Species-group Names Research Articles

On the authorship of Canthon rutilans cyanescens ‘Harold, 1868’ (Coleoptera, Scarabaeidae, Scarabaeinae, Deltochilini): the treatment of varieties and nomina nuda by a 19th-century zoologist

Ever since Schmidt’s (1922) review of Canthon, the authorship of the species-group name Canthon rutilans cyanescens (Coleoptera, Scarabaeidae) has been almost unanimously attributed to Harold (1868). Here, we argue that this is mistaken. This species-group name first entered the literature as a nomen nudum in Lacordaire’s 1830 memoirs of his South American beetle explorations, and there its authorship was credited to Dejean. Later, it was listed in Harold’s (1868) Canthon revision as an unavailable name referring to what he considered to be a blue variety of Canthon rutilans Castelnau, 1840. In keeping with his usual nomenclatural treatment of nomina nuda as unavailable and with his procedure of not naming infraspecific taxa, Harold did not consider ‘cyanescens Dejean’ as the valid name of that ‘blue variety’ of C. rutilans. His citation, therefore, does not make the name available by not complying with Article 11.5 of the Code. The first author to fulfil all the availability requirements of the Code was Schmidt (1922), who both treated the name as valid for a taxon and accompanied it with an indication—more specifically, an indication to Harold’s description of the blue variety of C. rutilans. It is to Schmidt (1922) that the authorship of cyanescens must, thereby, be credited. One of the specimens used by Harold to produce the description indicated by Schmidt was found by us in the Paris Museum and it is here designated as the lectotype of Canthon rutilans cyanescens.

A taxonomic revision of the New World genus Agamopus Bates, 1887 (Coleoptera: Scarabaeinae: Ateuchini)

Differential diagnoses, updates on distribution ranges, and an illustrated identification key are given for all species of Agamopus. The first record for A. castaneus from Brazil is given. Type specimens of all valid species-group names are studied and illustrated. Notes on the natural history of Agamopus species, as well as brief phylogenetic comments, are presented. The new species Agamopus joker sp. nov. is described based on a male from Paraná, southern Brazil. Lectotypes are designated for Canthon unguicularis Harold, 1883 and Agamopus lampros Bates, 1887. We also propose the transfer of Agamopus convexus Balthasar, 1965 to the Ateuchus ovalis species group (Ateuchus convexus comb. nov.) based on morphological features of the protibiae, clypeal teeth, and the posterior margin of the pronotum and pygidium.

Revalidation, synonymies, and notes in American Callichromatini (Coleoptera, Cerambycidae, Cerambycinae).

The unpublished formal synonymy of Callichroma chiriquina Bates, 1879, C. opiparum Bates, 1874, C. compressipes Casey, 1912, and C. regalis Casey, 1912 is commented. Schwarzerion carinatum Schmidt, 1924 is removed from the synonym of Callichroma holochlorum holochlorum Bates, 1872, and transferred to the synonymy of Callichroma euthalia Bates, 1879. The species group name euthalium is corrected. The type-localities of Schwarzerion carinatum are reported as Colombia and West Indies, and the institution where one of the syntypes is deposited is corrected. Callichroma collarti Fuchs, 1959 is revalidated, and Callichroma magnificum Napp Martins, 2009 is considered its junior synonym. The status of the specimens of the type series of Callichroma collarti is commented. Additionally, the holotype of Callichroma opiparum Bates, 1874, and the lectotype of Callichroma chiriquina Bates, 1879 are illustrated for the first time.

Annotated catalogue of the Tachinidae (Insecta, Diptera) of Chile.

Annotated catalogue of the Tachinidae (Insecta, Diptera) of Chile.

Description of four new species of Anelaphus Linsley (Coleoptera, Cerambycidae, Cerambycinae, Elaphidiini), correction of the spelling of species-group names, and description of the female of Anelaphus pilosus Chemsak and Noguera.

