Seymour Island Research Articles

Miocene and Plio-Pleistocene foraminiferal assemblages from Seymour Island, Antarctica

AbstractHere we describe new microfossil assemblages for the Miocene Hobbs Glacier Formation and the first possibly indigenous assemblages for the Plio-Pleistocene Weddell Sea Formation on Seymour Island, West Antarctica. The assemblages are composed mainly of foraminifers, but radiolarians, calcitarchs and poriferan sclerites are also present. For the Hobbs Glacier Formation, we report the foraminifers Bolivina sp., Oolina globosa and Rosalina cf. globularis; and for the Weddell Sea Formation, we report Favulina hexagona, Globigerinita uvula, Globocassidulina cf. subglobosa and Psammosphaera fusca. The low abundance and diversity of microfossils, allied with the complex taphonomical processes that prevailed in Antarctic glacial–marine palaeoenvironments, make it impossible to define whether the assemblages are composed of a mixture of indigenous and re-elaborated specimens or exclusively of re-elaborated remains. Nevertheless, the indigenous nature of some specimens is suggested by their inherent fragility, excellent preservation and/or taxonomic association with indigenous assemblages from correlated strata. The taxonomic compositions are not directly comparable with other Antarctic assemblages, although most of the species were previously reported from pre-Quaternary or modern deposits of both West and East Antarctica. This lack of correspondence is probably due to preservation biases, but any further significance is hidden by the complex taphonomy of the deposits.

Dinoflagellate cysts of the La Meseta Formation (middle to late Eocene), Antarctic Peninsula: implications for biostratigraphy, palaeoceanography and palaeoenvironment

AbstractDinoflagellate cyst assemblages recovered from the La Meseta Formation cropping out in Seymour Island, Antarctic Peninsula, are studied herein and their distribution is compared with the biostratigraphic scheme available for the Palaeogene of the Southern Ocean and other high-latitude regions. In this way, the La Meseta Formation is dated as middle Lutetian to Priabonian (46.2–36 Ma), which differs from the age provided by other fossils, isotopes and also with the magnetostratigraphic scheme recently performed for the unit. The dinoflagellate cyst data support the proposal of ocean circulation patterns on the South American Shelf prior to the opening of Drake Passage. Assemblages from the La Meseta Formation contain Antarctic-endemic taxa which are also dominant in several circum-Antarctic sites, located south of 45° S. Their distribution reflects an ocean-circulation scheme with wide clockwise gyres surrounding Antarctica that were disrupted as a consequence of the deepening and definitive apertures of the Tasmanian Gateway and Drake Passage towards the Eocene/Oligocene transition. The palaeoenvironmental inference based on the S/D ratio (sporomorphs versus dinoflagellate cysts) and the P/G ratio (peridinioid versus gonyaulacoid dinoflagellate cysts) suggests an overall trend through the section from marine-dominated assemblages with poorly productive waters in the lower part of the section to more terrestrially dominated assemblages with increasing productivity in the upper part of the unit, reflecting a shallowing trend to the top.

New fossil woods from lower Cenozoic volcano‐sedimentary rocks of the Fildes Peninsula, King George Island, and the implications for the trans‐Antarctic Peninsula Eocene climatic gradient

AbstractTen embedded fossil logs sampled in situ from the middle Eocene volcano‐sedimentary rocks close to Suffield Point in the Fildes Peninsula, King George Island, Antarctica, are assigned to Protopodocarpoxylon araucarioides Schultze‐Motel ex Vogellehner, Phyllocladoxylon antarcticum Gothan, Agathoxylon antarcticum (Poole & Cantrill) Pujana et al., A. pseudoparenchymatosum (Gothan) Pujana et al. and an unidentified angiosperm wood. Differences in the taxonomic representation and growth‐ring characters of the Eocene woods on King George Island and coeval assemblages from Seymour Island, on the western and eastern sides of the Antarctic Peninsula respectively, are interpreted to result from environmental and climatic gradients across the Peninsula Orogen during the early Palaeogene. In particular, a precipitation gradient inferred across the Peninsula at that time might have been induced by a rain‐shadow effect.

