Shell Beds Research Articles

Palaeoecological Analysis of Benthic Molluscs from the Eocene of Kutch, Gujarat Reveals an Event of Storm Induced Concentration of Shells in a Quiet Marginal Marine Environment

The Eocene deposits of the Kutch basin are characterized by predominance of plane-laminated shales and low diversity mollusc faunas. These features reflect a low energy, restricted environment. A thin molluscan shell bed stands out in this backdrop because it produced a very diverse assemblage of bivalves and gastropods. Palaeosynecological analysis of the benthic mollusc assemblages from different stratigraphic levels of the Eocene of Kutch, including the shell bed, was carried out by testing for standard ecological models of rank-abundance. The specific abundance data from the shell bed c onform to the log-normal model indicating stable, resource-rich environment. On the other hand, low diversity and often high abundance of opportunist species from other levels reflect harsh ecological conditions for molluscs. Analyses of habitat preferences and habits of the genera from the shell bed indicate shallow to very shallow shelf environment. It is inferred that this exotic assemblage of shells was brought in this quiet, restricted environment by storm surges from open part of the sea. High degree of shell fragmentation and high content of micrite matrix corroborate this interpretation.

Ecological snapshot of a population of Panopea within their traces (Pliocene, Agua Amarga subbasin, SE Spain)

Pliocene deposits cropping out in the Agua Amarga subbasin (Almería, SE Spain) include a composite shell bed made up of variously preserved and densely packed mollusks. The characteristics of the shell bed indicate deposition in shallow marine settings under a changeable sedimentation rate. The composite shell bed was formed through the amalgamation of several depositional events, mostly connected with storm events. During relatively slow sedimentation the bivalve Panopea colonized the substratum. The colonization took place over different stages of the shell bed formation, as reflected by differently preserved Panopea scattered throughout the shell bed. The upper part of the shell bed contains several tens of Panopea preserved within their burrows: Scalichnus cf. phiale Hanken et al., 2001. All bivalves in S. cf. phiale are articulated and preserved in life position. They are confined to a single horizon and most probably represent a single fossil population composed of adult individuals of Panopea resulting from anastrophic burial by storm deposits. As a consequence, the bivalves in their burrows succumbed at the same time; that is, the study case represents an “ecological snapshot” or ecological census.

Removal of Chromium from Aqueous Solution using Multilayer-Mixed (Reduction/Sorption) Bed from Iron Scrap and Nutshells

In this research removal of chromium from an aqueous solution by multilayer-mixed (reduction/sorption) bed via scrap iron as zero valent iron and charred nutshell (walnut shell) as adsorber were studied. Both batch and continuous system were conducted. The batch system reveals that the adsorption process was of favorable type. Three models were used to describe the adsorption process. Freundlich model is matching well to the data with (R2=0.975). In the continuous system , two sets of experimental were considered, the first set (multilayer bed) conducted by fixing the characteristics of the scrap iron bed (bed height 0.2 cm, particle size 500 µm) and varying the walnut shell parameters, the optimum value of this set recorded as (bed height 2.5 cm, particle size250µm). While for the second set of experimental after fixing the optimum values of the walnut shell bed and varying the scrap iron bed characteristics, the optimum values were (height 1cm, particle size 500µm as filling). Also, the Mixed bed configuration were studied by conducting the optimum conditions for the previous experiment. it was found that at any certain time the mixed layer offers the maximum efficiency of removal rather than the multi-layer bed.

