The research results of the problem of definition of the principles of the ecological competence formation of the future biology teachers during the professional training have been defined in the article. It has been specified, that the conception of the ecological competence formation of the future biology teachers is based on general pedagogical and specific principles that are normative guidelines, basic foundations for organization and performing the professional training of students. We have highlighted the fact, that the effectiveness of the implementation of the system of the students' ecological competence is achieved due to clear compliance with the chosen principles and approaches to the process of the future teachers' training. In this article the principles of the ecological competence formation of the future biology teachers are considered as the most common starting points that define the strategic ways and tactic for the professional training of higher-education students of the qualification mentioned above in order to form the studied quality of personality. The author has defined general pedagogical (principles of science, regularity and sequence, unity of consciousness, activity and self-discipline, connection of studying with practical activity and life's realities) and specific principles of ecological competence formation (principles of interdisciplinary integration, the creation of the ecologically friendly educational environment, professional orientation, variability, the relationship of local, emotional values of perception of the natural environment in practice, regional and global approaches in the educational process). It has been pointed out, that the specific principles of the ecological competence formation of the future biology teachers have been defined by the author based on the synergetic connection of the principles of the professional education, competence approach, the principles of ecological education and upbringing, which have points of intersection for individual positions. We have highlighted the fact, that these principles are used during the professional training of the future biology teachers as interconnected and complementary elements of a single pedagogical system.