bootstrap Spanning Network approaches that we apply to a large database including 772 ammonite species from 16 northwestern Tethyan and Arctic basins. Our results confirm a robust faunal dichotomy between Euro-Boreal and Mediterranean areas throughout the Pliensbachian, with the first emergence of an Arctic biome during the cooling regressive event of the Spinatum Zone. Whatever its complexity, Pliensbachian provincialism could be directly linked to paleogeographical barriers and to latitudinal paleoclimatic and paleoecological contrasts. During the Early Toarcian, this pattern was progressively lost, with northward expansions of Mediterranean ammonites during the Tenuicostatum Zone, followed by a strong interprovincial mixing during the Falciferum Zone. This faunal homogenization results from the combination of several parameters including a major sea-level rise facilitating basinal connections, a global warming event stretching the spatial range limits of southern taxa, and a mass extinction preferentially removing endemic species. Ammonite provincialism, although slightly different, was progressively re-established during the cooling regressive trend of the Middle Toarcian. These results therefore suggest a paramount influence of paleoclimatic, eustatic, and extinction constraints on the paleobiogeography of Early Jurassic ammonites, even if some threshold effects or independent biological factors may sporadically complicate the patterns.

Microstructure and mineralogy of the outer calcareous layer in the lower jaws of Cretaceous Tetragonitoidea and Desmoceratoidea (Ammonoidea)

Tanabe, K., Landman, N.H. & Kruta, I. 2011: Microstructure and mineralogy of the outer calcareous layer in the lower jaws of Cretaceous Tetragonitoidea and Desmoceratoidea (Ammonoidea). Lethaia, Vol. 45, pp. 191–199. Based on the differences in their relative size, overall shape, structure and the degree of development of an outer calcified covering, lower jaws of the Ammonoidea have been classified into four morphotypes: normal, anaptychus, aptychus and rhynchaptychus types. However, detailed microstructural and mineralogical comparison of these morphotypes has not yet been addressed. This article documents the results of SEM and XRD observations of the lower jaws of three Late Cretaceous ammonoid species belonging to the Tetragonitoidea (Anagaudryceras limatum) and Desmoceratoidea (Pachydiscus kamishakensis and Damesites aff. sugata), based on excellent material preserved in situ within the body chamber, and retaining an aragonitic shell wall. The lower jaws of the three species are assigned to an intermediate form between anaptychus and aptychus types for the first two species and the rhynchaptychus type for the third species. Their black, presumably originally chitinous outer lamella is wholly covered with a calcareous layer. The calcareous layer is composed of aragonite in D. aff. sugata and A. limatum, and calcite in P. kamishakensis. The microstructure of the outer calcareous layer differs among the three species, i.e. granular in A. limatum, spherulitic prismatic in D. sugata, and prismatic in P. kamishakensis, all of which can be distinguished from the lamellar and spongy structure of the outer-paired calcitic plates of the typical aptychus-type lower jaws in some Jurassic and Cretaceous Ammonitina and Ancyloceratina. Our study suggests that most Jurassic and Cretaceous ammonoids possessed an outer calcareous layer in their lower jaws, although its mineralogy, microstructure and relative thickness vary among different taxa. □Ammonoidea, Cretaceous, Desmoceratoidea, lower jaw, microstructure, Tetragonitoidea.

Late Albian (Early Cretaceous) ammonites from the Provincial Formation of central Cuba

A late Albian ammonite assemblage from the Provincial Formation of Villa Clara Province, Cuba is described. The Provincial Formation is a lithostratigraphic unit of Albian–Cenomanian age extensively exposed in central Cuba and formed within a part of the Caribbean Tethys that was volcanic during the Cretaceous. The formation is mainly composed of calcareous, terrigenous marine, and volcano-sedimentary deposits characterized by a series of micritic limestones intercalated with marls, sandstones, calcareous conglomerates, ash, and tuffaceous material. A rich assemblage of ammonites recovered from the calcareous biomicrites and marls is of late Albian ( Stoliczkaia dispar Zone, Mortoniceras rostratum Subzone) age. The ammonite fauna shows a strong Tethyan affinity, and only a single hoplitid ammonite species was recorded. Although scarce, the first Cuban report of this and other boreal ammonite species now allows precise correlations to be made between Cuba and Albian sediments elsewhere in the world.