Gigantoproductid and allied productid brachiopods from the “Calcaires à Productus” (late Viséan-Serpukhovian; Montagne Noire, southern France): Taxonomy and palaeobiogeographical position in the Palaeotethys

The “Calcaires à Productus” or “Calcaires à Productus giganteus” on the southern slope of the Montagne Noire (Hérault, France) consist of numerous isolated olistolitic limestone blocks, generally transported by gravitational flows into siliciclastic basinal deposits. The present paper is the first systematic monographic study of gigantoproductids, semiplanids, and some other large-sized productid brachiopods from the upper Viséan to Serpukhovian strata of the Montagne Noire. Gigantoproductids are represented by abundant Datangiasemiglobosa and Kansuella spp.; a single Globosoproductus specimen is described. The semiplanids are represented by Latiproductus aff. latissimus, L. latissimus complicatus and L. edelburgensis, and the linoproductids by Balakhonia aff. kokdscharensis. The Montagne Noire productid faunas are found in very different facies including shell beds and microbial mounds. They are most abundant in assemblages from agitated, shallow water environments with a firm substrate where rugose corals are rare or absent. The palaeobiogeographical relations to the other regions in the western Palaeotethys are often limited to taxa with an almost cosmopolitan character. Worth noting is the absence of Gigantoproductussensu stricto in the Montagne Noire. This genus, and other brachiopod taxa, migrated probably during the late Serpukhovian into southern France. On a larger scale, the presence of Kansuella indicates faunal exchanges with the eastern Palaeotethys (South China) via a westward migration path along North China, the Central Asian Orogenic Belt and south-eastern Laurussia. The studied taxa, even at the genus level, did not migrate into North America even if the Alleghanian Isthmus was still open.

Microbialite fields developed in a protected rocky coastline: The shallow carbonate ramp of the Aptian Romualdo Formation (Araripe Basin, NE Brazil)

The Aptian Romualdo Formation is a siliciclastic-dominated sedimentary unit recording the last marine ingression within the Cretaceous interior basins of northeastern Brazil. At the western margin of the Araripe Basin, rocks of the Romualdo Formation are mainly represented by carbonates resting abruptly over the Proterozoic crystalline basement. Detailed mapping and stratigraphic analysis revealed two stromatolite fields that were described and discussed for the first time. Several bioherms, biostromes and isolated stromatolites characterized by distinct microbialite morphologies associated with echinoid-rich strata have been identified, suggesting that hypersalinity, water depth and hydraulic conditions were the main factors controlling stromatolite morphogenesis. A cm-thick amalgamated bivalve rudstone, resting directly on the basement and representing a shell concentration formed above the fair-weather wave base was also recorded. Based on the regional distribution of the stromatolite types and associated sedimentary facies, we interpret the depositional environment as a local low-gradient carbonate ramp deepening to the east. Our data robustly indicated that the western rocky shorelines of the Araripe Basin during the Aptian were populated by microbial mats and stromatolites in a condition analogous to the modern world-famous Hamelin Pool, Shark Bay, Western Australia. Finally, the development of stromatolites and echinoid-bearing microbialites at the western margin of the basin may be correlated with the formation of bakevelliid- and cassiopid-rich shell beds in the upper part of the Romualdo Formation at the eastern margin.

Underwater Shell Middens: Excavation and Remote Sensing of a Submerged Mesolithic site at Hjarnø, Denmark

Shell middens, or shell-matrix deposits, occur in large numbers across the coastlines of the world from the mid-Holocene (ca. 6000–5000 cal BC) onwards, often forming substantial mounds. However, they become smaller, rarer or absent as one goes back into earlier periods, suggesting a world-wide process of economic intensification. Since sea level was generally much lower during these earlier periods, a critical question is the extent to which mounded shell middens could have accumulated on now-submerged palaeoshorelines, and if so, how they were affected by the potentially destructive impact of sea-level rise. Further, and important to modern practice, it is essential that archaeologists consider how such sites can be discovered through underwater investigation. Here we offer a proof of concept that shell middens can survive submergence and can be detected, using systematic investigation of a rare example of a confirmed underwater shell midden at the Mesolithic site of Hjarnø (ca. 5300–4300 cal BC) in Denmark. We compare the excavation results with the results of geophysical survey, explore the problems of distinguishing underwater cultural shell middens from natural shell beds and conclude that shell middens can survive inundation by sea-level rise and can be detected by remote sensing, but require at least minimally invasive sampling to establish their cultural status. We suggest the methods developed may be applicable to coastal and marine sites impacted by postglacial sea-level rise worldwide.

