Open or closed tone holes are usually represented as shunt impedances in the air column transmission line equations. Nederveen and van Wulfften Pathe (1963) pointed out that the lumped circuit for a closed tone hole includes a negative inertance in series with the shunt compliance. The open and closed tone hole impedance parameters have been calculated as a function of frequency for various tone hole proportions using variational method, and experimentally confirmed in detail. Large diameter short chimney holes (as on the sax) behave differently from small diameter long chimney holes (e.g., clarinet). Additional viscous and thermal losses near the sax-type open holes lessen their otherwise higher radiation efficiency. A single sax-type open hole at a pressure node radiates as a dipole while clarinet-type holes do not, since the evanescent modes are more weakly generated at the main bore-tone hole junction and more rapidly attenuated in the long chimney. The series impedance for an open hole, like the closed hole, is a negative inertance larger in magnitude than for the closed hole. The behavior of an open hole lattice near its cutoff frequency differs for the two types of holes. Adjacent open or closed sax holes interact within the main bore, while clarinet holes do not. External interaction via radiation from adjacent open holes raises the shunt inertance by a few per cent. Internal and external hole interactions modify overall instrument tuning and radiation efficiency. [Work supported by NSF.]