Although Butler (1907) discovered more than thirty years ago that species of Pythium could be isolated from the soil, it was not until the application of Butler's cultural methods by Harvey (1925) that the widespread occurrence of the Saprolegniaceae and other water molds in the soil was generally appreciated. Subsequently, the phycomycetous soil flora has been studied in various parts of the world. Coker and Braxton (1926), Coker (1927), and Raper (1928) have made a series of studies of terrestrial forms of the Saprolegniaceae and closely related groups in North Carolina, Couch (1927) has isolated aquatic fungi from soils in New York, and Harvey (1927, 1928, 1930) has investigated the water molds occurring in Wisconsin soils, as well as those of Oklahoma, Mississippi, and Kentucky. The studies of Apinis (1930) in Latvia, Dissmann (1931) in Austria, Barnes and Melville (1932) in England, Cook and Morgan (1934) and Morgan (1938) in Wales, Hohnk (1935) and Richter (1937) in Germany, Nagai (1931) in Japan, and Cookson (1937) in Australia have shown that saprolegniaceous fungi as well as other groups of the aquatic Phycomycetes are widely distributed in soils throughout various parts of the world. These investigations have further demonstrated that while certain species previously found only in water are also quite common in the soil, other forms, including species of the genera Brevilegnia and Geolegnia, are apparently adapted for a strictly terrestrial mode of existence, and are to be found only as constituents of the soil microflora. H6hnk (1935) has clearly correlated the number of motile stages of the zoospores in various genera of the Saproleg-
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