A first report of microtektites from the shell beds of southwestern Florida

AbstractThe Plio‐Pleistocene Upper Tamiami Formation (Pinecrest beds) of Florida is well known for its fossiliferous shell beds, but not for its extraterrestrial material. Here we report the first occurrence of tiny (~200 μm in diameter) silica‐rich microspherules from this unit and from the state. This material was analyzed using petrographic and elemental methods using energy dispersive X‐ray spectroscopy (EDS). The majority of microspherules are glassy and translucent in reflected light with some displaying “contact pairs” (equal‐sized micro‐spherules attached to each other). Broken microspherules cleave conchoidally, often with small internal spherical vesicles, but most lack any other evidence of internal features, such as layering. Using the EDS data, the microspherules were compared to volcanic rocks, microtektites, and cosmic spherules (micrometeorites). Based on their physical characteristics and elemental compositions these are likely microtektites or a closely related type of material. The high Na content in the examined material deviates significantly from the abundances usually found in micrometeorites and tektite material; this is enigmatic and requires further study. This material may be derived from a nearby previously unknown impact event; however, more material and sites are required to confirm the source of this material. Because of the focus on molluscan fossils in southwestern Florida shell beds, microtektite material has likely been overlooked in the past, and it is probable that these microspherules are in abundance elsewhere in these units and possibly throughout the region.

A holoplanktic gastropod in a raised reef: Hopegate Formation, Jamaica (upper Pliocene)

Of the two highly fossiliferous but contrasting sedimentary successions in the upper Pliocene of Jamaica, only the fossils of the Bowden shell beds have received adequate study. The Hopegate Formation of the island’s central north coast is a thick and areally extensive unit, a raised reef rich in fossil scleractinian corals and benthic molluscs. However, it is well lithified and in part dolomitised, so it is difficult to collect the fossils, which are commonly indifferently preserved. To the fauna of the Hopegate Formation is added its first holoplanktic gastropod, Diacria trispinosa (de Blainville), a unique specimen preserved as an internal mould. All other molluscs from this formation are benthic. However, the presence of D. trispinosa is unexpected, as it likely spent most of its life in the deeper water zooplankton.

Distinguishing core and flank facies based on shell fabrics in Lower Jurassic lithiotid shell beds

Lower Jurassic larger bivalves, mostly represented by the monospecific lithiotid genera Lithiotis, Cochlearites and Lithioperna, formed large shell accumulations in the shallow-water carbonate Tethyan and Panthalassa margins. A quantitative analysis of lithiotid accumulations from the Trento Platform (northern Italy) was carried out in order to distinguish the distribution of autochthonous, parautochthonous and allochthonous lithiotids in the accumulation core and flanks. Various representative accumulations are characterized with respect to systematic content, shell cover, shell density, disarticulation, shell fragmentation and orientation of individual shells. Lithioperna shows a high shell cover within the accumulation core while that of Cochlearites is variable and Lithiotis is rare. All three lithiotids occur in high densities in the accumulation flanks. As expected, autochthonous individuals of all three genera are frequent in the accumulation cores where they are preserved in life position. In the flanks, most bivalves are parautochthonous except for Lithioperna which can grow in life position. All bivalves in allochthonous shell beds are highly fragmented and disarticulated. The studied traits are potentially useful as proxies for distinguishing core or flank facies when the accumulation does not crop out as a whole.