Eight species-group names of longhorned woodboring beetles in the genus Anelaphus are corrected. Four new species of Anelaphus Linsley, 1936 are described: A. steveni Santos-Silva, sp. nov., from Mexico (Chiapas); A. erici Santos-Silva, sp. nov., from Nicaragua (Granada); A. izabalensis Santos-Silva, sp. nov., from Guatemala (Izabal); and A. zacapensis Santos-Silva, sp. nov., from Guatemala (Zacapa). New Mexican state records for Anelaphus undulatus (Bates, 1880) are provided. The female of Anelaphus pilosus Chemsak and Noguera, 2003 is described for the first time, and the species is newly recorded from Guatemala. The inclusion of Anelaphus nitidipennis Chemsak and Linsley, 1968 in this genus is questioned.

The green mole,Astromycter prasinatusT. M. Harris, 1825 (Mammalia: Eulipotyphla: Talpidae): an origin story

Thaddeus William Harris described the green mole of Maine, Condylura prasinata (Mammalia: Eulipotyphla: Talpidae), in the July 1825 issue of the Boston Journal of Philosophy and the Arts, and this was considered the original description of the species throughout most of the nineteenth century. In the early twentieth century, taxonomists began instead to cite an earlier notice in the June 1825 issue of the American Journal of Science and Arts. This short article also described the species, but also established a separate genus for it, Astromycter, despite Harris’s indications elsewhere that the species was congeneric with the star-nosed mole, Condylura cristata ( Linnaeus, 1758 ). Moreover, the American Journal of Science and Arts article cited as its source the “ Machias Star”, indicating the possibility of an even earlier description of the animal. With Astromycter prasinata in synonymy with C. cristata for over a century, little effort has been exerted to determine whether earlier source materials exist or why, within two months, the green mole was allocated to two different genera by its describer. The question is taxonomically relevant today because C. prasinata predates C. cristata nigra Smith, 1940 , as an available name for north-eastern populations of star-nosed moles. If subspecies of C. cristata are to be recognized, the north-eastern subspecies should correctly bear the name C. cristata prasinata. In fact, authority for both genus- and species-group names for the green mole have been misattributed since 1825. The descriptions of C. prasinata by Thaddeus William Harris in the Boston Journal of Philosophy and the Arts and American Journal of Science and Arts were preceded by at least three published descriptions of A. prasinatus by Thaddeus Mason Harris, his father.

Case 3847 – Simopithecus oswaldi Andrews, 1916 (currently Theropithecus oswaldi; Mammalia, Primates, Cercopithecidae), proposed conservation by reversal of precedence with Cynocephalus atlanticus Thomas, 1884

The purpose of this application, under Articles 23.9.3 and 81.1 of the Code, is to conserve the usage of the species-group name Simopithecus oswaldi Andrews, 1916 by giving it precedence over its senior subjective synonym Cynocephalus atlanticus Thomas, 1884. Theropithecus is a common to dominant member of the extinct primate community across Africa after 4 million years ago (Jablonski & Frost, 2010) and often co-occurred with extinct humans (Hominini); fossils are also known rarely across Eurasia (Roberts et al., 2014). Most fossil samples are currently included in Theropithecus oswaldi (Andrews, 1916), which is often divided into chrono-geographic subspecies. Cynocephalus atlanticus Thomas, 1884 was not recognized as a member of Theropithecus until 1973, and this nomen has seldom been used, but if it were shown (as seems likely) to be conspecific with Simopithecus oswaldi Andrews, 1916, widespread paleontological usage would be upset. It is thus proposed to give conditional precedence to the later name, which would still permit the use of Theropithecus atlanticus as a distinct species or subspecies of Theropithecus oswaldi. Lectotypes are designated for Simopithecus oswaldi olduvaiensis Leakey & Whitworth, 1958 and Simopithecus oswaldi hopefieldensis Singer, 1962.