A giant elasmosaurid (Sauropterygia; Plesiosauria) from Antarctica: New information on elasmosaurid body size diversity and aristonectine evolutionary scenarios

Aristonectines show a highly derived morphology among elasmosaurid plesiosaurs, including some species with large body size. A new postcranial skeleton is described from the uppermost Maastrichtian levels of the López de Bertodano Formation, Seymour Island (= Marambio), Antarctica, being referred to as cf. Aristonectes sp; the most striking feature of the specimen described is its large body size, among the largest elasmosaurids worldwide. The occurrence of this specimen, located approximately 2.3 m or less below the Cretaceous-Paleogene (K/Pg) boundary, indicates the persistence of aristonectines at high latitudes and also it verifies their chronostratigraphical distribution until the end Cretaceous, before the mass extinction. Elasmosaurid diversity in terms of body size, possible relation of this body size, the trophic niche and abiotic drivers in aristonectine evolution are discussed. The body size inferred for MLP 89-III-3-1 seems to indicate an environment with high primary productivity, suggesting that these conditions persisted until the K/Pg mass extinction.

New records of mummified crabeater seals on islands bordering Admiralty Sound, Weddell Sea

AbstractNinety-six mummified crabeater seals discovered at Seymour Island (Isla Marambio) are reported. Each specimen was georeferenced, photographed and assigned to five different taphonomic states. Previous work stated that seals at Seymour Island get stranded inland around the breeding season. However, it is not clear if the species breeds in this area. The abundance of crabeater seals and the ice condition along Admiralty Sound (Estrecho Bouchard) were obtained by aerial surveys during spring (2015–17). It appears that the species uses the strait as a passage to breeding grounds. Under heavy ice conditions, the seals become stranded in the middle section of this strait and wander inland through a valley that represents the mouth of an ephemeral stream that ends at the pack ice level. This situation was observed in 2014 and 2015 when recently dead seals were found, evidencing that this natural trap is still active. Nonetheless, in 2016 and 2017, during an early breakup of Admiralty Sound, the seals that remained in the area were more numerous than in 2015 but they did not get stranded inland. This early breakup may encourage the seals to breed there in the presence of open water areas with ice floes.

Indigenous agglutinated foraminifera from the Eocene La Meseta Formation, Seymour Island, West Antarctica

Indigenous agglutinated foraminifera from the Eocene La Meseta Formation, Seymour Island, West Antarctica

Gigantic mysticete predators roamed the Eocene Southern Ocean

AbstractModern baleen whales (Mysticeti), the largest animals on Earth, arose from small ancestors around 36.4 million years ago (Ma). True gigantism is thought to have arisen late in mysticete history, with species exceeding 10 m unknown prior to 8 Ma. This view is challenged by new fossils from Seymour Island (Isla Marambio), Antarctica, which suggest that enormous whales once roamed the Southern Ocean during the Late Eocene (c. 34 Ma). The new material hints at an unknown species of the archaic mysticete Llanocetus with a total body length of up to 12 m. The latter is comparable to that of extant Omura's whales (Balaenoptera omurai Wada et al. 2003), and suggests that gigantism has been a re-occurring feature of mysticetes since their very origin. Functional analysis including sharpness and dental wear implies an at least partly raptorial feeding strategy, starkly contrasting with the filtering habit of living whales. The new material markedly expands the size range of archaic mysticetes, and demonstrates that whales achieved considerable disparity shortly after their origin.