Temporal patterns in disparity and diversity of the Jurassic ammonoids of southern Germany

A morphometric analysis of the characteristic whorl cross-sections of 1,200 Jurassic ammonoid species from southern Germany enabled us to characterise their morphospace. The successive Jurassic ammonoid faunas of southern Germany show characteristic patterns in morphospace occupation. While Early and Middle Jurassic ammonoids occupy limited areas of the morphospace range, the Late Jurassic ammonoids cover the entire spectrum. The ammonoids are characterised by an overall increase of both taxonomic diversity and morphological disparity in the course of the Jurassic. Strong fluctuations occur until the middle Late Jurassic, followed by a diversity decrease in the early Kimmeridgian and a disparity reduction in the early Tithonian. While diversity and disparity show similar progression during most of the Early Jurassic, they diverge subsequently and show only poor correlation until the end of the Jurassic. Particularly in the Middle Jurassic diversity and sea level changes correlate strongly. Neither temporal patterns in diversity nor disparity support the hypothesis of a mass extinction event in the early Toarcian. Significant changes in diversity and disparity in the early Callovian support a putative migration event of Boreal ammonoids into the Tethyan realm. <br><br> doi:<a href="http://dx.doi.org/10.1002/mmng.201000016" target="_blank">10.1002/mmng.201000016</a>

Temporal patterns in disparity and diversity of the Jurassic ammonoids of southern Germany

Abstract. A morphometric analysis of the characteristic whorl cross-sections of 1,200 Jurassic ammonoid species from southern Germany enabled us to characterise their morphospace. The successive Jurassic ammonoid faunas of southern Germany show characteristic patterns in morphospace occupation. While Early and Middle Jurassic ammonoids occupy limited areas of the morphospace range, the Late Jurassic ammonoids cover the entire spectrum. The ammonoids are characterised by an overall increase of both taxonomic diversity and morphological disparity in the course of the Jurassic. Strong fluctuations occur until the middle Late Jurassic, followed by a diversity decrease in the early Kimmeridgian and a disparity reduction in the early Tithonian. While diversity and disparity show similar progression during most of the Early Jurassic, they diverge subsequently and show only poor correlation until the end of the Jurassic. Particularly in the Middle Jurassic diversity and sea level changes correlate strongly. Neither temporal patterns in diversity nor disparity support the hypothesis of a mass extinction event in the early Toarcian. Significant changes in diversity and disparity in the early Callovian support a putative migration event of Boreal ammonoids into the Tethyan realm. doi: 10.1002/mmng.201000016

An unusually diverse mollusc fauna from the earliest Triassic of South China and its implications for benthic recovery after the end-Permian biotic crisis

A single carbonate coquinoid lens from the Griesbachian (Early Triassic) of Shanggan, South China, yielded 11 bivalve species described in this study in addition to four gastropod and one ammonoid species reported elsewhere. This makes the Shanggan fauna one of the richest mollusc faunas from the early post-extinction interval after the end-Permian mass extinction event. Four of the present genera are long-term survivors, five are holdovers that went extinct at the end of the Griesbachian or later in the Early Triassic, and seven first appear in the Griesbachian. Three new bivalve species are described: Myalinella newelli nov. sp., Scythentolium scutigerulus nov. sp., and Eumorphotis shajingengi nov. sp. The genus Astartella, previously assumed to have vanished at the end of the Permian, is reported for the first time from the Early Triassic, which also removes Astartidae from Early Triassic Lazarus taxa. The small growth size of the Astartella specimens supports an earlier hypothesis that many of the Early Triassic Lazarus taxa did not survive in unknown refuges but were simply overlooked due to the scarcity of easily observable large-sized specimens. Ecologically, a comparatively high proportion of infaunal bivalve species (4/11) is remarkable for the early post-extinction interval, supporting the impression of a relatively advanced recovery state. Moreover, abundance-data of the bivalve-gastropod community reveal a remarkably low dominance index (D = 0.17) that is suggestive for advanced recovery and stable environmental conditions. It is proposed that the Shanggan fauna represents a late Griesbachian benthic recovery event that coincided with the appearance of similarly diverse benthic faunas in Oman and Primorye. A high proportion of genera that have previously not been reported from the Early Triassic indicate that the prevalence of poor preservation conditions is a major obstacle in identifying early phases of recovery from the greatest crisis in the history of metazoan life. The early recovery of benthic faunas reported in this study questions previous claims of a prolonged lag phase as a consequence of the extraordinary extinction magnitude or the persistence of adverse environmental conditions.

Revision of some problematic Early Cretaceous ammonite species described by H. Coquand (1880)

The finding and analysis of the type specimens of some Barremian ammonite species described, but not depicted by Coquand (1880, Bull. de l’Acad. d’Hippone 15, 1-449), has led us to the revise their current interpretation. The purpose of the manuscript is to re-describe the three most commonly used Early Cretaceous species described by Coquand (1880, Bull. de l’Acad. d’Hippone 15, 1-449) and is to consider one species as a nomen dubium. The original type material of Holcodiscus diversecostatus, Holcodiscus metamorphicus and Heinzia heinzi shows that these species have been usually misidentified in the literature. We designate lectotypes for these species, thus amending some previous invalid designations, and fully discuss their actual status and synonymy. In addition, we conclude that the use of the nominal species “Ammonites mazuca” should be fully avoided, it is considered as a nomen dubium. The type material is apparently seems to be lost so its application is very doubtful.