Isaac Lea's (1792–1886) Substitutions and Other Modifications of His Own Names of Molluscan Species

The American conchologist Isaac Lea had a long and productive career during which he introduced more than 1,800 names of molluscan species between 1827 and 1874, the majority North American land and freshwater species. His idiosyncratic way of publishing, by describing new taxa multiple times in duplicated and variously modified journal papers as well as in retitled collections of extracts, has led to considerable difficulties in determining accurate dates of his original descriptions. He considered the reading and presentation of a manuscript before a “learned society” as the date of its original publication and therefore his own recorded dates must be approached with caution. The problem of interpreting Lea's names was compounded by the fact that Lea frequently modified his own new taxonomic names in his subsequent publications, and often without providing justification for the change. The various name versions were inconsistently applied by subsequent authors. The current paper analyses Lea's substitutions and other modifications of his own names of molluscan species and discusses 131 of his publications in this context, for which attempts at precise dating were made. The status of these “replacements” of Lea's taxonomic names were evaluated under the current ICZN Code (1999). A few additional cases of changes of Lea's names by other authors are included; these are instances in which Lea had missed a primary homonym or a secondary homonym was discovered later. While some of the names here discussed are currently resting in synonymy, others are of greater current relevance as they are in use as valid names for taxa of conservation/management concern, including IUCN and U.S.-federally listed species. The following 98 discussions of often-complex name changes involve 348 species-group names and their subsequent spellings, 231 of which were introduced by Lea. Based on their current taxonomic status, the names belong to 4 bivalve and 11 gastropod families, with the vast majority concentrated in Unionidae and Pleuroceridae. Under the current ICZN Code, the investigated changes fall into many different categories, which impacts their nomenclatural availability: 35 names changed by Lea (plus 6 by other authors) are interpreted as substitute names for junior primary homonyms, 7 of which were unnecessary; 1 name (plus 6 by others) as substitute names for junior secondary homonyms, 1 of which was unnecessary; 13 names as justified emendations; 10 names (plus 1 by another) as unjustified emendations that have not entered prevailing usage; 1 name determined by his first reviser action; 37 names (plus many by others) as incorrect subsequent spellings that are not in prevailing usage; and 1 incorrect subsequent spelling that is shown to be in prevailing usage and thus considered a correct original spelling. The concept of “prevailing usage,” ill-defined under the current (1999) ICZN Code, is addressed in this context. The following names in current use are corrected as follows: Helix balasteriana I. Lea, 1840 (taxon inquirendum, ?Dyakiidae), Lioplax cyclostomatiformis (I. Lea, 1844), Diplodon demararaensis (I. Lea, 1859), Iheringella isocardiodes I. Lea, 1856, Dilatata brongniartiana (I. Lea, 1843), Elliptio nasutidus (I. Lea, 1863), Elliptio pullata (I. Lea, 1857), Pleurocera picta (I. Lea, 1841) [instead of P. curta (Haldeman, 1841), which is preoccupied] and Beringiana youconensis (I. Lea, 1847); many others are confirmed in their recently applied spellings. This work serves as an example that the basic endeavor of summarizing available and valid names from the published literature is not always a straightforward task.

World Catalogue of Hypophlaeini Billberg, 1820 (Coleoptera: Tenebrionidae: Diaperinae)

This catalogue includes all valid genus and species-group names of Hypophlaeini (Coleoptera: Tenebrionidae), their available synonyms, and misspellings. Type depository, distribution, and basic morphological data are given for every treated species-group name. The introductory part of this publication contains a summary of the current knowledge on larval and adult morphology within the tribe. The following new synonymies are proposed: Cheilopoma castaneumMurray, 1867 =Cheilopoma kraatziGrouvelle, 1914syn. nov.; Corticeus maehleri (Kulzer, 1957) =Hypophloeus mayumiaeMasumoto, 1983syn. nov. The lectotypes of Hypophloeus rufipesFabricius, 1801 and Hypophloeus constrictusGebien, 1921 are designated. Corticeus africanus uhligiBremer, 1995 is recognized as an independent species: Corticeus uhligiBremer, 1995stat. nov. Subgeneric classification of Corticeus is adjusted. According to the results Hypophlaeini is composed of 261 species classified in the following six genera: CheilopomaMurray, 1867 (monotypic), CorticeusPiller & Mitterpacher, 1783 (255 species and subspecies), IschnarthronGebien, 1921 (monotypic), Myonophloeus Bremer & Lillig, 2017 (monotypic), PogonoxenusWasmann, 1899 (monotypic), and TyphlophloeusJeannel & Paulian, 1945 (two species).