Eocene bryozoan assemblages from the La Meseta Formation of Seymour Island, Antarctica

Early to Late Eocene bryozoans from the La Meseta Formation of Seymour Island were collected at two localities within the Cucullaea I Allomember (Telm4 and Telm5) on the northwestern side of the island and in two localities within the Submeseta Allomember (Telm6 and Telm7) on the northeastern side. This fauna is represented by cyclostomes of the suborders Tubuliporina and Cerioporina and suborders of Neocheilostomata, among which nine species have been recognized. The following new species are introduced: Micropora nordenskjoeldi sp. nov., Lunulites marambionis sp. nov., Otionellina antarctica sp. nov. and Otionellina eocenica sp. nov. Some other taxa recognized in the studied material, such as Reticrescis plicatus , Uharella seymourensis and Celleporaria mesetaensis , were previously described from the lowermost (Telm1) or uppermost parts (Telm6–7), thus their stratigraphical ranges within the La Meseta Formation are extended. The diverse growth-forms of the bryozoans include a sole ball-shaped celleporiform colony and reticulated and bilamellar-foliaceous colony, as well as rich encrusting and free-living forms (so-called sand faunas), indicating the existence of locally restricted shallow-marine environments. This is particularly true in the middle and upper parts of the La Meseta Formation (Telm4–7). Reticulated, spheroidal and robust, branched colonies, which thrived in the environmental conditions of the lower part (Telm1), are represented only by a sparse bryozoan biota in the upper part of the La Meseta Formation. Lunulitiforms, such as Lunulites and Otionellina which are warm water, free-living bryozoans, dominate in the siliciclastic sediments of Telm5, but erect folded sheets forming a shell bed composed of ? Goodonia occur in Telm6–7. These three genera are recognized in Antarctica for the first time. The austral genus Otionellina has its earliest fossil record here, showing close biogeographical links with the Late Eocene–Miocene faunas of Australia and New Zealand. The taxonomic composition of the studied fauna together with their growth forms is a very good tool for reconstructing palaeoenvironmental conditions in the middle and upper parts (Telm4–7) of the La Meseta Formation, deposited during the Late Ypresian–Priabonian.

Indirect effects of climate change altered the cannibalistic behaviour of shell-drilling gastropods in Antarctica during the Eocene.

The fossil record from Seymour Island, Antarctic Peninsula, provides a record of biotic response to the onset of global climatic cooling during the Eocene. Using drilling traces—small, round holes preserved on prey shells—we examined the effect of a cooling pulse 41 Ma on the cannibalistic behaviour of predatory naticid gastropods. We predicted that cannibalistic attacks would decline in response to the cooling climate, reflecting reduced activity levels, energy requirements and constraints on the chemically aided drilling process of the naticids. Surprisingly, however, cannibalism frequencies did not change. This counterintuitive result is best explained by a sharp reduction in durophagous (shell-crushing) predation in shallow-benthic communities in Antarctica that also occurred as the climate cooled. Reduced durophagous predation may have created a less-risky environment for foraging naticids, stimulating cannibalistic behaviour. The change in the top-down control exerted by shell-crushing predators on naticids may have counteracted the direct, negative effects of declining temperatures on the predatory performance of naticids. Our results suggest that the long-term consequences of climate change cannot be predicted solely from its direct effects on predation, because the temperature can have large indirect effects on consumer–resource interactions, especially where risk-effects dominate.

High benthic methane flux in low sulfate oceans: Evidence from carbon isotopes in Late Cretaceous Antarctic bivalves