High‐frequency cycles of brachiopod shell beds on subaqueous delta‐scale clinoforms (early Pliocene, south‐east Spain)

AbstractDuring the early Pliocene, subaqueous delta‐scale clinoforms developed in the Águilas Basin, in a mixed temperate carbonate–siliciclastic system. The facies distribution is consistent with the infralittoral prograding wedge model. Stacking patterns and bounding surfaces indicate that the clinoforms formed during the highstand and falling sea‐level stages of a high rank cycle. Twenty‐two prograding clinothems were recognized over a distance of ≥1 km. Biostratigraphic data indicate a time span shorter than 700 kyr for the whole unit (MPl3 biozone of the Mediterranean Pliocene). Cyclic skeletal concentrations and occasional biostromes of suspension feeders (terebratulid brachiopods, modiolid bivalves and adeoniform bryozoan colonies), slightly evolved glauconite and occasionalGlossifungitesichnofacies formed on the clinoforms during high‐frequency pulses of relative sea‐level rise. During such stages, increased accommodation space in the topsets of the clinoforms caused a strong reduction of terrigenous input into the foresets and bottomsets. This provided favourable conditions for the development of these suspension feeder palaeocommunities. During stillstand stages, however, reduced accommodation space in the topsets eventually resumed progradation in the foresets. There, the abundance ofDitrupatubes indicates frequent siltation events that extirpated the terebratulid populations and other epifaunal suspension feeders in the foreset and bottomset subenvironments. The occurrence of shell beds on the clinoforms suggests that this case study represents lower progradation rates than standard examples where shell beds bound the clinobedded units at their base and top only. Importantly, the distributions of biofacies and ichnoassemblage associations contribute significantly to the understanding of the effects of relative sea‐level fluctuations on the evolution of subaqueous delta‐scale clinoform systems.

Global geoheritage significance of Ordovician stratigraphy and sedimentology in the Cliefden Caves area, central western New South Wales

Globally significant geoheritage features of the Cliefden Caves area, in the Belubula River Valley between Orange and Cowra in central western New South Wales, comprise a richly fossiliferous shallow-water limestone succession of Late Ordovician age (the Cliefden Caves Limestone Subgroup) overlain by deep-water laminites and allochthonous limestones of the Upper Ordovician Malongulli Formation. Key features of the Ordovician geology of the Cliefden Caves area that have been identified using the Geoheritage Toolkit as being of international significance are the abundance of unique and exceptionally diverse fossils in the Fossil Hill Limestone (forming the lower part of the Cliefden Caves Limestone Subgroup), which supplement detailed interpretation of carbonate-dominated deposition within an Ordovician volcanic island setting. The fossiliferous limestones preserve biostromes and local small bioherms of stromatoporoids and corals, and recurrent in situ and disarticulated/imbricated Eodinobolus shell beds formed in shallow, quiet-water, dominantly muddy carbonate sediments that passed up-sequence to clay-free carbonate environments. These mud-dominated carbonate sediments are interspersed with higher-energy conditions, represented by skeletal, lithoclastic and calcrete-ooid grainstones overlying disconformities, leading to the identification of subaerial disconformities and associated diagenesis in the Fossil Hill Limestone. The Fossil Hill Limestone is succeeded by massive limestones in the middle part of the Cliefden Caves Limestone Subgroup and then, in turn by the Vandon Limestone and the deeper-water graptolitic laminites of the Malongulli Formation—this completes a succession that is rarely preserved in the geological record, further enhancing the geoheritage significance of the Cliefden Caves area.

Ordovician strata in the Cliefden Caves area, New South Wales: a case study in the preservation of a globally significant paleontological site