A modularity analysis helps improving the structure of the International Code of Zoological Nomenclature.

BackgroundIn a recent work I transformed a complex and integrated text like the International Code of Zoological Nomenclature into a network of interconnected parts of text. This new approach allowed understanding that a continuous body of text cannot accurately reflect the true structure of the Code, and provided a scientific methodology to identify a priori parts that could be affected by future revisions. In this next step, I investigate further the structure of the Code, seeking to use the network in order to identify the various conceptual communities grouping the various articles and other text items of the Code.MethodsUsing the first version of the network of the Code, I perform a comprehensive modularity analysis in two rounds: the first round aims to identify the fewest and largest communities or modules for the entire network, whereas the second round identifies the sub-modules within each larger module. The potential conflicts between the current structure of the Code and the module composition are evaluated with a parcellation analysis.ResultsThe optimal modularity search identified 10 different modules in the entire network of varying size (ranging from 75 to 200 nodes). Each module can be further divided into smaller modules, that all-together allow describing the 65 conceptual groups of text items in the Code. Parcellation analysis revealed that two-thirds of the current chapters of the Code are in excellent or good accordance with the recovered conceptual modules, whereas the current composition of six chapters is in serious conflict with the conceptual structure of the Code.DiscussionJudging only the composition and not the order of appearance of the Articles in the Chapters of the Code, I show that in many cases the current structure of the Code is found to correspond quite well to the concepts presented therein. The most important conflict is found on the provisions related to the various groups of names governed by the Code: family-, genus-, and species-group names. Currently, these provisions are spread out in different Articles in different Chapters, along the entire length of the Code. The modularity analysis suggests that re-organizing the Code in chapters that will deal with all aspects related to a given group (e.g., chapters including information on name formation, availability, typification, and validity for a given group), could potentially improve reader experience and, consequently, the applicability of the Code.

On the availability of the name Poecilia kempkesi Poeser 2013(Cyprinodontiformes: Poeciliidae).

The poeciliid species, Poecilia kempkesi Poeser, 2013, was the fourth species of the subgenus Acanthophacelus Eigenmann, 1907 to be described, based on individuals from a single urban anthropized locality close to Paramaribo, Suriname (Poeser, 2013). The description itself lacked any section clearly distinguishing the new species from the remaining species of Poecilia Bloch Schneider 1801, and in particular from the species of the subgenus Acanthophacelus, type species Poecilia reticulata Peters, 1859. According to Article 13 of the International Code of Zoological Nomenclature (ICZN, 1999) the criteria of availability for a species-group name are:.

Catalogue of rhysodine beetles (Coleoptera: Carabidae) from East Asia.

An annotated catalogue for 22 extant species of the subfamily Rhysodinae (Coleoptera: Carabidae) from East Asia is presented, with 7 genera classified into 3 tribes: I. Clinidiini Bell Bell, 1978: Clinidium Kirby, 1830 (1 sp.) and Rhyzodiastes Fairmaire, 1895 (4 spp.); II. Omoglymmiini Bell Bell, 1978: Arrowina Bell Bell, 1978 (1 sp.), Yamatosa Bell Bell, 1979 (6 spp.), Shyrodes Grouvelle, 1903 (1 sp.), and Omoglymmius Ganglbauer, 1891 (8 spp.); III. Rhysodini Laporte, 1840: Rhysodes Germar, 1822 (1 sp.). For each genus- and species-group name, synonymy, type locality, type depository, distribution, and bibliography are provided where appropriate. A modified key to genera of rhysodine beetles from East Asia is compiled.

Validation of Stenus (Hypostenus) primivenatus and Stenus (Hypostenus) yiae (Coleoptera, Staphylinidae, Steninae).