The shell material of marine benthic bivalves provides a sensitive archive of water chemistry immediately above the sediment–water interface, which in turn is affected by sedimentary geochemistry and redox reactions. Sulfate has a major controlling effect on sedimentary carbon cycling, particularly the processes of methane production and oxidation, with lower concentrations of sulfate likely resulting in an increase in sedimentary methane production. Whilst it is accepted that ocean sulfate varied markedly across the Phanerozoic, evidence of changes in methane production in sediments has so far been lacking. There is potential for the oxidation products of sedimentary methane to be preserved and detected in marine fossils. Here we present the results of high resolution carbonate isotope records from two taxa of well-preserved shallow-infaunal bivalve (Lahillia and Cucullaea) collected from the marine shelf succession across the Cretaceous–Paleogene (K–Pg) boundary in Seymour Island, Antarctica. The succession has pre-existing subtle indications of more abundant methane, and the time period is characterized by much lower marine sulfate concentrations than modern.These shell carbonate–carbon isotope records vary widely: at one extreme, shells have typical average values and small ranges compatible with a contemporaneous marine dissolved inorganic carbon (DIC) source and modern-style sedimentary carbon cycling. At the other, the shells have large-amplitude annual cycles of carbon isotopic variability of up to 23.8‰ within a single year of growth and shell carbonate δ13C compositions as negative as −34‰. Shells with these increased ranges and unusually negative values are found at discrete intervals and across both bivalve taxa. The contribution of methane required to explain the most negative carbonate–carbon isotopic values in the bivalve shells is extremely high (between 30 to 85% of bottom-water DIC based on mass balance calculations). Records of organic-carbon isotopes from the same succession remained between −26.1 and −21.7‰ throughout, suggesting that methane influence was restricted to bottom-waters. A lack of authigenic carbonate in the section indicates that methane oxidation progressed aerobically and may have provided a significant driver for transient bottom water de-oxygenation.Where methane addition is indicated, the seasonal sensitivity precludes control by methane hydrates. We argue that these data represent the increased importance and sensitivity of methanogenesis in the sediments, enabled by lower ocean sulfate concentrations during the Late Cretaceous. The tendency towards a more dynamic role for marine methane production and oxidation is likely to apply to other times of low marine sulfate in Earth's history.

A critical review of Antarcticeras Doguzhaeva, 2017 – teuthid affinities can explain the poorly mineralized phragmocone

Antarcticeras from the Early Eocene of Seymour Island (Antarctica) has recently been described as a representative of the ‘Paracoleoidea’, a previously unknown cephalopod subclass. After reinvestigation of the type material of Antarcticeras nordenskjoeldi Doguzhaeva, 2017, the justification of this new cephalopod subclass is here rejected. Phragmocone characteristics allow alternative interpretations and these justify to consider the poorly mineralized phragmocone of Antarcticeras being transitional between a heavily mineralized, fully functional phragmocone and a completely demineralized gladius of an oegopsid coleoid. Eocene Antarcticeras therefore might represent the first evidence of teuthid shell remains in the fossil record. Its poor state of mineralization accordingly suggests remarkably late development of teuthid gladii.

An intertidal balanomorph Hexaminius venerai sp. nov. (Austrobalanidae) colonizing a log of Podocarpoxylon from the La Meseta Formation (Eocene), Seymour Island, Antarctica: a glimpse of Antarctic antiquity

The sessile barnacle Hexaminius venerai sp. nov. (Tetraclitoidea: Austrobalanidae) is described from the middle Eocene La Meseta Formation of Seymour Island, Antarctic Peninsula. Hexaminius venerai sp. nov. is the earliest known record of the genus in the Antarctic, the first occurrence of Hexaminius from outside Australian waters and the first record of a fossil cirripede attached to the substrate from the Antarctic. Exceptional preservation of more than 200 specimens, some of which retain opercula within the shell, is discussed. In life, the cirripedes were attached to a tree trunk tentatively identified as Podocarpoxylon, a South Hemisphere conifer. Hexaminius venerai sp. nov. is a survivor of an early phase in balanid radiation, prior to the development of strong radially-interlocked parietes.https://zoobank.org/urn:lsid:zoobank.org:pub:EAA2EB5C-BC57-41FA-93A0-046AF5FD59EB

“Business as usual”: Drilling predation across the K-Pg mass extinction event in Antarctica