Diversity dynamics of ammonoids during the latest Bajocian and Bathonian (Middle Jurassic) in the epicratonic Polish Basin

Results of the analysis of Middle Jurassic (latest Bajocian–Bathonian) ammonoid diversity patterns in the Polish Basin are reported. The data used in this study are based on a large number of ammonoid specimens, collected bed-by-bed, from the Polish Jura in south-central Poland, as well as on existing literature. The ammonoid diversities, both at the genus and species levels, have been calculated for particular ammonite zones and subzones and compared with the regional transgressive–regressive cycles for the Polish Lowlands and Hallam’s global sea-level curve. The patterns of ammonoid diversity dynamics seem to be well correlated with global sea-level fluctuations. Particular diversity peaks correspond with major transgressive episodes. Three main regional bio-events related to transgressions have been distinguished for the Polish Basin: (1) the Latest Bajocian (Bomfordi Subchron) bio-event is related to a short-lasting immigration of Tethyan ammonoids; (2) a late Early Bathonian (Tenuiplicatus Chron) bio-event corresponds to a proliferation of Asphinctites tenuiplicatus (Brauns), most probably as a result of transgression-driven eutrophication of a shallow-marine environment; during this time, the immigration (passive dispersion?) of some single Tethyan species is also observed; and (3) a Late Bathonian (Hodsoni and Orbis chrons) bio-event corresponds to the highest ammonoid species diversity peaks and most probably is related to the major transgression during the Bathonian, which allowed easy migration of several species via several newly opened sea-ways.

Size distribution of the Late Devonian ammonoid Prolobites: indication for possible mass spawning events

Worldwide, the ammonoid genus Prolobites is only known from a few localities, and from these fossil beds almost all of the specimens are adults as shown by the presence of a terminal growth stage. This is in marked contrast to the co-occurring ammonoid genera such as Sporadoceras, Prionoceras, and Platyclymenia. Size distribution of specimens of Prolobites from three studied localities show that, unlike in the co-occurring ammonoid species, most of the material belongs to adult individuals. The morphometric analysis of Prolobites delphinus (Sandberger & Sandberger 1851) demonstrates the intraspecific variability including variants with elliptical coiling and that dimorphism is not detectable. The Prolobites material shows close resemblance to spawning populations of Recent coleoids such as the squid Todarodes filippovae Adam 1975. Possible mass spawning events are discussed in the context of the size distribution and limited geographic range of Prolobites. Finally, the potential fecundity and brooding behaviour of Prolobites is hypothesized using the examples of post spawning egg care in Recent coleoids.

Intraspecific variation of hatchling size in Late Cretaceous ammonoids from Hokkaido, Japan: implication for planktic duration at early ontogenetic stage

Tajika, A. & Wani, R. 2011: Intraspecific variation of hatchling size in Late Cretaceous ammonoids from Hokkaido, Japan: implication for planktic duration at early ontogenetic stage. Lethaia, Vol. 44, pp. 287–298. Intraspecific variations of the early shell dimensions (ammonitella and protoconch diameters) of two Late Cretaceous (earliest Campanian) ammonoid species (Gaudryceras tenuiliratum and Hypophylloceras subramosum) from the Haboro and Ikushumbetsu areas, Hokkaido, Japan, show no significant difference between these areas that are approximately 110 km apart. The geographic distributions of G. tenuiliratum and H. subramosum are supposed to be mainly controlled by the flotation and transportation during the embryonic stage within floating egg masses and/or post-embryonic stage because of their small hatchling sizes (1.18–1.46 mm in diameter for G. tenuiliratum, and 0.91–1.13 mm in diameter for H. subramosum), suggesting these two ammonite species at the embryonic and/or post-embryonic stages were transported at least 110 km. Postulating that the velocity of palaeocurrent around the Haboro and Ikushumbetsu areas during the Cretaceous Period was 0.25 m/s, similar to those in the modern ocean current flowing off the eastern Pacific coast of Hokkaido, the egg masses and/or hatchlings of G. tenuiliratum and H. subramosum were buoyant and transported more than 5 days. The preliminary comparison of hatchling size through time suggests that the hatching sizes of H. subramosum in Hokkaido increased slightly from the Middle Turonian until the earliest Campanian (during about 7 Myr). □ammonoid, hatchling, paleoecology, variation, Cretaceous.

New species of Dzhulfian (Late Permian) ammonoids from the Hambast Formation of Central Iran

New ammonoid species of the genus Araxoceras (A. abarquense sp. nov. and A. iranense sp. nov.) are described from the Early Dzhulfian (Early Wuchiapingian) beds of the Hambast Formation of Central Iran (Abade). The data on the distribution of Paraceltites (Paraceltites sp.) and Vedioceras (V. umbonovarum Ruzhencev), for the first time discovered in the Dzhulfian beds of Abade, and of Eoaraxoceras are provided. The correlation of the Late Permian assemblages of ammonoids of Iran and Transcaucasia is amended.