Holotype depository was not indicated for the new species, Stenus (Hypostenus) primivenatus and Stenus (Hypostenus) yiae, described as new in the publication Zhao Zhou (2008). Missing this important message made these two species-group names unavailable according to the fourth edition of the International Code of Zoological Nomenclature (the Code, ICZN, 1999: Article 16.4). This problem was first denoted by Schülke, M. Smetana, A. (2015).

Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue.

412 species-group names (including 11 replacement names), and 14 genus-group names of the Alycaeidae have been introduced to date. Type materials of 85% (336) of the known species and subspecies were examined, a further 5% (19) of the taxa were studied using available non-type material, and for another 6% (22) the original descriptions were sufficiently detailed to evaluate their taxonomic status. Only 3% of the taxa (12) could not be examined. Special attention was paid to the sculpture of the embryonic whorls and the sutural tube-microtunnel system in order to provide a novel classification for this group.In this study 363 taxa (320 species or 43 subspecies) are accepted within the family Alycaeidae. Of these, 22 have been described by the lead author and his coauthors in previous publications. In addition, there are 18 species that were formerly classified in Cycloryx and now belong to Pincerna due to its synonymy with Cycloryx. Among the remaining 323 species, 209 (65%) are transferred here to another genus, whilst 114 (35%) have remained in their original genus.Seven genera are accepted. While some questions (e.g., the distinction between Pincerna and Alycaeus) remained unanswered, this revision made three main achievements: (1) The Dicharax species were identified based on the absence of spiral striation on the entire shell; (2) the Metalycaeus species were identified based on the spiral striation of the protoconch; (3) and Stomacosmethis was separated from Alycaeus based on the extremely short sutural tube.Five nominal species are being synonymised with other species, and eight species are now treated as subspecies. The following replacement names are proposed: Dioryx urnula niosiensis Páll-Gergely, nom. nov. for Alycaeus urnula var. daflaensis Godwin-Austen, 1914; Dioryx urnula rotundus Páll-Gergely, nom. nov. for Alycaeus urnula var. globosus Godwin-Austen, 1914; Pincerna crenilabris juttingae Páll-Gergely, nom. nov. for Alycaeus crenilabris laevis van Benthem Jutting, 1959; Pincerna crenilabris korintjiensis Páll-Gergely, nom. nov. for Alycaeus crenilabris latecostatus van Benthem Jutting, 1959; Dicharax conicus jatingaensis Páll-Gergely, nom. nov. for Alycaeus conicus var. nanus Godwin-Austen, 1914; Metalycaeus godwinausteni Páll-Gergely, nom. nov. for Alycaeus neglectus Godwin-Austen, 1914; and finally Metalycaeus suhajdai Páll-Gergely, nom. nov. for Alycaeus varius Godwin-Austen, 1914.

Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae).

The genus Peronia Fleming, 1822 includes all the onchidiid slugs with dorsal gills. Its taxonomy is revised for the first time based on a large collection of fresh material from the entire Indo-West Pacific, from South Africa to Hawaii. Nine species are supported by mitochondrial (COI and 16S) and nuclear (ITS2 and 28S) sequences as well as comparative anatomy. All types available were examined and the nomenclatural status of each existing name in the genus is addressed. Of 31 Peronia species-group names available, 27 are regarded as invalid (twenty-one synonyms, sixteen of which are new, five nomina dubia, and one homonym), and four as valid: Peronia peronii (Cuvier, 1804), Peronia verruculata (Cuvier, 1830), Peronia platei (Hoffmann, 1928), and Peronia madagascariensis (Labbé, 1934a). Five new species names are created: P. griffithsi Dayrat & Goulding, sp. nov., P. okinawensis Dayrat & Goulding, sp. nov., P. setoensis Dayrat & Goulding, sp. nov., P. sydneyensis Dayrat & Goulding, sp. nov., and P. willani Dayrat & Goulding, sp. nov.Peronia species are cryptic externally but can be distinguished using internal characters, with the exception of P. platei and P. setoensis. The anatomy of most species is described in detail here for the first time. All the secondary literature is commented on and historical specimens from museum collections were also examined to better establish species distributions. The genus Peronia includes two species that are widespread across the Indo-West Pacific (P. verruculata and P. peronii) as well as endemic species: P. okinawensis and P. setoensis are endemic to Japan, and P. willani is endemic to Northern Territory, Australia. Many new geographical records are provided, as well as a key to the species using morphological traits.