A survey of 759 predatory drill holes in Late Cretaceous and Paleogene molluscan and serpulid worm prey from Seymour Island (Antarctica) has allowed exploration of the effects of the K-Pg mass extinction on predator-prey dynamics at this high latitude site. Circular holes (0.62–6.41 mm in diameter), suggesting a large gastropod predator, most probably “Vanikoropsis” arktowskiana, occur throughout the study interval. Analysis suggests that the driller was a catholic predator capable of attacking a range of prey types, showing some degree of size and stereotypic handling behaviour. Although there were changes in prey choice across the extinction event, these reflect changes in available prey, and in particular limited options immediately post extinction. There were no significant changes in failure rate (Prey Effectiveness) over the mass extinction. Our findings suggest that at this site, for this particular predator, aside from menu choice, predation dynamics post-extinction was “business as usual”.

The impact of the Cretaceous–Paleogene (K–Pg) mass extinction event on the global sulfur cycle: Evidence from Seymour Island, Antarctica

The Cretaceous–Paleogene (K–Pg) mass extinction event 66 million years ago led to large changes to the global carbon cycle, primarily via a decrease in primary or export productivity of the oceans. However, the effects of this event and longer-term environmental changes during the Late Cretaceous on the global sulfur cycle are not well understood. We report new carbonate associated sulfate (CAS) sulfur isotope data derived from marine macrofossil shell material from a highly expanded high latitude Maastrichtian to Danian (69–65.5 Ma) succession located on Seymour Island, Antarctica. These data represent the highest resolution seawater sulfate record ever generated for this time interval, and are broadly in agreement with previous low-resolution estimates for the latest Cretaceous and Paleocene. A vigorous assessment of CAS preservation using sulfate oxygen, carbonate carbon and oxygen isotopes and trace element data, suggests factors affecting preservation of primary seawater CAS isotopes in ancient biogenic samples are complex, and not necessarily linked to the preservation of original carbonate mineralogy or chemistry. Primary data indicate a generally stable sulfur cycle in the early-mid Maastrichtian (69 Ma), with some fluctuations that could be related to increased pyrite burial during the ‘mid-Maastrichtian Event’. This is followed by an enigmatic +4‰ increase in δ34SCAS during the late Maastrichtian (68–66 Ma), culminating in a peak in values in the immediate aftermath of the K–Pg extinction which may be related to temporary development of oceanic anoxia in the aftermath of the Chicxulub bolide impact. There is no evidence of the direct influence of Deccan volcanism on the seawater sulfate isotopic record during the late Maastrichtian, nor of a direct influence by the Chicxulub impact itself. During the early Paleocene (magnetochron C29R) a prominent negative excursion in seawater δ34S of 3–4‰ suggests that a global decline in organic carbon burial related to collapse in export productivity, also impacted the sulfur cycle via a significant drop in pyrite burial. Box modelling suggests that to achieve an excursion of this magnitude, pyrite burial must be reduced by >15%, with a possible role for a short term increase in global weathering rates. Recovery of the sulfur cycle to pre-extinction values occurs at the same time (∼320 kyrs) as initial carbon cycle recovery globally. These recoveries are also contemporaneous with an initial increase in local alpha diversity of marine macrofossil faunas, suggesting biosphere-geosphere links during recovery from the mass extinction. Modelling further indicates that concentrations of sulfate in the oceans must have been 2 mM, lower than previous estimates for the Late Cretaceous and Paleocene and an order of magnitude lower than today.