The Cliefden Caves area in central-western New South Wales includes the scientifically most important and irreplaceable examples of fossiliferous Ordovician rocks in the State. Exposures of the stratigraphically lower parts of the Cliefden Caves Limestone Subgroup on the aptly named Fossil Hill are world-famous among paleontologists and internationally significant for preserving the earliest in situ shell beds documented in the literature. They also contain some of the oldest known rugose corals, and an exceptional example of one of the oldest coralline biostromes, as well as many examples of invertebrate fossils and cyanobacterial mat structures that either are unique to this locality or were first described from here. Other stratigraphic levels throughout the total 363 m-thick Cliefden Caves Limestone Subgroup are similarly endowed with highly significant fossils, such as a globally unique in situ shell bank with rare examples of the trimerellide brachiopod Belubula spectacula, a wealth of shelly fossils and trilobites on Dunhill Bluff (adjacent to Fossil Hill to the east), and the appropriately named Trilobite Hill. Less well known to the general public, but of international importance to paleontologists, is the unique deep-water sponge fauna of the overlying Malongulli Formation that occurs at several levels in limestone lenses within this unit. Fossils from the Cliefden Caves Limestone Subgroup and the Malongulli Formation have been documented in more than 60 scientific papers and monographs since paleontological investigations into the site were first published in 1895. Despite concerted scientific endeavour in the region over the past 50 years, much more study needs to be done to fully document the paleontological riches of the Cliefden Caves area. These sites are interpreted as the remains of a tropical island, fringed by limestone and flanked by deep-water environments in which the Malongulli Formation was deposited. Preservation of such islands is exceptionally rare in the geological record. It is therefore vital for the area to remain accessible to scientific researchers to continue their studies. Flooding of the Belubula Valley by a proposed dam downstream from the Cliefden Caves area would hinder future research work on this unique geoheritage resource. Fortunately, a successful public campaign has led to listing of the site on the State Heritage Register that will provide essential protection of the caves from inundation while ensuring continued access to researchers.

Remarkable preservation of shell microstructures from the Late Ordovician of the Cincinnati Arch region, USA, and the success of nacre among Ordovician mollusks

AbstractExceptional sub-micrometer details of shell microstructure are preserved in phosphatic micro-steinkerns representing several phyla from shell beds of the Upper Ordovician of the Cincinnati Arch region, USA. These fossils provide the most detailed record of Ordovician mollusk shell microstructures, as well as exceptional details on the earliest cases of undisputed nacre. The trend towards nacre in the Mollusca is one aspect of the surge in escalation between mollusks and their predators during the Great Ordovician Biodiversification Event.

Taphonomy and Sedimentological Significance of Oyster Shell Beds within Cretaceous Transgressive Sediments in Japan

Through taphonomical analyses of Crassostrea shell beds included in several Japanese Cretaceous strata, paleoecological and sedimentological significance of intertidal muddy and sandy bottom dwelling oysters are examined. The Upper Cretaceous Crassostrea shell beds show a variety of modes of occurrence, lithological changes and sedimentary structure, suggesting composite formative, paleoecological and sedimentary processes. Crassostrea seems to have adapted from muddy substratum during early Cretaceous to coarser sandier substratum often influenced by physical turbulence such as tidal currents and waves during the mid-Cretaceous time, associated with increasing in shell size and thickness, and colony dimension.

AN IN SITU PRESERVED EARLY CARBONIFEROUS (SERPUKHOVIAN) BRACHIOPOD COMMUNITY IN SOUTHERN GUIZHOU, CHINA

A brachiopod shell bed from the Lower Carboniferous (Serpukhovian) in Guizhou Province of southern China is reported as representing an in situ preserved brachiopod community. The community is characterized by yielding more than 80% complete and articulated specimens preserved in life position and poor size sorting. A new spiriferide species, Weiningia ziyunensis n. sp., is described in the community, which contains the other eight species belonging to six genera. Morphology and preservation analysis of Weiningia ziyunensis n. sp. suggests that it lived in dense clusters attached to living and dead shells and stabilizing its position with thickened posterior shell. Size-frequency distribution and survivorship curve are applied to the population dynamics investigation of this species. Large number of juvenile shells accompanied by high mortality reflects that many juvenile individuals suffered from the limited life space and turbid environment generated by dense clusters. The same high adult mortality is the result of more pressure from neighbors that lead to shell malformation and eventually low feeding and cleaning efficiency, whereas the low senior mortality is attributed to their abilities to cope with these problems. Members of the community show great difference in numeric frequency, with Weiningia ziyunensis n. sp. being one of the dominant species that was characterized by crowded life strategy. By living in dense clusters on Striatifera striata (Fischer de Waldheim, 1837) or other shell fragments, Weiningia ziyunensis n. sp. could resist the water current and gradually expand its population.