New replacement name for a homonym in Lepeophtheirus (Crustacea: Copepoda: Caligidae).

Markevich (1940) established Pseudolepeophtheirus Markevich, 1940 for Pseudolepeophtheirus longicauda Markevich, 1940 based on copepods collected from the pleuronectid fish Platichthys stellatus (Pallas, 1787). Dojiri Ho (2013) synonymized the genus and the species with Lepeophtheirus Nordmann, 1832 and Lepeophtheirus parvicruris Fraser, 1920, respectively. Later, Homma et al. (2020) resurrected Markevich's species as a member of Lepeophtheirus, i.e., as L. longicauda (Markevich, 1940). The last component of the names of both genera is 'phtheirus' (transliterated from the Greek φθειρ; Nordmann 1832: 30), a masculine noun, and thus under Article 30.1.2 of the International Code of Zoological Nomenclature (hereinafter, Code; International Commission on Zoological Nomenclature 1999), both generic names are also masculine. The species-group name longicauda might be regarded as either a noun in apposition or as an adjective in the feminine gender, and Markevich (1940) did not specify his intention in this regard. Bearing in mind that 'cauda', meaning 'tail', actually is a feminine Latin noun and that Markevich did not change the final '-a' to '-us' to match the masculine gender of the genus, we deem that longicauda Markevich, 1940 is a noun in apposition, a position supported by Article 31.2.2 of the Code.

Key to South American genera of Acanthocinini (Coleoptera, Cerambycidae, Lamiinae) with erect setae on elytral surface.

A key for identification of the 45 genera of Acanthocinini with erect setae on the elytra and which occur in South America is provided. A new synonymy is proposed for Trichonyssodrys Gilmour, 1957 (junior synonym of Pentheochaetes Melzer, 1932), resulting in new combinations for the following species: Pentheochaetes aureopilosa (Monné, 1990), P. cincta (Delfino, 1981), P. maculata (Gilmour, 1957), P. melasma (Delfino, 1981), and P. nessimiani (Monné Monné, 2012). The gender of the species-group names in Pentheochaetes is corrected. Diagnosis for each genus is provided, as well as type-locality and geographical distribution of the type-species.

Elrathia hensonensis nomen novum, new replacement name for Elrathia groenlandica Geyer and Peel, 2017 (Trilobita, Ptychopariacea)

In an article by Geyer and Peel (2017, p. 288), we inadvertently coined a species-group name, Elrathia groenlandica Geyer and Peel, 2017 for a trilobite species of the superfamily Ptychopariacea. This name is a junior homonym of Elrathia? groenlandica Poulsen, 1927 and is thus invalid (ICZN Article 57.2; see International Commission on Zoological Nomenclature, 1999). We therefore propose Elrathia hensonensis new name as the replacement name for E. groenlandica Geyer and Peel, 2017. The new species name is derived from the Henson Gletscher area, North Greenland, the region where the type locality of this species is located.

Every sponge its own name: removing Porifera homonyms.

The occurrence of different sponge species bearing the same Linnean binomial name combination, i.e. homonyms, is to be avoided for obvious reasons. In a review of sponge taxon names of the World Porifera Database, we detected 121 homonymic cases (115 species-group names, 6 genus-group names), involving a total of 272 nominal taxa. It is the object of the present study to remove their occurrence by proposing new names for the junior homonyms following the rules of the International Commission of Zoological Nomenclature as laid down in the Code (ICZN, 1999) and the on-line edition http://iczn.org/iczn/index.jsp . Homonym cases are discussed and, where applicable, junior homonyms are either replaced by nomina nova or reassigned to their earliest available synonyms. The order in which the homonyms are treated is alphabetical on original species name, with genus names separately treated at the end. A summary table with all proposed name changes is also presented to allow quick access to the junior homonyms and their proposed new names. A total of 116 nomina nova are proposed, including five new genus names.