Terrestrial and marine floral response to latest Eocene and Oligocene events on the Antarctic Peninsula

ABSTRACTPalynological results from opposite sides of the northernmost Antarctic Peninsula provide insight on terrestrial vegetation and sea-surface conditions immediately before the Eocene–Oligocene transition (EOT), through Early Oligocene glacial conditions and the subsequent Late Oligocene interglacial interval. A latest Eocene sample set from the uppermost La Meseta Formation on Seymour Island, James Ross (back-arc) Basin, records a low-diversity Nothofagus (southern beech)-dominated vegetation with some podocarp conifers similar to Valdivian-type forest found today in Chile and Argentina. Marine organic-walled phytoplankton include leiospheres and Eocene dinoflagellate cysts such as Vozzhennikovia rotunda, V. apertura, Senegalinium asymmetricum and Spinidinium macmurdoense. Immediately before the EOT near the top of the section the decrease in terrestrial palynomorphs, increase in reworked specimens, disappearance of key dinocysts, and overwhelming numbers of sea-ice-indicative leiospheres plus the small dinoflagellate cyst Impletosphaeridium signal the onset of glacial conditions in a subpolar climate. Early to Late Oligocene samples from the Polonez Cove and Boy Point formations on King George Island, South Shetland Islands (magmatic arc), yielded an extremely depauperate terrestrial flora, likely resulting in part from poor vegetation cover during the Polonez Glaciation but also because of destruction of vegetation due to continued regional volcanism. The prevalence of sea-ice-indicative leiospheres in the marine palynomorph component is consistent with polar to subpolar conditions during and following the Polonez Glaciation.

Conifer Fossil Woods from the Sobral Formation (Lower Paleocene, Western Antarctica)

Conifer fossil woods represent 54% of an assemblage of 116 specimens collected from sediments of the Sobral Formation in Seymour (Marambio) Island, Western Antarctica. These woods are anatomically described in detail and assigned to seven fossil-species of the following fossil-genera: Agathoxylon (Araucariaceae), Podocarpoxylon, Phyllocladoxylon, Protophyllocladoxylon (Podocarpaceae), and Cupressinoxylon (Podocarpaceae/Cupressaceae). The conifer wood assemblage reveals that the most common woods are those of Agathoxylon, therefore indicating a relative abundance of the Araucariaceae in the Antarctic Paleocene forests. This abundance of the Araucariaceae woods is locally continued in the overlying Cross Valley Formation (Paleocene). Podocarpaceae woods are found in almost similar proportions to those of the Araucariaceae. Almost the other half of the fossil woods are dicotyledon woods. The proportions of the identified fossil wood taxa are consistent with those of the palynological studies of the same stratigraphic unit.

Skates and rays (Elasmobranchii, Batomorphii) from the Eocene La Meseta and Submeseta formations, Seymour Island, Antarctica

Eocene deposits of the famous La Meseta Formation of Seymour Island, Antarctic Peninsula, yielded the most diverse Paleogene fossil elasmobranch association of the Southern Hemisphere. In this assemblage, sharks clearly dominate the fauna, whereas batoids are very rare components. Herein, we describe two new taxa of cold water tolerant skates, Marambioraja leiostemma gen. et sp. nov., and Mesetaraja maleficapelli gen. et sp. nov., two new species of the genus Raja, Raja amphitrita sp. nov. and Raja manitaria sp. nov., as well as remains of warm water adapted myliobatiforms. It is, however, not possible to unambiguously assign these remains either to Myliobatidae or Rhinopteridae, or to any specific genus. Previously reported remains of Raja/Bathyraja sp. are assigned to the new described species Raja manitaria sp. nov. The biogeographic distribution of extant and extinct rays and skates clearly shows that both groups are more widely distributed today than in the past, and additionally seem to have been more diverse in the Northern than the Southern Hemisphere. The occurrence, albeit rare of isolated teeth of skates (Rajidae) and rays (Myliobatidae) in the La Meseta Formation representes a minimum age constraint for their first appearance in the Southern Ocean.http://www.zoobank.org/urn:lsid:zoobank.org:pub:E507D86C-FFEC-4047-A337-AE77606BB1A6

Health Status of Great Frigatebirds (Fregata minor) Determined by Haematology, Biochemistry, Blood Gases, and Physical Examination.