PERFORMANCE AND SHORT TERM DURABILITY OF PALM KERNEL SHELL AS A SUBSTRATE MATERIAL IN A PILOT HORIZONTAL SUBSURFACE FLOW CONSTRUCTED WETLAND TREATING SLAUGHTERHOUSE WASTEWATER

Wastewater treatment using constructed wetlands is one of the effective and low-cost technologies to improve the quality of slaughterhouse effluent. This study was carried out to investigate the suitability of palm kernel shell as a substrate material for constructed wetlands treating slaughterhouse wastewater. Rhizomes of Thalia Geniculata and Typha Latifolia were grown in four pilot horizontal subsurface flow constructed wetland beds filled with palm kernel shell and grave, and their growth and treatment performance evaluated. The results of the study showed that Thalia Geniculata survives and proliferates in palm kernel shell bed. The mean removal rates of 72.81% (BOD5), 89.87% (TSS), 39.42% (NH4-N), 60.79% (NO3-N) and 42.52% (PO43-) for the palm kernel shell were comparable to the values obtained for the gravel bed. The study proved that palm kernel shell, as a substrate material in constructed wetlands had the potentials to sustain the growth of some macrophytes, as well as the capacity to remove contaminants from wastewater.

Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)

ABSTRACTShell beds from the Caballas Formation in the East Pisco Basin of southern Peru have yielded an assemblage of molluscs numerically dominated by oysters and cerithioid and eulimid gastropods, with lesser numbers of neritid, ampullinid, and pyramidelloid gastropods and noetiid, mytilid, anomiid, and cardiid bivalves. The shell beds mark a transition from continental to marine-influenced environments. Palaeontological evidence, including plant twig moulds on the attachment scars of oysters and a high diversity of potamidid gastropods, is consistent with an interpretation of a brackish-water setting. The majority of molluscan species from the Caballas Formation show a close affinity with Late Cretaceous species from northern Peru, but no unequivocally Cretaceous invertebrate taxa (e.g. ammonites, rudists) have been found. A morphologically distinctive bivalve species that occurs in the Caballas Formation, Carolia (Parinomya) parinensis, only occurs elsewhere in the upper lower Eocene Pariñas and Chacra formations of the Talara Basin of northern Peru. Moreover, several other Caballas Formation molluscan taxa closely resemble early Eocene species from the Talara Basin. These equivocal biostratigraphic data indicate an early Paleogene age as most likely for the Caballas Formation.http://www.zoobnk.org/urn:lsid:zoobank.org:author:25E21A31-0230-48C7-A2ED-A48A98A3F231

Vendian Macrofossils of the Yudoma Group, Southeast of the Siberian Platform

An assemblage of Late Precambrian macrofossils consisting of five species (Aspidella terranovica, Beltanelliformis brunsae, Nenoxites curvus, Palaeopascichnus renarius, and Suvorovella aldanica), was discovered in the type sections of the Yudoma Group of the Siberian Platform. A unique feature of the assemblage is the presence of carbonate discoidal remains of Suvorovella, formed as a result of secondary incrustation of basal structures, similar to Aspidella. The accumulations of redeposited fragments of Suvorovella formed the rock resembling the Phanerozoic shell beds. Except for this taxon, all members of the Yudoma assemblage have a wide geographic distribution. Their stratigraphic distribution tends to occur in the youngest deposits of the Upper Vendian, directly underlying the beds with small shelly fauna (SSF) of the Nemakit-Daldynian Stage, Cloudina tubes, and traces of the Treptichnus pedum Zone.