The world woodlouse flies (Diptera, Rhinophoridae).

The world Rhinophoridae are catalogued, recognising 33 genera and 177 species. Nomenclatural information is provided for all genus-group and species-group names, including lists of synonyms and name-bearing type data. Species distributions are recorded by country. A key to the world genera is presented. Four new genera are erected to accommodate five new species, which do not fit within any of the current generic concepts in Rhinophoridae, according to the results of a morphology-based phylogenetic analysis: Marshallicona Cerretti & Pape with type species Marshallicona quitu Cerretti & Pape, gen. et sp. nov. (Ecuador); Maurhinophora Cerretti & Pape with type species Maurhinophora indoceanica Cerretti & Pape, gen. et sp. nov. (Mauritius); Neotarsina Cerretti & Pape with type species Neotarsina caraibica Cerretti & Pape, gen. et sp. nov. (Trinidad and Tobago) and Neotarsina andina Cerretti & Pape, sp. nov. (Peru); Kinabalumyia Cerretti & Pape with type species Kinabalumyia pinax Cerretti & Pape, gen. et sp. nov. (Malaysia, Sabah). The genus Aporeomyia Pape & Shima (type species Aporeomyia antennalis Pape & Shima), originally assigned to Tachinidae, is here reassigned to Rhinophoridae based on a reassessment of the homologies of the male terminalia. The following five species-group names, which were previously treated as junior synonyms or nomina dubia, are recognised as valid species names: Acompomintho caucasica (Villeneuve, 1908), stat. rev. [from nomen dubium to valid species]; Acompomintho sinensis (Villeneuve, 1936), stat. rev. [from nomen dubium to valid species]; Stevenia bertei (Rondani, 1865), stat. rev. [from nomen dubium to valid species]; Stevenia sardoa Villeneuve, 1920, stat. rev. [from junior synonym of Rhinophora deceptoria Loew, 1847 to valid species]; Stevenia subalbida (Villeneuve, 1911), stat. rev. [from junior synonym of Rhinophora deceptoria Loew, 1847 to valid species]. Reversal of precedence is invoked for the following case of subjective synonymy to promote stability in nomenclature: Rhinophora lepida (Meigen, 1824), nomen protectum, and Musca parcus Harris, 1780: 144, nomen oblitum. New generic and specific synonymies are proposed for the following two names: Mimodexia Rohdendorf, 1935, junior synonym of Tromodesia Rondani, 1856, syn. nov. and Ptilocheta tacchetti Rondani, 1865, junior synonym of Stevenia obscuripennis (Loew, 1847), syn. nov. The following new combinations are proposed: Acompomintho sinensis (Villeneuve, 1936), comb. nov. [transferred from Tricogena Robineau-Desvoidy, 1830]; Tromodesia guzari (Rohdendorf, 1935), comb. nov. [transferred from Mimodexia Rohdendorf, 1935]; Tromodesia intermedia (Rohdendorf, 1935), comb. nov. [transferred from Mimodexia Rohdendorf, 1935]; Tromodesia lindneriana (Rohdendorf, 1961), comb. nov. [transferred from Mimodexia Rohdendorf, 1935]; Tromodesia magnifica (Rohdendorf, 1935), comb. nov. [transferred from Mimodexia Rohdendorf, 1935]; Tromodesia obscurior (Rohdendorf, 1935), comb. nov. [transferred from Mimodexia Rohdendorf, 1935]; Tromodesia pallidissima (Rohdendorf, 1935), comb. nov. [transferred from Mimodexia Rohdendorf, 1935]; Tromodesia setiventris (Rohdendorf, 1935), comb. nov. [transferred from Mimodexia Rohdendorf, 1935] and Tromodesia shachrudi (Rohdendorf, 1935), comb. nov. [transferred from Mimodexia Rohdendorf, 1935].