The great frigatebird, Fregata minor, is a widely distributed seabird native to the Galápagos archipelago. Haematology and blood chemistry parameters have been published for this species but not from the San Cristóbal and North Seymour great frigatebird breeding colonies. Analyses were run on blood samples drawn from 25 great frigatebirds captured by hand at their nests at Punta Pitt on San Cristóbal Island and 30 birds on North Seymour Island, Galápagos Islands. A portable blood analyser (iSTAT) was used to obtain near immediate field results for pH, pO2, pCO2, TCO2, HCO3- , haematocrit (Hct), haemoglobin (Hb), sodium (Na), potassium (K), chloride (Cl), ionized calcium (iCa), creatinine, urea nitrogen, anion gap and glucose. Blood lactate was measured using a portable Lactate Plus™ analyser. Average heart rate, respiratory rate, body weight, body temperature, biochemistry and haematology parameters were comparable to healthy individuals of other Fregatidae. The reported results provide baseline data that can be used for comparisons among populations and in detecting changes in health status among Galápagos great frigatebirds.

Stratigraphy, sedimentology and paleontology of Upper Cretaceous deposits of Day Nunatak, Snow Hill Island, Antarctica

Day Nunatak exposes an actively emergent stratigraphic section located on Snow Hill Island to the east of the Antarctic Peninsula. Strata exposed on Snow Hill Island were deposited in the James Ross Basin, which includes marine units of Cretaceous–Paleogene (K–Pg) age. Here we provide the first report of the sedimentology and paleontology of Day Nunatak, and place it into the broader stratigraphic context of the basin. Day Nunatak was previously unexplored due to difficulties accessing the site, and the historically poor exposure which has recently improved due to warming of the Antarctic Peninsula. Deposits exposed at Day Nunatak are assigned to the Karlsen Cliffs Member (KCM) of the Snow Hill Island Formation, and are better preserved than deposits at the type section of the KCM at Karlsen Cliffs, which has been altered by cross-cutting basaltic dikes. Correlation of lithostratigraphic and biostratigraphic data from Day Nunatak with other outcrops in the basin allows the section to be placed within a previously developed stratigraphic framework, and assigned an early Maastrichtian age. Our observations, and previous descriptions of the KCM, are consistent with a middle- to inner-shelf depositional environment below storm-wave base. Ammonites of the genus Gunnarites are very common, with other ammonite genera and benthic mollusks an order of magnitude less common. Stable carbon and oxygen isotope values obtained from bivalve shells are similar to values previously reported from Seymour Island, and suggest seawater temperatures of ∼7 °C. Measured bedding orientations suggest the presence of a structural offset between Day Nunatak and other sections exposed further north on Snow Hill Island. Day Nunatak preserves a similar depositional environment to deposits reported from the uppermost Maastrichtian on Seymour Island, and is the deeper-water equivalent of contemporaneous proximal sections reported from Vega Island.

Recognition of a likely two phased extinction at the K-Pg boundary in Antarctica

The southernmost Cretaceous – Paleogene (K-Pg) outcrop exposure is the well-studied exposure on Seymour Island, Antarctica. Deposition across the K-Pg boundary there is uninterrupted, and as a consequence the ammonite fossil record is commonly used to test statistical methods of evaluating mass extinctions to account for the incompleteness of the fossil record. Numerous detailed fossil data sets from Seymour Island, comprised dominantly of mollusks, have been published over the last 30 years, but in most cases have not received statistical treatment. Here a previously published statistical technique is modified, automated, and applied to all published macrofossil data sets available from Seymour Island. All data sets reveal likely evidence of two separate multi-species extinctions, one synchronous with bolide impact evidence at the K-Pg boundary, and another 45 ± 15 meters (~140–290 ky) below the boundary. The apparent earlier extinction primarily affects benthic mollusks, while the boundary extinction primarily affects ammonites. While there is no unique sedimentological change over the interval where the earlier extinction is identified, it is impossible to exclude the possibility that this pattern is stratigraphically controlled. The automation of this technique allows it to be applied easily to other large fossil data sets.