Analysis of a Cretaceous (late Aptian) high-stress ecosystem: The Romualdo Formation of the Araripe Basin, northeastern Brazil

The Cretaceous (upper Aptian) Romualdo Formation of the Araripe Basin, northeastern Brazil, is a marine-influenced, predominantly fine-grained siliciclastic unit. In the eastern part of the basin, it corresponds to a depositional sequence composed of two transgressive-regressive cycles. Low-diversity benthic macrofaunas document high-stress conditions. Their palaeoecological and taphonomic analysis allows for the detailed reconstruction of the palaeoenvironments and of the temporal changes in the palaeoecosystem. The transgressive part of the lower cycle is a black shale with calcareous concretions that contain excellently preserved fishes and other vertebrates (the famous “Santana fossils”), but apart from abundant ostracods and rare decapod crustaceans, benthic faunal elements are very rare due to anoxic to dysoxic conditions. A low-diversity parautochthonous molluscan fauna of small bivalves and gastropods records fluctuating dysoxic conditions during early regression. Towards the top of the measured sections (late regression), thin gastropod shell beds become conspicuous. At the top of the sections decimetre-thick shell concentrations show signs of reworking and a greater marine influence (transgressive part of the upper cycle). The distribution pattern of three benthic associations and two assemblages is almost equally shaped by three environmental parameters: The low overall species diversity, small size of the fauna, and the lack of stenohaline elements suggest lowered and fluctuating salinity conditions. Fluctuating redox conditions with anoxia to dysoxia, identified especially in the lower parts of the succession, and substrate were the other two factors shaping the faunal distribution. The late Aptian sedimentary and faunal history of the eastern part of the basin indicates restricted marine influence, coupled with increasing freshwater input during phases of sea-level lowstand, and low-energy conditions offshore, which led to widespread anoxia, especially in the lower part of the formation. Thus, palaeoecological analyses are an excellent tool to disentangle the various environmental parameters that shaped the palaeoenvironments of the basin, to understand more closely its Fossil-Lagerstätten, and to corroborate and fine-tune the sequence stratigraphic framework.

Eocene bryozoan assemblages from the La Meseta Formation of Seymour Island, Antarctica

Early to Late Eocene bryozoans from the La Meseta Formation of Seymour Island were collected at two localities within the Cucullaea I Allomember (Telm4 and Telm5) on the northwestern side of the island and in two localities within the Submeseta Allomember (Telm6 and Telm7) on the northeastern side. This fauna is represented by cyclostomes of the suborders Tubuliporina and Cerioporina and suborders of Neocheilostomata, among which nine species have been recognized. The following new species are introduced: Micropora nordenskjoeldi sp. nov., Lunulites marambionis sp. nov., Otionellina antarctica sp. nov. and Otionellina eocenica sp. nov. Some other taxa recognized in the studied material, such as Reticrescis plicatus , Uharella seymourensis and Celleporaria mesetaensis , were previously described from the lowermost (Telm1) or uppermost parts (Telm6–7), thus their stratigraphical ranges within the La Meseta Formation are extended. The diverse growth-forms of the bryozoans include a sole ball-shaped celleporiform colony and reticulated and bilamellar-foliaceous colony, as well as rich encrusting and free-living forms (so-called sand faunas), indicating the existence of locally restricted shallow-marine environments. This is particularly true in the middle and upper parts of the La Meseta Formation (Telm4–7). Reticulated, spheroidal and robust, branched colonies, which thrived in the environmental conditions of the lower part (Telm1), are represented only by a sparse bryozoan biota in the upper part of the La Meseta Formation. Lunulitiforms, such as Lunulites and Otionellina which are warm water, free-living bryozoans, dominate in the siliciclastic sediments of Telm5, but erect folded sheets forming a shell bed composed of ? Goodonia occur in Telm6–7. These three genera are recognized in Antarctica for the first time. The austral genus Otionellina has its earliest fossil record here, showing close biogeographical links with the Late Eocene–Miocene faunas of Australia and New Zealand. The taxonomic composition of the studied fauna together with their growth forms is a very good tool for reconstructing palaeoenvironmental conditions in the middle and upper parts (Telm4–7) of the La Meseta Formation, deposited during the Late Ypresian–Priabonian.